ValimYeKavanaughDavid H.ShiHongliangLiangHongbinA key to species of subgenus Lithochlaenius (Coleoptera, Carabidae, Chlaeniini, Chlaenius), with descriptions of three new speciesZookeys9920112011128155210.3897/zookeys.128.1804 Chlaenius (Lithochlaenius) noguchii http://species-id.net/wiki/Chlaenius_(Lithochlaenius)_noguchii BatesFigs 28–29587286105–106129–130158Chlaenius noguchii Bates, 1873:251; Chaudoir, 1876:192; Kryzhanovskij, 1976:12; Kirschenhofer, 1997:116; Kirschenhofer, 2005:491Specimens examined.

Total 47 specimens: Japan: 10 males and 11 females (IZCAS), "Shiga, Takashima, Biwa Lake, 2008.3.30, Liang H.B. coll.”; 1 male and 1 female (IZCAS), "Yadorigi-zawa, Tanzawa, Kanagawa”; 1 male (IZCAS), "Sado-shima Island”/ "1968.VIII.20”; 2 males and 2 females (CASC), "Harima Japan May 1916”; 1 female (CASC), "Hozukyo Kyoto 1951.9.21”; 4 males and 2 females (CASC); "Kobe V-1912”/ "L. Gressitt Collector”; 1 female (CASC), "Kyoto, Yase, Oct. 21, 1951, Col. T. Horio”; 2 females (CASC), "Mimasaka Japan VII. J. E. Lewis”; 2 males and 2 females (CASC), "Mimasaka Japan VII-1912”/ "Coll. by J. C. Thompson”; 1 male and 1 female (CASC), " Nikko Japan VII.30.23”/ "ECVan Dyke collector”; 2 females (CASC), "Nikko Hondo Japan VII-1912”/ "Coll. by J. C. Thompson”; 1 female (CAS), Tokyo, Japan VI-6-31.”/ "L. Gressitt Collector”.

Diagnosis.

Antennomere 1 coniform (Fig. 58); intervals 1–7 strongly convex, carinate, glabrous medially (Fig. 86).

Description.

Total length = 14.50–16.00 mm (mean = 15.33), width = 5.07–5.60 mm (mean = 5.33); HW = 2.50–2.90 mm (mean = 2.67), EYL = 1.00–1.10 mm (mean = 1.02), ratio Ant3/Ant1 = 1.56–1.88 (mean = 1.72), PL/PW = 0.83–0.90 (mean = 0.86), EL/EW = 1.75–1.91 (mean = 1.80), EL/ PL = 1.29–1.40 (mean = 1.35).

Head and pronotum black with strong green or bluish green metallic luster; elytra black with very weak blue luster; ventral surface black; legs, antennae, mandibles and palpomere dark brown.

Head with vertex coarsely punctate behind eyes; antennomere 1 coniform (Fig. 58); clypeus and labrum nearly truncate at apex; mentum tooth emarginate. Pronotum with disk finely punctate, nearly glabrous; basal foveae deep, finely pubescent and punctate (punctations much smaller than those on vertex). Elytra with intervals strongly convex, intervals 1–7 carinate, smooth and glabrous medially, with a row of punctation laterally (Fig. 86), intervals 8–9 flat, densely pubescent and punctate; striae deep, punctate; humeral angle obtuse (Fig. 72). Abdominal sterna IV-VI sparsely pubescent medially; apex of sternum VII more distinctly truncate in female than in male (Figs 105–106). Aedeagus slender, flattened (Fig. 130), lamella broadened to the left side, asymmetric (Fig. 129), thin and bent ventrally (Fig. 130).

Geographical distribution.

Fig. 158. Confirmed only from Japan.

Remarks.

Mensural data cited in the description were based on measurements obtained from 5 males and 5 females selected for maximum variation.

This species may well be endemic to Japan, and its reported occurrence in Korea (Kwon and Lee 1986) and Vietnam (Paik et al. 2006) requires further clarification. The slender aedeagus of males of this species is very different from the stout aedeagi of other species of the agilis group.

Chlaenius (Lithochlaenius) spp28 Chlaenius noguchii Bates, male, in IZCAS, dorsal view29 Chlaenius noguchii Bates, male, in IZCAS, ventral view30Chlaenius wrasei Kirschenhofer, male, in IZCAS, dorsal view31 Chlaenius wrasei Kirschenhofer, male, in IZCAS, ventral view32 Chlaenius agiloides Jedlička, male, in IZCAS, dorsal view33 Chlaenius agiloides Jedlička, male, in IZCAS, ventral view. Scale line = 5.0 mm.

Antennomere 1 of Chlaenius (Lithochlaenius) species52 Chlaenius chuanqianensis sp. n., holotype53 Chlaenius linwensini sp. n., holotype54 Chlaenius propeagilis sp. n., holotype55 Chlaenius agilis Chaudoir, holotype56 Chlaenius formosensis Lorenz, CCCC57 Chlaenius leishanensis Kirschenhofer, IZCAS58 Chlaenius noguchii Bates, IZCAS59 Chlaenius wrasei Kirschenhofer, IZCAS60 Chlaenius agiloides Jedlička, IZCAS61 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS62 Chlaenius anchomenoides Bates, paratype63 Chlaenius nuristanus Jedlička, paratype64 Chlaenius nuristanus a. rubridipes Jedlička, paratype65 Chlaenius rambouseki Lutshnik from Taiwan,CCCC. Scale line = 1.0 mm.

