(A) Average action potentials (APs) per light pulse stimulation of each voltage trace (for all light intensities) revealed differences in circuit strength across frontal cortical trans-claustral circuits depending on type I and II claustrum projection neuron target (Kruskal Wallis test for multiple comparisons: p < 0.0001; n = 570 type I cells and n = 525 type II cells total).
(B and C) Clustered frontal cortico-claustro-cortical circuits were detected by subgraph extraction (cluster marked in black) for type I and type II claustrum neuron subtypes. Type I: detected subgraph from ACC and plPFC to PtA and V1/V2. Type II: detected subgraph from ACC and plPFC to ACC, plPFC, PtA, and V1/V2. Detected subgraphs p < 0.001 under permutation test.
(D) Select frontal cortico-claustro-cortical circuits preferentially activated type II neurons compared with type I neurons: ACC > CL > ACC (Wilcoxon rank-sum test: p = 0.027); ACC > CL > plPFC (p < 0.0001); ACC > CL > ilPFC (p = 0.003); ACC > CL > OFC (p = 0.006); ACC > CL > V1/V2 (p = 0.016); ACC > CL > PtA (p = 0.009); plPFC > CL > V1/V2 (p = 0.041); ilPFC > CL > ilPFC (p = 0.016); ilPFC > CL > V1/V2 (p = 0.215); and OFC > CL > RSC (p = 0.015). n = 15 cells in each circuit for each subtype. n = 1,050 cells total. Error bars: standard error of the mean.