TABLE 6.

Organic osmotic solutes within moderately halophilic bacteria^a

Speciesb	Glycine betaine	Ectoine	Hydroxyectoine	Trehalose	Others
Micrococcus halobius		++	+	+	Glucose, glutamate
“Micrococcus varians subsp. halophilus”		++	+		Glucose, glutamate
Marinococcus halophilus		++	++		Glucose
Marinococcus albus		++	+		Alanine
Planococcus citreus					Proline, N-acetylornithine, N-acetyllysine
Halobacillus halophilus		++			Proline, N-acetyllysine
Salinicoccus roseus*	+				Proline
Salinicoccus hispanicus*	+				Proline
Halomonas halophila		++	+		Glutamate
Halomonas elongata		++	+		Glutamate, glucose
Halomonas halmophila		++	+		Glutamate, alanine
Halomonas halodenitrificans		++			Glutamate, glucose
Halomonas salina		++	+		Glutamate
Halomonas variabilis*	++	++	++	+
Halomonas eurihalina	++	+			Glutamate
Salinivibrio costicola	++				Glutamate
Chromohalobacter marismortui*		+	++
“Pseudomonas halosaccharolytica”		++	+
Bacillus haloalkaliphilus	+	++			N-Acetylornithine
Bacillus halophilus		++			Proline, N-acetylornithine

^aData were derived from references 43, 87, and 306. 

^b++, present in high concentrations; +, present in minor amounts; ∗, yeast extract was present in the medium, enabling the accumulation of glycine betaine.