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. 2022 Sep 27;135(16):1894–1912. doi: 10.1097/CM9.0000000000002391

Table 2.

A summary of recommended biomarkers for testing.

Adjuvant therapy (evidence level)
Biomarkers Function Tumor location Clinical significance (evidence level) Diagnosis (evidence level) Radiation therapy Systemic therapy Follow-up (evidence level) Test method Recommendation grade
Copy number alterations
 22q A critical role in meningiomagenesis by dysregulating oncogene and tumor suppressor activity Convexity and spine 22q loss is significantly associated with worse clinical outcome (evidence 2B) Unknown Unknown Sensitive to sunitinib (evidence 3) Annually within 5 years (evidence 3) Karyotyping, FISH, Array-CGH, NGS Grade I
 1p Unknown Unknown 1p loss is associated with worse clinical outcome (evidence 2A). 1p/14q loss increases the risk of tumor recurrence (evidence 2A) Unknown Unknown Sensitive to sunitinib (evidence 3) Every 6 months within 5 years (evidence 3) Karyotyping, FISH, Array-CGH, NGS Grade I
 14q Unknown Unknown 1p/14q loss increases the risk of tumor recurrence (evidence 2A) Unknown Unknown Unknown Every 6 months within 5 years (evidence 3) Karyotyping, FISH, Array-CGH, NGS Grade I
 18q Unknown Unknown 18q loss increases the risk of tumor short-term recurrence (evidence 2B) Unknown Unknown Unknown Annually within 5 years (evidence 3) Karyotyping, FISH, Array-CGH, NGS Grade III
Gene mutation signatures
NF2 NF2 produces merlin that functions as a tumor suppressor by inhibiting cell growth Convexity and spine NF2 mutation is associated with worse clinical outcome, increases the risk of tumor recurrence (evidence 2A) Unknown Unknown Unknown Annually within 5 years (evidence 3) NGS, Sanger sequence, MLPA, FISH, IHC Grade I
TERT promoter TERTp regulates TERT gene expression and maintains normal cell mitosis Unknown TERTp mutation is associated with worse clinical outcome, and increases the risk of tumor recurrence (evidence 1A) Histological criteria of WHO grade 3 (evidence 3) TERTp alteration is associated with radiation sensitivity in de novo high-grade meningiomas (evidence 2B) Unknown Every 3–6 months indefinitely (evidence 3) NGS, Sanger sequence Grade I
CDKN2A/B CDKN2A/B regulates cell cycle and inhibits tumor cell proliferation Unknown CDKN2A/B mutation is associated with worse clinical outcome, and increases the risk of tumor recurrence (evidence 2A) Histological criteria of WHO grade 3 (evidence 3) Unknown Unknown Every 6 months within 5 years (evidence 3) NGS, Sanger sequence Grade I
AKT1, PIK3CA PI3K pathway serves to disconnect the control of cell growth, survival and metabolism from exogenous growth stimuli Middle fossa skull base AKT1 and PIK3CA mutation increases the risk of tumor recurrence (evidence 2A) Unknown Unknown Everolimus-octreotide (evidence 2A); everolimus-bevacizumab (evidence 2A) Every 6 months within 5 years (evidence 3) AKT1: NGS, RT-PCR; PIK3CA: NGS, Sanger sequence, RT-PCR Grade I
SMO, SUFU Driver genes in Hedgehog pathway Middle fossa skull base SMO and SUFU mutation increases the risk of tumor recurrence (evidence 2A) Unknown Unknown Unknown Annually within 5 years (evidence 3) SMO: NGS, Sanger sequence; SUFU: NGS, IHC Grade I
TRAF7 Transduction molecules in TNFR mediated signaling pathway Middle fossa skull base TRAF mutation increases the risk of tumor short-term recurrence (evidence 2A) Histological criteria of secretory subtype (evidence 3) Unknown Unknown NGS, Sanger sequence Grade I
KLF4 Zinc finger protein transcription factor, which involves in the regulation of cell proliferation, differentiation and embryonic development Non-midline anterior and central skull base KLF4 mutation is associated with better 5-year progression-free survival and peritumoral brain edema (evidence 2A) Histological criteria of secretory subtype (evidence 3) Unknown Unknown NGS, Sanger sequence Grade I
SMARCE1 Chromatin remodeling complex SWI/SNF involves in encoding part of ATP Spine Common mutation in clear cell subtype (evidence 2A) Reference for diagnosis of secretory subtype (evidence 3) Unknown Unknown NGS, IHC Grade I
BAP1 A tumor suppressor gene that encodes a deubiquitylating enzyme, and inhibits tumor growth by binding to the ring finger domain of BRCA1 Unknown BAP1 mutation is associated with worse clinical outcome, and increases the risk of tumor recurrence (evidence 2A) Reference for diagnosis of rhabdoid subtype (evidence 3) Unknown Unknown Every 6 months within 5 years (evidence 3) NGS, IHC Grade I
DMD DMD locates at Xp21.1-p21.3, encoding dystrophin Unknown DMD mutation is associated with worse clinical outcome, and increases the risk of tumor recurrence (evidence 2B) Unknown Unknown Unknown Annually within 5 years (evidence 3) NGS, IHC Grade I
PBRM1 A tumor suppressor gene encoding the BAF180 subunit of the SWI/SNF complex, and involving in the regulation of tumor cell proliferation and migration Unknown Common mutation in papillary subtype (evidence 2B) Unknown Unknown Unknown NGS Grade III
POLR2A The protein that mediates transcription of all protein-coding genes in eukaryotes Anterior skull base, especially tuberculum sellae POLR2A mutation is associated with better clinical outcome (evidence 2B) Unknown Unknown Unknown NGS, Sanger sequence Grade III
Epigenomic alterations
 H3K27me3 A chromatin modification that is tightly linked to gene repression and plays an essential role in the development and progression of intracranial tumors Unknown H3K27me3 loss increases the risk of tumor short-term recurrence (evidence 2A) Unknown Unknown Unknown Every 6 months within 5 years (evidence 3) IHC Grade I
TIMP3 methylation TIMP3 methylation causes a down-regulation in transcription product and then loss of tumor suppressor activity Unknown TIMP3 methylation increases the risk of tumor short-term recurrence (evidence 2B) Unknown Unknown Unknown Every 6 months within 5 years (evidence 3) Pyrosequencing Grade II
TP73 promoter methylation TP73 promoter methylation regulates downstream transcripts and inhibits tumor growth Unknown TP73 promoter methylation increases the risk of tumor short-term recurrence (evidence 2B) Unknown Unknown Unknown Every 6 months within 5 years (evidence 3) Pyrosequencing Grade II
Hormone receptors
 ER ER participates in regulation of multiple hormones, inhibits cell proliferation and promotes cell apoptosis Unknown High expression of ER is associated with worse clinical outcome (evidence 2B) Unknown Unknown Unknown IHC Grade III
 PR PR participates in regulation of multiple hormones, inhibits cell proliferation and promotes cell apoptosis Unknown High expression of PR is associated with worse clinical outcome (evidence 2B) Unknown Unknown Unknown IHC Grade III
 SSTR2 SSTR2 participates in regulation of multiple hormones, inhibits cell proliferation and promotes cell apoptosis Unknown SSTR2 alterations increase the risk of tumor recurrence (evidence 2B) Octreotide-scanning (evidence 2B) Unknown Everolimus-octreotide (evidence 2B) IHC Grade I
Tyrosine kinases receptors
 VEGFR VEGFR involves in regulation of lymphatic endothelial cells and vascular endothelial cells, and promotes the formation of lymphatic vessels and blood vessels Unknown Expressien of VEGFR is associated with higher WHO grade and shorter progression-free survival (evidence 2B) Unknown Unknown Bevacizumab (evidence 2A) IHC Grade II

