JamesonMary LizDrumontAlainAroid scarabs in the genus Peltonotus Burmeister (Coleoptera, Scarabaeidae, Dynastinae): key to species and new distributional dataZookeys31720132013320639510.3897/zookeys.320.5352 Peltonotus morio http://species-id.net/wiki/Peltonotus_morio Burmeister, 1847Figs 12, 24, 65, 80, 92Diagnosis (male and female).

Length 14.0–18.0 mm, color overall black or castaneous, elytra black or castaneous and shining (Fig. 12), head with unisetigerous punctures, labrum weakly sinuate (Fig. 24), mentum quadrate in apical half, labial palpomere 2 not enlarged and not dorsoventrally flattened, mala lacking lamellate setal brush, maxillary stipes without setae curled at apices, male protibia tridentate with basal tooth weakly developed, form of male parameres (Fig. 65), female epipleuron weakly, quadrately incised (Fig. 80) and with moderately developed dorsal pillow.

Distribution

(Fig. 92). Bhutan, India (northeastern), Myanmar, Nepal, Thailand, Vietnam.

Peltonotus species dorsal habitus. 1 Peltonotus animus, male 2–3 Peltonotus cybele, male and female (respectively) 4 Peltonotus favonius, male 5–7 Peltonotus fujiokai, males (showing variation) 8 Peltonotus fujiokai, female 9 Peltonotus karubei, male 10–11 Peltonotus malayensis, male and female (respectively) 12 Peltonotus morio, male.

Head (dorsal view) showing characters of the labrum, mandible, and clypeus. 20 Peltonotus karubei (apex of labrum deeply bi-lobed; apex of mandible rounded laterally) 21 Peltonotus malayensis (apex of labrum bi-emarginate; apex of mandible rounded laterally) 22 Peltonotus vittatus (apex of labrum bi-emarginate; apex of mandible rounded laterally) 23 Peltonotus nasutus (apex of labrum weakly sinuate; apex of mandible quadrate laterally with broadly truncate apex; apex of clypeus with weak tubercle in male) 24 Peltonotus morio (apex of labrum weakly sinuate; apex of mandible quadrate laterally with broadly truncate apex; apex of clypeus without tubercle in male).

Male parameres (with or without phallobase), dorsal and lateral views, in Peltonotus. Male parameres are highly asymmetrical, and we illustrate the lateral view that best assists in identification. 61 Peltonotus fujiokai 62 Peltonotus gracilipodus and Peltonotus podocrassus 63 Peltonotus karubei 64 Peltonotus malayensis 65 Peltonotus morio 66 Peltonotus nasutus.

Female elytral epipleuron (gray, ventral view) and position relative to metacoxa in Peltonotus. 73 Peltonotus brunnipennis 74 Peltonotus cybele 75 Peltonotus favonius 76 Peltonotus fujiokai 77 Peltonotus gracilipodus, Peltonotus podocrassus and Peltonotus silvanus 78 Peltonotus kyojinus 79 Peltonotus malayensis 80 Peltonotus morio 81 Peltonotus mushiyaus 82 Peltonotus nasutus 83 Peltonotus nethis 84 Peltonotus pruinosus 85 Peltonotus rubripennis 86 Peltonotus similis 87 Peltonotus sisyrus

Distribution of Peltonotus morio (green icon) and Peltonotus nasutus (blue icon) in southeast Asia. Icons with stars indicate new country records for each species. Map was generated using data in Table 1.

Peltonotus Locality Table. Locality information for Peltonotus morio and Peltonotus nasutus. The Appendix file can be used for dynamic mapping using EarthPoint and GoogleEarth.

