ValimYeKavanaughDavid H.ShiHongliangLiangHongbinA key to species of subgenus Lithochlaenius (Coleoptera, Carabidae, Chlaeniini, Chlaenius), with descriptions of three new speciesZookeys9920112011128155210.3897/zookeys.128.1804 Chlaenius (Lithochlaenius) propeagilis urn:lsid:zoobank.org:act:0DB721CA-0192-49D6-BD14-A9320D1D574C http://species-id.net/wiki/Chlaenius_(Lithochlaenius)_propeagilis Liu & Kavanaughsp. n.Figs 14–1520–2154688298–99121–122148158Types.

Holotype: male (IZCAS),"Southwest Yunnan, Gongshan, Dulongjiang, 0.5km N of Dizhengdang, 28.08442, 98.32652”/ "1880 m, 2004.10.29, David Kavanaugh Coll.”/ "Holotype Chlaenius (Lithochlaenius) propeagilis Liu & Kavanaugh sp. n.” [red label]. Paratypes: Total 452 specimens (IZCAS, CASC): 1 male, "China, Yunnan Provin. Gongshan County, Cikai town, along street, N27°44'43", E98°39'53"”/ "1500 m, 2006.5.5, Liang H.B, Ba Weidong”; 2 males, "China, Yunnan, Gongshan County, Cikai Township, Nu Jiang at Dashaba, N27.73845, E098.67092”/ "1430 m, 8–9 October 2002, Stop #DHK 2002–40, D.H. Kavanaugh, P.E. Marek & H.B. Liang collectors”; 17 males and 11 females, "China, Yunnan Provin. Gongshan County, Cikai town, Pulahe, N27°46'08", E98°39'12"”/ "1510 m, 2002.9.21–24, Liang Hongbin, Ba Weidong”; 1 male and 1 female, "China, Yunnan, Gongshan, Cikai Township, 3.3 airkm NW of Gongshan above hydropower diversion dam, 1530 m”/ "N27.77175, E098.64924, 24 September 2002, Stop # DHK 2002–028, D.H. Kavanaugh collector”; 2 males, "China, Yunnan Prov. Gongshan, Cikai Town, Pulahe joint of Nujiang, 27.74843N, 98.66498E”/ "1530m, 2004.10.23, D. Kavanaugh, Dong D.Z.”; 6 males and 6 females, "China, Yunnan Provin. Gongshan County, Cilou (Power Station), N27°46'14", E98°39'16"”/ "1510m, 2002.5.6, Liang H.B, Ba Weidong, Yang Guodong, Li X.Q”; 5 males and 3 females, "China, Yunnan Provin. Gongshan County, Cikai town, Gazu Station, N27°44'35", E98°36'17"”/ "1600–1750 m, 2002.5.5, Liang H.B, Ba Weidong”; 2 males, "China, Yunnan Provin. Gongshan County, Cikai town to Qiqi Station, N27.43086, E98.34150”/ "1700–2000m, 2002.4.29, Liang Hongbin, Ba W.D.”; 7 males and 26 females, "China, Yunnan, Gongshan, Cikai Township, 3.0 airkm N of Gongshan on Pula He at hydropower diversion dam, 1500 m”/ "N27.77055°, E098.65446°, 24 September 2002, Stop # DHK 2002–027, D.H. Kavanaugh, P.E. Marek & D.Z. Dong collectors”; 3 males and 4 females (CASC), "CHINA, Yunnan Province, Gaoligong Shan, Nujiang Prefecture, Gongshan County, Qiqi He just above hydroelectric plant, 1500m”/ "N27.75748°, E98.66073°, 22 July 2000, Stop #00–269, D.H. Kavanaugh, Liang H.-B., & Dong D.-Z. collectors”; 2 males and 4 females, "China, Yunnan, Gongshan, Bingzhongluo Township, 34km N of Gongshan at junction of Shuangla He and Nu Jiang, 1550m”/ "E27.96918°, E098.66198°, 25 September 2002, stop # DHK 2002–039, D.H. Kavanaugh, P.E. Marek & D.Z. Dong collectors”; 4 males and 6 females, "China, Yunnan Prov. Gongshan, Bingzhongluo, Shuangla He, beach, 27.96817N, 98.66187E”/ "1520 m, 2004.10.22, D. Kavanaugh collector”; 12 males and 10 females, "China, Yunnan, Gongshan, Bingzhongluo, Shuanglahe, riverside, N27°58'59", E98°39'15"”/ "1588 m, 2002.9.22–26, Liang H.B. Li Xiangqian”; 2 males and 5 females, "China, Yunnan, Gongshan, Bingzhongluo, Shuanglahe, riverside, N27°58'59", E98°39'15"”/ "1588 m, 2002.7.20, Ba Weidong”; 5 males and 2 female, "China, Yunnan