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Proposed biosynthetic pathway of oxaleimides. The biosynthesis requires HRPKS (PoxF) with PoxM and PoxL to generate a nonproteinogenic amino acid 58, which is incorporated by PKS-NRPKS (PoxE) assembly line. Knoevenagel condensation followed by hydroxylation and a semipinacol-like rearrangement complete the formation of the substituted succinimide.
