Plantae Lamiales Scrophulariaceae KhamarHamidOualidiJalal ElTouhamiAmina OuazzaniCiveyrelLaureNomenclature and typification in Verbascum (Scrophulariaceae) from North AfricaPhytoKeys2704202322511515210.3897/phytokeys.225.99356 6B140D66-3975-5BAB-AA90-D02CAD961D41 Verbascum rotundifolium subsp. rotundifolium , in Lunds Univ. Arsskrift, 2n.f. 29 (2): 398. 1933. = Verbascumnumidicum Pomel, in Nouv. Mat. Fl. Atlantique (1): 95. 1874. Type: [Algeria]. Rochers au sommet du Mécid à Constantine 23 Mai 1857, S. Choulette 369 (Lectotype designated here: MPU [MPU004892]!, isolectotype JE [JE00013694]!). ≡ Verbascumrotundifoliumvar.numidicum (Pomel) Murb., in Bull. Soc. Hist. Nat. Afrique N. 18: 83. 1927. [image of lectotype available at https://herbier.umontpellier.fr/zoomify/zoomify.php?fichier=MPU004892] (2) = Verbascumkabylianum Debeaux, in Rev. Bot. Bull. Mens. Soc. Franç. de Botan. 8: 265. 1890. Locality citation in the protologue of Type: [Algeria]. In sylyaticis montium kabylie surper. Prope fort-national, ad viam Taourirt-Amokran, 950 m, 6 Junio 1859, O. Debeaux s.n.: [n. v.] ≡ Verbascumrotundifoliumvar.kabylianum (Debeaux) Murb. in Bull. Soc. Hist. Nat. Afrique N. 18: 83. 1927. = Verbascumrotundifoliumvar.castellorum Maire, in Bull. Soc. Hist. Nat. Afrique N. 29:, 438. 1938. Type: [Morocco]. In rupestribus calcareis Atlantis Medii infra Ksiba, 900 m, 22 Juinio 1936, R. Maire, s.n. (Lectotype designated here: P [P00083085]!; isolectotype: MPU [MPU004005, MPU004006]!, P [P01167373]!). [image of lectotype available at https://plants.jstor.org/stable/viewer/10.5555/al.ap.specimen.mpu004006](3)Notes.

This taxon was established by Murbeck (1933) under the recombination V.rotundifolium subsp. eu-rotundifolium. According to Ferguson (1972) and Benedí (2009), this subspecies is distributed in southern Italy, Sicily, Algeria, and Tunisia. However, for Fennane and Ibn Tattou (2005), Ibn Tattou (2007), and Dobignard and Chatelain (2013), this taxon is spontaneous in Morocco. Under its current recombination, V.rotundifoliumsubsp.rotundifolium it is accepted in the Partical flora of Morocco (Fennane and Ibn Tattou 2005, Ibn Tattou, 2007), in the index of synonyms for the flora of North Africa (Dobignard & Chatelain, 2013), and by the APD (African Plant Database 2022).

(2)Pomel (1874) described on Verbascumnumidicum based on the collections of Choulette exs. [exsiccata] 369, but did not provide collection date nor the name of the herbarium where the original material were housed. After a thorough search, two specimens were found, one at Herb. MPU (MPU004892) and the other at Herb. JE (JE00013694), both of which matched the data provided in the protologue. Thus, the sheet MPU004892 is chosen here as the lectotype of the name V.numidicum, because it is better preserved and shows more diagnostic features described in the protologue.

(3) The protologue of Verbascumrotundifoliumvar.castellorum (Maire 1938) is composed of a Latin diagnosis, and the details of the collection locality on limestone rocks below Ksiba region at an altitude of about 900 m in the Middle Atlas, Morocco, but there is no mention of the name of the herbarium that houses the original material. Four herbarium sheets deposited in Herbs MPU (MPU004005, MPU004006) and P (P01167373, P00083085) were traced. These specimens can be considered as the original material according to the type details provided in the protologue. The specimen P00083085 persevered at Herb. P is better conserved and best represents the diagnostic features of the taxon given in the protologue; hence it is selected here as the lectotype for the name V.rotundifoliumvar.castellorum.

MurbeckSS (1933) Monographie der Gattung Verbascum.Acta Universitatis Lundensis29: 1630.FergusonIK (1972) Verbascum L. In: TutinTGHeywoodVHBurgesNAMooreDMValentineDHWaltersSMWebbDA (Eds) Flora Europaea (Vol 3).Cambridge University Press, 205216.BenedíC (2009) Verbascum L. In: BenedíCRicoEGüemesJHerreroA (Eds) Flora Iberica, vol.XIII. Real Jardín Botánico, Csic, Madrid, 4996.FennaneMIbn TattouM (2005) Flore vasculaire du Maroc inventaire et chorologie 1. Travaux de l’Institut Scientifique.Série Botanique37: 1405.Ibn TattouM (2007) Scrophulariaceae. In: FennaneMIbn TattouMOuyahyaAEl OualidiJ (Eds) Flore Pratique du Maroc (Vol. 2).Travaux de l’Institut Scientifique, Série Botanique38: 503554.DobignardAChatelainC (2013) Index synonymique de la flore d’Afrique du Nord. Dicotyledoneae: OleaceaeZygophyllaceae, (Vol. 5).Conservatoire et Jardin Botaniques Ville de Genève – ECWP, Genève, 451 pp.African Plant Database (APD) (2022) African Plant Database (version 3.4.0). Conservatoire at Jardin Botaniques de la Ville de Genèva and South Africa National Biodiversity Institute, Pretoria. http://www.ville-ge.ch/musinfo/bd/cjb/africa/ [Accessed on 12.12.2022]PomelA (1874) Nouveaux matériaux pour la flore atlantique. Imprimerie V. Aillaud et Cie, Alger, 95.MaireR (1938) Contributions à l'étude de la Flore de l'Afrique du Nord.Bulletin de la Société d'Histoire Naturelle de l'Afrique du Nord29(1): 438.