Mannose receptor
(MR, CD206) glycopolymer ligands. (a) Structure
of CD206-binding multivalent glycans designed in this study. These
glycopolymer ligands possess variable chain length (n = 32 and 187) and either mannose (Man) or SO4-3-galactose
(SO4-3-Gal) CD206-binding sugar units. Glycopolymer contents
in Man (x) and SO4-3-Gal (y) are systematically varied (0, 33, 66, and 100%), using galactose
(Gal) as the remaining non-CD206-binding polymer repeating units.
Glycopolymers containing only non-CD206-binding Gal repeating units,
G32 and G187, were used in this study as negative
control ligands. All glycopolymers are fluorescently labeled with
1% mol/mol of either 2,4,5,7,7′-pentafluorofluorescein (Oregon
Green (OG), λex/em 488/530 nm) or Nile blue (λex/em 629/670 nm) to enable detection in in vitro and in vivo
assays. Table S1 shows the full composition
and characterization of the glycopolymers. (b) Schematic representation
of the mannose receptor (MR, CD206). CR: Cysteine-rich domain which
binds 3/4-O-sulfate galactose (SO4-3/4)-Gal)
and N-(acetyl galactosamine) (SO4-3/4)-GalNAc);
FNII: collagen-binding fibronectin type II domain; CTLD: C-type lectin-like
domain which binds mannose (Man), fucose (Fuc), and N-(acetyl glucosamine) (GlcNAc). (c) Binding of glycopolymers to CTLD4-7
and CR CD206 fragments as assessed by ELISA assay. Glycopolymers were
first immobilized, and then for (i) lectin ELISA tests, they were
treated with CTLD4-7-Fc or CR-Fc CD206 Fc-chimeras, while for (ii)
inhibition ELISA tests, CTLD4-7-Fc and CR-Fc fragments were first
co-incubated with 25 mM 2′-azidoethyl-O-α-D-mannopyranoside
or 3-O-sulfo-2′-azidoethyl-O-β-D-galactopyranoside monovalent ligands and then added to
the immobilized glycopolymers. In both sets of experiments, CD206
fragments were detected with anti-human IgG Fc-specific, alkaline
phosphatase conjugates. G32 = (G100%)32, G187 = (G100%)187, M187 = (M100%)187, and S187 = (S100%)187. Results are from three independent experiments
performed in quadruplicate. Data represent mean ± s.d. CSA: chondroitin
sulfate A.