Acidic residues in the membrane-proximal stalk region of vaccinia virus protein B5 are required for glycosaminoglycan-mediated disruption of the extracellular enveloped virus outer membrane, by Kim L. Roberts, Adrien Breiman, Gemma Carter, Helen Ewles, Michael Hollinshead, Mansun Law and Geoffrey L. Smith

Journal of General Virology vol. 90, part 7, pp. 1582–1591