LikhitrakarnNatdanaiGolovatchSergei I.PanhaSomsakRevision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae) Zookeys29920112011131116110.3897/zookeys.131.1921 Orthomorpha subelevata urn:lsid:zoobank.org:act:75C015CB-D0D4-4745-8B4D-4FF1901F0892 http://species-id.net/wiki/Orthomorpha_subelevata sp. n.Figs 103105Holotype.

♂ (CUMZ), Malaysia, Kedah State, Gunung Baling, 6°06'N, 101°29'E, 27.02.2008, leg. C. Sutcharit.

Paratypes.

2 ♀ (CUMZ), same data, together with holotype. 1 ♂ (CUMZ), Malaysia, Perlis State, Kaki Bukit, 7°04'32"N, 100°20'55"E, 28.03.2011, leg. R. Chanabun. 1 ♂, 1 ♀ (CUMZ), Penang State, Penang Dam, 5°41'08"N, 100°28'65"E, 26.03.2009, leg. C. Sutcharit. 1 ♂, 1 ♀, 1 juv. (CUMZ), Perak State, Gunung Genting, 4°35'58"N, 102°20'04"E, 30.03.2009, leg. C. Sutcharit.

Name.

To emphasize the strong similarity to Orthomorpha elevata sp. n.

Diagnosis.

Comes closest to Orthomorpha elevata sp. n., but differs in the paraterga lying below the dorsum, coupled with pleurosternal carinae present at least as a caudal denticle until segment 16 (♂) or 13 (♀) (see also Key below).

Description.

Length 40–42 (♂) or 41–45 mm (♀), width of midbody pro- and metazona 3.0–3.7 and 4.0–5.6 mm (♂), 3.0–4.2 and 4.9–6.1 mm (♀), respectively. Holotype 3.95 mm long, 3.9 and 4.0 mm wide on midbody pro- and metazona, respectively.

Coloration of alcohol material after long-term preservation brown to dark brown with contrasting whitish calluses, lateral parts of paraterga close to calluses and, sometimes, entire epiproct, as well as light yellow to yellow-brown legs and epiproct; two paramedian spots divided by a narrow, brown or grey-brown axial stripe on prozona and central parts of metaterga slightly lighter than background, light brown, except for an infuscate cross composed of transverse sulcus and axial line, the latter often extending from collum to tip of epiproct (Fig. 103A-G).

All other characters as in Orthomorpha elevata sp. n., except as follows.

Antennae barely reaching end of segment 3 (♂) or 2 (♀) dorsally. Ribs at caudal margin of metaterga often vague, sometimes represented by low bosses. Axial line always visible on collum, as well as on following prozona, metaterga and epiproct. Paraterga very strongly developed (Fig. 103A-G), but always lying below dorsum. Transverse sulcus incomplete on metatergum 4, complete, but not reaching bases of paraterga on metaterga 5–18. Hypoproct (Fig. 103G) subtrapeziform, caudal margin with two stronger and well-separated setiferous knobs.

Sterna densely setose. Legs long and slender, slightly incrassate in ♂, midbody ones ca 1.7–1.8 (♂) or 1.2–1.3 times (♀) as long as body height; ♂ tarsal brushes visible on legs 1–8, thereafter absent.

Gonopods (Figs 104 & 105) virtually same as in Orthomorpha elevata sp. n.

Orthomorpha subelevata sp. n., ♂ holotype. A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively D, E, F posterior part of body, lateral, dorsal and ventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively.

Orthomorpha subelevata sp. n., ♂ holotype. A, B right gonopod, mesal and lateral views, respectively C–F distal part of right gonopod, mesal, lateral, suboral and subcaudal views, respectively. Scale bar: 0.2 mm.

Orthomorpha subelevata sp. n., ♂ holotype. A, B left gonopod, mesal and lateral views, respectively.Scale bar: 0.2 mm.

Remark.

This new species is rather widespread in northern Malaysia (Map 2).

Distributions of the new species of Orthomorpha. Crossed circle: Orthomorpha picturata sp. n.; Asterisk: Orthomorpha tuberculifera sp. n.; Inverted open triangle: Orthomorpha subtuberculifera sp. n.; Open diamond: Orthomorpha communis sp. n.; Crossed square: Orthomorpha atypica sp. n.; Filled triangle: Orthomorpha latiterga sp. n.; Open square: Orthomorpha suberecta sp. n.; Open circle: Orthomorpha isarankurai sp. n.; Open triangle: Orthomorpha elevata sp. n.; Filled circle: Orthomorpha subelevata sp. n.; Filled square: Orthomorpha similanensis sp. n.; Inverted filled triangle: Orthomorpha spiniformis sp. n.