Prov. Gongshan, Dulongjiang, 0.5 km N of Dizhengdang, 28.08442N, 98.32652E”/ "1880 m, 2004.10.29; D. Kavanaugh, Dong D.Z.”; 7 males and 4 females, "China, Yunnan Provin. Gongshan County, Dulongjiang, Kongdang, headlamp, 27.87764N, 98.33618E”/ "1510m, 2006.8.27, David Kavanaugh”; 12 males and 17 females, "China, Yunnan Provin. Gongshan, Dulongjiang, Xianjiudang village, 27.94092N, 98.33340E”/ "1580m, 2004.11.4, D. Kavanaugh, Dong D.Z.”;1 male, "China, Yunnan, Fugong County, Shangpa Town, west bank of Nu Jiang, 1185 m, N26.90668, E098.86339”/ "13 October 2002, Stop #DHK 2002–047, D.H. Kavanaugh, P.E. Marek & H.B. Liang, D.Z. Dong collectors”; 5 males and 3 females, "China, Yunnan Prov. Fugong, Shangpa Town, Nujiang, River, 26.90650N, 98.86397E”/ "1175 m, 2004.4.20, Liang H.B. coll.”; 9 males and 4 females, "China, Yunnan Prov. Fugong, Shangpa Town, Beach of Nujiang, 26.90650N, 98.86397E”/ 1175 m, 2005.8.20, Liang H.B. Zhang J.F. Dong D.Z.”; 7 males and 5 females, "China, Yunnan Prov. Fugong, Shangpa Town, Mugujia, riverside, 26.86203N, 98.87142E”/ 1177 m, 2005.8.22, Dong Dazhi collector”; 1 male and 2 females, "China, Yunnan Prov. Fugong, Shangpa, Mugujia, round waterfall, 26.86203N, 98.87142E”/ 1177 m, 2005.8.22, Liang H.B. Zhang J.F.”; 1 female, "China, Yunnan, Fugong, Lumadeng Township, 2km airkm S of Aludi on Nu Jiang, 1245 m, N27.09037, E098.87359”/ "20 September 2002, Stop # DHK 2002–022, D.H. Kavanaugh & H.B. Liang collectors”; 1 female, "China, Yunnan Prov., Fugong, Pihe at junction of Nujiang River, 26.53177N, 98.89753E”/ "1060 m, 2004.4.20, D. Kavanaugh, C. Griswold”; 2 males and 6 females, "China, Yunnan, Lushui, Liuku, west bank of Nu Jiang, 960 m, N25.854, E098.852/ "15 October 2002, stop # DHK 2002–050, D.Z. Dong collector”; 1 male and 2 females, "China, Yunnan Provin. Lushui, Beach of Nujiang River, under stone, N25°51'20", E98°50'58"”/ 800 m, 2002.9.19, Liang Hongbin”; 2 males and 2 females (CASC), "CHINA, Yunnan Province, Gaoligongshan Mountains, Nujiang Prefecture, Lushui County, Salween River, 17 km N of Liuku, 970m”/ "25°58.7'N, 98°50.4'E, 21 October 1998, Stop #98–120, D.H. Kavanaugh & C.-L. Long collectors”; 2 females (CASC), "CHINA, Yunnan Province, Nujiang Prefecture, Lushui County, Liuku Township, Liuku, 800m”/ "25.86010°N, 98.85155°E, 25–26 June 2000, Stop #00–7, D.H. Kavanaugh & Liang H.-B. collectors”; 4 males and 5 females (CASC), "CHINA, Yunnan Province, Nujiang Prefecture, Lushui County, San jiang Township, Nu Jiang, 790m”/ "25.72964°N, 98.87180°E, 26 June 2000, Stop #00–9, D.H. Kavanaugh & Liang H.-B. collectors”; 7 male and 7 females, "China, Yunnan Prov. Baoshan, Longyang, Bawan, Dongfengqiao, 24.98742N, 98.87047E”/ "670 m, 2005.6.1, D. Kavanaugh, Dong D.Z.”; 15 male and 16 females, "China, Yunnan Prov. Baoshan, Longyang, Bawan, Dongfengqiao, 24.98535N, 98.87382E”/ "670 m, 2005.5.29–6.1, Liang H.B. Dong D.Z.”; 1 male, "China, Yunnan Prov. Tengchong, Mangbang, Longwenqiao, 25.02329N, 98.67710E”/ "1290 m, 2006.6.5, Liang H.B. Hu P.”; 6 males and 7 females, "China, Yunnan Prov. Tengchong, Mangbang, Longwenqiao, beach, 25.02396N, 98.67675E”/ "1285 m, 2006.6.5, David Kavanaugh”; 3 males and 1 female, "China, Yunnan Province, Tengchong, Shangying, N25°02'29.7", E98°40'22.9"” / "1335 m, 2003.10.19, Liang H.B, Shi X.C.”; 2 males and 8 females, "China, Yunnan Prov. Tengchong Co., Wuhe Town, Longjiangqiao, 24.89499N, 98.67510E”/ "1205 m, 2005.V.24, Kavanaugh D. Dong D.Z.”; 1 female, "China, Yunnan Province, Tengchong, Wuhe Township, Longjiang Bridge on Longchuanjiang”/ "N24.89889, E098.67667, 1215 m, 28 October 2003, under rocks, Dong D.Z. collector”; 17 males and 9 females, "China, Yunnan Prov. Tengchong, Wuhe, Longjiangqiao, beach, 24.89176N, 98.67551E”/ "1230 m, 2006.6.3, Kavanaugh D. Brett R.”; 4 males and 4 females, "China, Yunnan Prov. Tengchong Co., Wuhe Town, Longjiangqiao, 24.89284N, 98.67439E”/ "1210 m, 2005.V.24, Liang H.B. Yang J.L”; 1 male, "China, Yunnan Prov. Tengchong, Wuhe, Longjiangqiao, beach, 24.89293N, 98.67489E”/ "1220 m, 2006.6.3, Liang H.B. Hu P.”; 3 males, "China, Yunnan Province, Tengchong Co. Qushi Township, Shuang He Cun, N25.32555, E098.60861”/ "1464 m, 21 October 2003, under rocks, Dong D.Z. collector”; 2 males, "China, Yunnan Prov. Tengchong, Qushi, Xiangyangqiao, beach, 25.21221N, 98.57836E”/ "1500 m, 2006.V.24, David Kavanaugh”; 2 females, "China, Yunnan Prov. Tengchong, Qushi, Xiangyangqiao, beach, 25.23939N, 98.62723E”/ "1440 m, 2006.V.24, David Kavanaugh”; 11 males and 14 females (CASC), "CHINA, Yunnan Province, Gaoligongshan Mountains, Baoshan Prefecture, Tengchong County, Longchuan River at Longkou village”/ ”25°16.9'N, 98°35.5'E, 1500 m, 25 October 1998, Stop #98–126, D.H. Kavanaugh, C.E. Griswold, & C.-L. Long collectors”; 6 males and 1 female, "China, Yunnan Prov. Tengchong, Qushi, Longkou, beach, 25.28175N, 98.59246E”/ "1500 m, 2006.6.6, David Kavanaugh”; 5 males and 1 female (CASC), "CHINA, Yunnan Province, Gaoligongshan Mountains, Baoshan Prefecture, Tengchong County, Longchuan River at Longkou village”/ "25°16.9'N, 98°35.5'E, 1500 m, 2 November 1998, Stop #98–128, D.H. Kavanaugh, C.E. Griswold, C.-L. Long, R. Li, & H.-X. He collectors”; 3 males and 5 females, "China, Yunnan Prov. Tengchong, Qushi, Qinqiao, 25.27236N, 98.60093E”/ "1460 m, 2006.6.2–6, David Kavanaugh, Brett R.”; 4 males and 4 females, "China, Yunnan Province, Tengchong, Zhoujiapo Village, N25.33222, E098.67611”/ "1740 m, 24 October 2003, under rocks Dong D.Z. collector”; 3 males, "China, Yunnan Prov. Tengchong, Jietou, Yonganqiao, beach, 25.32502N, 98.70459E”/ "1470 m, 2006.V.24, Liang H.B.”; 2 males and 1 female, "China, Yunnan Prov. Tengchong, Jietou, Yonganqiao, beach, 25.32504N, 98.60959E”/ "1470 m, 2006.5.24, Kavanaugh D. Brett R.”; 1 male, "China, Yunnan Prov. Tengchong, Hehua, Dengma, on beach, 24.92346N, 98.38612E”/ "1105 m, 2006.6.2, Kavanaugh D. Brett R.”; 3 males and 1 female, "China, Yunnan Province, Jingping Co., Mengla Town, Mengla-daqiao, River side 22˚39'45.7",103˚04'44.7"”/ "312m, 2003.12.15, day, Liang H B, Boris Kataev Colls.”; 12 males and 10 females (IZCAS), "Yunnan, Jingdong, 1100m”/ "1982.IV.29–V.2, Yu P.Y.& Liao S.B.”; 1 female, "Yunnan, Honghe Prefecture, Lvchun, Huanglianshan, 2009.V. 16, Bai X. X. coll.” 2 males and 1 female, "Yunnan, Menglun, Xishuanbanna Botanical Garden”, "2005.5.22, light trap, Zheng Guo leg.”; 3 males and 5 females, "China, Sichuan, Yajiang, Hekou Town, Shabeihou, Yalongjiang, N30.00020, E101.01526”/ "2583m, 2009.5.27–28, Liang H.B.”; 1 female (IZCAS), "China, Tibet, Bomi Yi"ong, Tangmai bridge, Beach of Yi"ong Zangbo, 30.09633N, 95.06577E”/ "2035m, 2006.8.30 N, Liang H.B., Song Z.S.”. Each paratype with an additional yellow label: "Paratype Chlaenius (Lithochlaenius) propeagilis Liu & Kavanaugh sp. n.”

