Table 2.

Linear mixed effect models with maximum likelihood population effects testing the influence of landscape resistance variables on genetic differentiation in devils, tumors, and tumor clusters 1 and 2. Model performance was evaluated by AICc averaged over 1,000 bootstrap iterations, with models with ΔAICc < 2 highlighted. Marginal R² (mR²) is the proportion of overall variation explained by the model fixed effects and % Top Model is the percentage of times the model was the top performing model over 1,000 bootstraps, as determined by AICc support.

	Variable	K	AICc	ΔAICc	mR2	% Top Model
Devils	Roads	5	−7364.67	0.00	0.379	38.4
	Precip. seasonality	4	−7359.56	4.66	0.269	1.7
	Elevation relief ratio	4	−7358.72	5.49	0.253	6.8
	Temp. annual range	4	−7357.35	6.87	0.255	0.0
	Distance	2	−7357.64	5.97	0.249	52.7
	Elevation	4	−7357.35	6.86	0.253	0.1
	Annual precip.	4	−7355.11	9.11	0.255	0.0
	Land cover	13	−7322.04	50.90	0.319	0.3
All tumors	Elevation	4	−7083.89	0.00	0.015	67.7
	Precip. seasonality	4	−7082.19	1.70	0.030	30.5
	Annual precip.	4	−7080.23	3.67	0.020	1.4
	Elevation relief ratio	4	−7072.25	11.64	0.012	0.3
	Distance	2	−7070.41	13.48	0.009	0.1
	Temp. annual range	4	−7069.27	14.62	0.006	0.0
	Land cover	13	−7067.75	16.14	0.024	0.0
	Roads	5	−7065.33	18.57	0.009	0.0
Tumor cluster 1	Elevation relief ratio	4	−7627.07	0.00	0.026	77.2
	Annual precip.	4	−7625.59	1.48	0.019	11.5
	Precip. seasonality	4	−7624.56	2.51	0.016	4.6
	Distance	2	−7624.19	2.88	0.009	6.4
	Elevation	4	−7623.73	3.35	0.021	0.2
	Temp. annual range	4	−7623.67	3.40	0.009	0.0
	Roads	5	−7622.07	5.01	0.020	0.1
	Land cover	13	−7616.24	10.83	0.016	0.0
Tumor cluster 2	Elevation	4	−7972.78	0.00	0.025	77.6
	Annual precip.	4	−7969.02	3.76	0.043	2.1
	Elevation relief ratio	4	−7967.41	5.37	0.044	10.3
	Precip. seasonality	4	−7965.74	7.04	0.024	8.6
	Distance	2	−7959.39	13.39	0.007	0.5
	Roads	5	−7958.98	13.80	0.047	0.9
	Land cover	13	−7957.66	15.12	0.053	0.0
	Temp. annual range	4	−7958.77	14.01	0.006	0.0