AKT1: Protein kinase B; Array-CGH: Array-comparative genomic hybridization; ATP: Adenosine triphosphate; BAF: BRM-associated factor; BAP1: BRCA1 associated protein 1; BRCA1: Breast cancer 1; CDKN2A/B: Cyclin-dependent kinase inhibitor 2A/B; DMD: Duchenne muscular dystrophy; ER: Estrogen receptor; FISH: Fluorescence in situ hybridization; H3K27me3: Trimethylation of lysine 27 on histone 3; IHC: Immunohistochemistry; KLF4: Krüppel-like factor 4; MLPA: Multiple ligation-dependent probe amplification; NF2: Neurofibromin 2; NGS: Next generation sequencing; PBRM1: Polybromo-1; PI3K: Phosphosphatidylinositol-3-kinase; PIK3CA: Phosphatidylinositol-4,5-bisphosphate 3-kinase catalytic subunit alpha isoform; POLR2A: RNA polymerase II; PR: Progesterone Receptor; RT-PCR: Reverse transcription-polymerase chain reaction; SMARCE1: SWI/SNF Related, matrix associated, actin dependent regulator of chromatin, subfamily E, member 1; SMO: Smoothened; SSTR2: Somatostatin receptor 2; SUFU: Suppressor of fused homolog; SWI/SNF: Switching defective/sucrose non-fermenting; TERT: Telomerase reverse tranase; TERTp: Telomerase reverse transcriptase promoter; TIMP3: Tissue inhibitor of matrix metalloproteinase 3; TNFR: Tumor necrosis factor receptor; TP73: Tumor protein p73; TRAF7: Tumor necrosis factor receptor associated factor 7; VEGFR: Vascular endothelial growth factor receptor; WHO: World Health Organization; -: Not available.