Diagnosis.

Antennomere 1 elongate ovoid (Fig. 54); color of antennomeres 1–3 lighter than antennomeres 4–11; intervals 1–5 almost glabrous medially, with one or two rows of pubescence at each lateral side, more evenly pubescent near apex (Fig. 82); apical lamella of aedeagus moderately triangular (Fig. 121), thick and reflexed in lateral view (Fig. 122).

Description.

Total length = 15.50–17.00 mm (mean = 15.71), width = 5.60–6.13 mm (mean = 5.81); HW = 2.75–2.95 mm (mean = 2.82), EYL = 1.00–1.15 mm (mean = 1.09), ratio Ant3/Ant1 = 1.73–1.93 (mean = 1.81), PL/PW = 0.78–0.85 (mean = 0.83), EL/EW = 1.62–1.79 (mean = 1.73), EL/ PL = 1.29–1.45 (mean = 1.36).

Head and pronotum black with green or coppery metallic luster; elytra black; ventral surface black, with some slight metallic reflection; coxae almost black; mandibles dark brown; antennomeres 4–11, palpomere, trochanters, tibiae at both ends, and tarsomeres reddish or brown; antennomeres 1–3, femora and middle portions of tibiae yellow or yellowish-brown.

Head rugose near eyes and occiput, vertex sparsely rugose; eyes prominent; antennomere 1 elongate–ovoid (Fig. 54); labrum concave at apex, with distinct microsculpture; mentum tooth emarginate apically.

Pronotum cordate, moderately convex; lateral margins bordered; disk almost smooth, sparsely rugose and setose near base and lateral margins; basal foveae moderately deep.

Elytra widest at mid-length; humeral angle obtuse (Fig. 68); striae deep, punctate; intervals moderately convex, intervals 1–5 almost glabrous medially, with a row of pubescence laterally, intervals more evenly pubescent near apex (Fig. 82); intervals 6–9 pubescent throughout and with intervals 7–9 more densely pubescent than 6.

Abdominal sterna bordered, densely pubescent laterally; sterna IV–VI sparsely pubescent medially; sternum VII rugose–pubescent, with apex narrowly rounded or subobtuse in males (Fig. 98), more broadly rounded in female (Fig. 99).

Apical lamella of aedeagus moderately triangular (Fig. 121), thick and bent ventrally (Fig. 122). Gonostylus (Figs 14–15).

Etymology.

The Latinized namepropeagilis refers to the similarity of members of this species to those of Chlaenius agilis.

