Growth-promoting effects of TSG perturbations in vitro. A–C, Representative trajectories of cell lineages expressing unique dual sgRNAs grown in full, minimal, and TGFβ1-supplemented media, respectively. D, Distribution of gene-level fitness of 1,378 perturbations in the three conditions. E, The top five single-gene effects on fitness in the three conditions. Data are presented as mean ± SEM. F, Relative percentage of DKO/SKO normalized counts in the three conditions. Data are presented as mean ± SEM. G, A heatmap that shows the comparison of growth-promoting effects in vitro and tumorigenic effects in vivo of SKOs.