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The “flip” and “flop” sides of a Holliday junction in the open conformation. The same junction is shown from the opposite sides. The 3′ and 5′ ends of DNA strands are marked. In the center, the 5′-to-3′ direction around the junctions is indicated by arrows. RuvC binds to the “clockwise” side of the junction, whereas RuvA binds to the “counterclockwise” side.