Geographical distribution.

Fig. 158. Known only from southeastern Tibet and Yunnan Province, China.

Remarks.

Mensural data cited in the description were based on measurements obtained from 5 males and 5 females selected for maximum variation.

Specimens of Chlaenius (Lithochlaenius) collected from western Yunnan were initially determined as Chlaenius agilis Chaudoir. However, after comparison of the male genitalia with those of type specimens of Chlaenius agilis, we are convinced that they represent a distinct new species. Males differ from those of Chlaenius agilis in having the median lobe of aedeagus gradually bent near the base in lateral view (Fig. 122), whereas Chlaenius agilis males have the median lobe abruptly bent with a depression near the base (Fig. 124).

This species is clearly very closely related to Chlaenius agilis. At present, the known ranges of these two species are broadly disjunct (Fig. 158). We know of no locality records for any Chlaenius (Lithochlaenius) species from the intervening area (i.e., between northcentral India and southeastern Tibet and western Yunnan Province. Whether this distributional gap represents a real disjunction or is only an artifact of inadequate collecting in the area to date can only be determined from additional sampling efforts in the region. It would be particularly informative to determine whether or not any populations representing the propeagilis/agilis lineage occur in the region and, if so, whether or not males display intermediate genitalic traits.

Character states of Chlaenius (Lithochlaenius) spp. Figs 1–13 Chlaenius rambouseki Lutshnik1 pronotum2 site of hind angle seta3 elytral base showing the site of basal pore4 abdominal sterna showing the pubescence5 male anterior tarsomere 4 in dorsal view6 male anterior tarsomere 4 in ventral view7 female anterior tarsomere 4 in dorsal view8 female anterior tarsomere 4 in ventral view9 antennomeres 1–5 showing antennomere 3 distinctly longer than 1 and 2 ones combined10 metepisternum11 aedeagus, showing the basic structure of the aedeagus in Lithochlaenius species12 female gonostylus in ventral view13 female gonostylus in lateral view14 female gonostylus of Chlaenius propeagilis sp. n. in ventral view15 female gonostylus of Chlaenius propeagilis sp. n. in lateral view.

Habitus of Chlaenius (Lithochlaenius) spp16 Chlaenius chuanqianensis sp. n., holotype, male, dorsal view17 Chlaenius chuanqianensis sp. n., holotype, male, ventral view18 Chlaenius linwensini sp. n., holotype, male, dorsal view19 Chlaenius linwensini sp.n., holotype, male, ventral view20 Chlaenius propeagilis sp. n., holotype, male, dorsal view21 Chlaenius propeagilis sp. n., holotype, male, ventral view. Scale line = 5.0 mm.

Antennomere 1 of Chlaenius (Lithochlaenius) species52 Chlaenius chuanqianensis sp. n., holotype53 Chlaenius linwensini sp. n., holotype54 Chlaenius propeagilis sp. n., holotype55 Chlaenius agilis Chaudoir, holotype56 Chlaenius formosensis Lorenz, CCCC57 Chlaenius leishanensis Kirschenhofer, IZCAS58 Chlaenius noguchii Bates, IZCAS59 Chlaenius wrasei Kirschenhofer, IZCAS60 Chlaenius agiloides Jedlička, IZCAS61 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS62 Chlaenius anchomenoides Bates, paratype63 Chlaenius nuristanus Jedlička, paratype64 Chlaenius nuristanus a. rubridipes Jedlička, paratype65 Chlaenius rambouseki Lutshnik from Taiwan,CCCC. Scale line = 1.0 mm.

Humeral region of elytra of Chlaenius (Lithochlaenius) spp66 Chlaenius chuanqianensis sp. n., holotype67 Chlaenius linwensini sp. n., holotype68 Chlaenius propeagilis sp. n., holotype69 Chlaenius agilis Chaudoir, holotype70 Chlaenius formosensis Lorenz, CCCC71 Chlaenius leishanensis Kirschenhofer, IZCAS72 Chlaenius noguchii Bates, IZCAS73 Chlaenius wrasei Kirschenhofer, IZCAS74 Chlaenius agiloides Jedlička, IZCAS75 Chlaenius rambouseki Lutshnik from Ussuri region, ZCAS76 Chlaenius anchomenoides Bates, paratype77 Chlaenius nuristanus Jedlička, paratype78 Chlaenius nuristanus Jedlička, paratype79 Chlaenius rambousekiLutshnik from Taiwan, CCCC. Scale line = 1.0 mm.

