Table 1. Systematic nomenclature.
| Clade | Nominal author | Definition | Composition and remarks | ICPN conversion and Regnum code |
|---|---|---|---|---|
| Tapejaroidea | Kellner (2003) | The least inclusive clade containing Tapejara wellnhoferi Kellner, 1989, Quetzalcoatlus northropi Lawson 1975, and Dsungaripterus weii Young 1964. | Includes the sister-taxa Dsungaripteridae and Azhdarchoidea. | This work, [820]. |
| Azhdarchoidea | Unwin (1995) | The least inclusive clade containing Tapejara wellnhoferi Kellner, 1989 and Quetzalcoatlus northropi Lawson 1975. | Includes the sister-taxa Tapejaromorpha and Azhdarchomorpha. | Andres (2021), [355]. |
| Tapejaromorpha | Andres, Clark & Xu (2014) | The most inclusive clade containing Tapejara wellnhoferi Kellner, 1989 but not Azhdarcho lancicollis Nessov 1984. | Includes the sister-taxa Tapejaridae and Thalassodromidae. | Andres (2021), [356]. |
| Thalassodromidae | Witton (2009) | The least inclusive clade containing Thalassodromeus sethi Kellner & Campos 2002 and Tupuxuara longicristatus Kellner & Campos 1988. |
Includes Thalassodromeus, Tupuxuara, and Kariridraco. | Andres (2021), [770]. |
| Tapejaridae | Kellner (1989) | The least inclusive clade containing Tapejara wellnhoferi Kellner, 1989, Sinopterus dongi Wang & Zhou, 2003, and Caupedactylus ybaka Kellner, 2013. | The first registered definition (Andres, 2021) is (unrestrictedly) emended here in order to stabilize the clade’s diagnosis, usage, and content, under the context of the present reference phylogeny. Characterized mainly by downturned rostra and tall rostral crests, it contains Caupedactylia and Eutapejaria. | Andres (2021), [357], unrestrictedly emended here. |
| Caupedactylia | This work. | The most inclusive clade containing Caupedactylus ybaka Kellner, 2013 but not Tapejara wellnhoferi Kellner, 1989. | Includes Caupedactylus and Aymberedactylus. This clade contains tapejarids which share a symphyseal shelf dorsoventrally steep and deep, and a flat dentary fossa. | This work, [821]. |
| Eutapejaria | This work. | The most inclusive clade containing Tapejara wellnhoferi Kellner, 1989 but not Caupedactylus ybaka Kellner, 2013. | This clade contains tapejarids which share a dorsal dentary eminence, encompassing Tapejarinae and Sinopterinae (sensu Andres, 2021). | This work, [822]. |
| Azhdarchomorpha | Pêgas et al. (2021) | The most inclusive clade containing Azhdarcho lancicollis Nessov 1984 but not Thalassodromeus sethi Kellner & Campos 2002 or Tapejara wellnhoferi Kellner, 1989. | Includes Keresdrakon, Chaoyangopteridae, Alanqidae, and Azhdarchidae. | Pêgas et al. (2021), [574]. |
| Chaoyangopteridae | Lü et al. (2008) | The most inclusive clade containing Chaoyangopterus zhangi Wang & Zhou, 2003 but not Quetzalcoatlus northropi Lawson 1975. | Includes Chaoyangopterus, Jidapterus, Shenzhoupterus, and Lacusovagus. | Andres (2021), [368]. |
| Azhdarchiformes | Andres (2021) | The most inclusive clade containing Quetzalcoatlus northropi Lawson 1975 but not Chaoyangopterus zhangi Wang & Zhou, 2003. | Under the present reference phylogeny, the Azhdarchiformes include Alanqidae and Azhdarchidae. | Andres (2021), [771]. |
| Alanqidae | Pêgas et al. (2021) | The most inclusive clade containing Alanqa saharica Ibrahim et al. 2010 but not Chaoyangopterus zhangi Wang & Zhou, 2003 or Azhdarcho lancicollis Nessov 1984. |
Includes Alanqa, Argentinadraco, Xericeps, Leptostomia, and Montanazhdarcho. Characterized by bowed-out lateral jaw margins in cross-section, and possibly by a pair of dentary occlusal ridges. | Pêgas et al. (2021), [576]. |
| Azhdarchidae | Padian (1986) | The least inclusive clade containing Azhdarcho lancicollis Nessov 1984, Phosphatodraco mauritanicus Pereda-Suberbiola et al. 2003, and Quetzalcoatlus northropi Lawson 1975. | Includes Eurazhdarcho, Aralazhdarcho, Phosphatodraco, Wellnhopterus, Zhejiangopterus, Azhdarcho, and Quetzalcoatlinae. Characterized by a vestigial cervical neural spine. | Andres (2021), [371]. Emended by Pêgas et al. (2021). |
Note:
Reference phylogeny: this work.