Humeral region of elytra of Chlaenius (Lithochlaenius) spp66 Chlaenius chuanqianensis sp. n., holotype67 Chlaenius linwensini sp. n., holotype68 Chlaenius propeagilis sp. n., holotype69 Chlaenius agilis Chaudoir, holotype70 Chlaenius formosensis Lorenz, CCCC71 Chlaenius leishanensis Kirschenhofer, IZCAS72 Chlaenius noguchii Bates, IZCAS73 Chlaenius wrasei Kirschenhofer, IZCAS74 Chlaenius agiloides Jedlička, IZCAS75 Chlaenius rambouseki Lutshnik from Ussuri region, ZCAS76 Chlaenius anchomenoides Bates, paratype77 Chlaenius nuristanus Jedlička, paratype78 Chlaenius nuristanus Jedlička, paratype79 Chlaenius rambousekiLutshnik from Taiwan, CCCC. Scale line = 1.0 mm.

Elytral intervals of Chlaenius (Lithochlaenius) spp. 80 Chlaenius chuanqianensis sp. n., holotype81 Chlaenius linwensini sp. n., holotype82 Chlaenius propeagilis sp. n., holotype83 Chlaenius agilis Chaudoir, holotype84 Chlaenius formosensis Lorenz, CCCC85 Chlaenius leishanensis, IZCAS86 Chlaenius noguchii Bates, IZCAS87 Chlaenius wrasei Kirschenhofer, IZCAS88 Chlaenius agiloides Jedlička, IZCAS89 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS90 Chlaenius anchomenoides Bates, paratype91 Chlaenius nuristanusJedlička, paratype92 Chlaenius nuristanus Jedlička, paratype93 Chlaenius rambouseki Lutshnik from Taiwan,CCCC. Scale line = 1.0 mm.

Abdominal sternum VII (ventral aspect) of Chlaenius (Lithochlaenius) species94 Chlaenius chuanqianensis sp. n., male, holotype95 Chlaenius chuanqianensis sp. n., female, paratype96 Chlaenius linwensini sp. n., male, holotype97 Chlaenius linwensini sp. n., female, paratype98 Chlaenius propeagilis sp. n., male, holotype99 Chlaenius propeagilis sp. n., female, paratype100 Chlaenius agilis Chaudoir, male, holotype101 Chlaenius formosensis Lorenz, male102 Chlaenius formosensis Lorenz, female103 Chlaenius leishanensis Kirschenhofer, male104 Chlaenius leishanensis Kirschenhofer, female105 Chlaenius noguchii Bates, male106 Chlaenius noguchii Bates, female107 Chlaenius wrasei Kirschenhofer, male108 Chlaenius wrasei Kirschenhofer, female109 Chlaenius agiloides Jedlička, male110 Chlaenius agiloides Jedlička, female111 Chlaenius rambouseki Lutshnik, from Ussuri region, male112 Chlaenius rambouseki Lutshnik from Ussuri region, female113 Chlaenius anchomenoides Bates, male, paratype114 Chlaenius nuristanus Jedlička, male, paratype115 Chlaenius nuristanus Jedlička, female, paratype116 Chlaenius formosanus Jedlička from Taiwan, male. Scale line = 1.0 mm.

Aedeagi of Chlaenius (Lithochlaenius) species117 Chlaenius chuanqianensis sp. n., holotype (dorsal view)118 Chlaenius chuanqianensis sp. n., holotype (left lateral view)119 Chlaenius linwensini sp. n., holotype (dorsal view)120 Chlaenius linwensini sp. n., holotype (left lateral view)121 Chlaenius propeagilis sp. n., holotype (dorsal view)122 Chlaenius propeagilis sp. n., holotype (left lateral view)123 Chlaenius agilis Chaudoir, holotype (dorsal view)124 Chlaenius agilis Chaudoir, holotype (left lateral view)125 Chlaenius formosensis Lorenz, holotype (dorsal view)126 Chlaenius formosensis Lorenz, holotype (left lateral view)127 Chlaenius leishanensis Kirschenhofer (dorsal view)128 Chlaenius leishanensis Kirschenhofer (left lateral view)129 Chlaenius noguchiiBates (dorsal view)130 Chlaenius noguchiiBates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130).

Map showing the known geographical distributions of Chlaenius (Lithochlaenius)spp.

KwonYJLeeSM (1986) Check list of superfamily Caraboidea from Korea (Coleoptera).Insecta Koreana 6: 1-56.PaikJKTracDHWillK (2006) Carabidae from Vietnam (Coleoptera).Journal of Asia-Pacific Entomology 9: 85-105.