Elytral intervals of Chlaenius (Lithochlaenius) spp. 80 Chlaenius chuanqianensis sp. n., holotype81 Chlaenius linwensini sp. n., holotype82 Chlaenius propeagilis sp. n., holotype83 Chlaenius agilis Chaudoir, holotype84 Chlaenius formosensis Lorenz, CCCC85 Chlaenius leishanensis, IZCAS86 Chlaenius noguchii Bates, IZCAS87 Chlaenius wrasei Kirschenhofer, IZCAS88 Chlaenius agiloides Jedlička, IZCAS89 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS90 Chlaenius anchomenoides Bates, paratype91 Chlaenius nuristanusJedlička, paratype92 Chlaenius nuristanus Jedlička, paratype93 Chlaenius rambouseki Lutshnik from Taiwan,CCCC. Scale line = 1.0 mm.

Abdominal sternum VII (ventral aspect) of Chlaenius (Lithochlaenius) species94 Chlaenius chuanqianensis sp. n., male, holotype95 Chlaenius chuanqianensis sp. n., female, paratype96 Chlaenius linwensini sp. n., male, holotype97 Chlaenius linwensini sp. n., female, paratype98 Chlaenius propeagilis sp. n., male, holotype99 Chlaenius propeagilis sp. n., female, paratype100 Chlaenius agilis Chaudoir, male, holotype101 Chlaenius formosensis Lorenz, male102 Chlaenius formosensis Lorenz, female103 Chlaenius leishanensis Kirschenhofer, male104 Chlaenius leishanensis Kirschenhofer, female105 Chlaenius noguchii Bates, male106 Chlaenius noguchii Bates, female107 Chlaenius wrasei Kirschenhofer, male108 Chlaenius wrasei Kirschenhofer, female109 Chlaenius agiloides Jedlička, male110 Chlaenius agiloides Jedlička, female111 Chlaenius rambouseki Lutshnik, from Ussuri region, male112 Chlaenius rambouseki Lutshnik from Ussuri region, female113 Chlaenius anchomenoides Bates, male, paratype114 Chlaenius nuristanus Jedlička, male, paratype115 Chlaenius nuristanus Jedlička, female, paratype116 Chlaenius formosanus Jedlička from Taiwan, male. Scale line = 1.0 mm.

Aedeagi of Chlaenius (Lithochlaenius) species117 Chlaenius chuanqianensis sp. n., holotype (dorsal view)118 Chlaenius chuanqianensis sp. n., holotype (left lateral view)119 Chlaenius linwensini sp. n., holotype (dorsal view)120 Chlaenius linwensini sp. n., holotype (left lateral view)121 Chlaenius propeagilis sp. n., holotype (dorsal view)122 Chlaenius propeagilis sp. n., holotype (left lateral view)123 Chlaenius agilis Chaudoir, holotype (dorsal view)124 Chlaenius agilis Chaudoir, holotype (left lateral view)125 Chlaenius formosensis Lorenz, holotype (dorsal view)126 Chlaenius formosensis Lorenz, holotype (left lateral view)127 Chlaenius leishanensis Kirschenhofer (dorsal view)128 Chlaenius leishanensis Kirschenhofer (left lateral view)129 Chlaenius noguchiiBates (dorsal view)130 Chlaenius noguchiiBates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130).

Photographs of habitats for Chlaenius (Lithochlaenius) spp. 146 Chlaenius chuanqianensis sp. n., Dabaitang, Xishui county, North Guizhou, China147 Chlaenius anchomenoides Bates, Goorais valley, Pakistan (Provided by Dr. Muhammad Abbas in Pakistan Museum of Natural History)148 Chlaenius propeagilis sp. n., Gaoligongshan, Yunnan, China149 Chlaenius leishanensis Kirschenhofer,Xiaodanjiang, Leigongshan, Leishan county, Southeast Guizhou, China150Chlaenius chuanqianensis sp. n. and Chlaenius agiloides Jedlička, Jinshajiang, Sichuan151 Chlaenius rambouseki Lutshnik, Ussri river, Heilongjiang, China.

Map showing the known geographical distributions of Chlaenius (Lithochlaenius)spp.