Shown from left to right are the schematics and representative images of GUVs 20 minutes after adding buffer only, anti-handle-conjugated PLY, PLY-DNA rings, and DNA-origami rings without PLY. Pseudo-colors: Cyan = Cy3 (DNA ring), red = Cy5 (GUV), and green = FITC (dextran). Scale bars: 5 μm. Bar graphs at the bottom denote the percentages of GUVs with observable dextran influx.