Latitude, LongitudeSpecies nameCollection or ReferenceLocality Information
16°40'27"N, 98°17'59"EPeltonotus morioFUJIS. Burma, Mt. Dawna, V.1992, 1 male, ele. 763m, NEW COUNTRY RECORD
12°05'N, 99°00'EPeltonotus morioFUJIS. Burma, Tenasserim, V.1992, 1 female, NEW COUNTRY RECORD
26°52'41"N, 88°17'25"EPeltonotus morioBMNHIndia, Kurseong Div., Lat Panchar, 4000 ft., VI.1934, 6 specimens, Col. Champion
27°39'N, 84°19'EPeltonotus morioDhoj et al. 2009Nepal, Chitwan Central region, Gunjanagar, 230 m
27°39'N, 84°21'EPeltonotus morioDhoj et al. 2009Nepal, Rampur, 230 m, amid maize-maize-vegetables in sandy soil from farming sites.
22°29'N, 103°57'EPeltonotus morioIRSBNVietnam, Lao Cai Prov., VI.10.1917, 1 male NEW COUNTRY RECORD
18°49'16"N, 98°55'11"EPeltonotus morioJameson and Wada 2004Thailand, Doi Suthep
27°18'42"N, 88°35'57"EPeltonotus morioJameson and Wada 2004Sikkim, India
24°39'32"N, 93°54'22"EPeltonotus morioJameson and Wada 2004India, Manipur
25°22'05"N, 91°45'13"EPeltonotus morioJameson and Wada 2004India, Meghalaya, Khasi Hills
27°09'33"N, 88°36'56"EPeltonotus morioJameson and Wada 2004India, Pedong
27°02'09"N, 88°14'08"EPeltonotus morioJameson and Wada 2004India, Darjeeling
28°16'N, 84°05'EPeltonotus morioJameson and Wada 2004Nepal, Chhachok
14°48'00"N, 106°49'59"EPeltonotus nasutusFUJIS. Laos, Attapu, V.13.2007, 1 male, 1 female, ele 450m
14°88'N, 105°87'EPeltonotus nasutusFUJIS. Laos, Champasak Province, 2 females,
16°42'18"N, 98°20'44"EPeltonotus nasutusFUJIS. Burma, Mt. Dawna, V.1992, 1 female
18°38'31"N, 94°42'56"EPeltonotus nasutusFUJIMyanmar, Arakan Province, Nianjyo, 1070m, 1 male, 1 female
15°N, 98°32'EPeltonotus nasutusBMNHW. Thailand, Kanchanaburi Prov., Thung Yai Wildlife Sanctuary, mixed riverside forest, M. Brendell, V.8.1988, 10 specimens, within spathe of Amorphophallus inflorescence
19°25'N, 103°30'EPeltonotus nasutusBMNHLaos, Xiankhouang Prov. V.18.1919, 1 male
26°51'22"N, 104°13'59"EPeltonotus nasutusDrumontChine, Guizhou, Mt. Ping-Qing-Liang-Zi, Weining county, 1-10/VII-2009, 1 male, 3 female NEW COUNTRY RECORD
23°28'5"N, 100°41'EPeltonotus nasutusDrumontChine, Yunnan, Mt. Longtanshan, Jinggu county, VI.11-20, 3 male, Col. Li Jingke NEW COUNTRY RECORD
22°35'N, 99°33'EPeltonotus nasutusDrumontChine, Yunnan, Mt. Daheishan, Menglian county, V.20-31-2009, Col. Li Jingke, 1 female NEW COUNTRY RECORD
22°47'56"N, 108°19'44"EPeltonotus nasutusARECChina, Guangxi Zhuang Automonus Region NEW COUNTRY RECORD
17°28'59"N, 101°4'0"EPeltonotus nasutusIRSBNThailand, Changwat Loei, Na Haeo Bio. Sta., V-15-19-2003, light trap, Col. Constant, Smets, and Grootaert, 1 male, 2 female
17°28'59"N, 101°4'0"EPeltonotus nasutusIRSBNThailand, Changwat Loei, Na Haeo Bio. Sta., V.17.2003, edge pond, Col. Constant and Smets, 2 female
17°28'59"N, 101°4'0"EPeltonotus nasutusIRSBNThailand, Changwat Loei, Na Haeo Bio. Sta., V.5-12-2001, light trap, Col. Constant and Grootaert, 2 female
19°27'N, 98°20'EPeltonotus nasutusIRSBNN. Thailand, Mae Hong Son Prov., 600 m, 28-V to 2-VI-1999, Col. D. Hauck, 2 male, 2 female
14°16'07"N, 98°59'12"EPeltonotus nasutusIRSBNThailand, Kanchanaburi Prov., Sai Yok NP, VI.4–5.2003, Constant and Smets, 1 male, 1 female
13°49'59"N, 106°57'0"EPeltonotus nasutusIRSBNCambodia, Ratanakiri Prov., Phumi Kalai Thum., VI.1-19.2007, Col. Li Jingke, 1 male, 2 female NEW COUNTRY RECORD
12°18'09"N, 102°59'20"EPeltonotus nasutusIRSBNCambodia, Pursat Prov., Phnum Samkos Wildlife Sanctuary, XI.15, 2005, light trapping, col. Smets and Van, 5 male, 4 female NEW COUNTRY RECORD
12°18'09"N, 102°59'20"EPeltonotus nasutusIRSBNCambodia, Pursat Prov., Phnum Samkos Wildlife Sanctuary, IV.13-14, 2005, light trapping, primary forest edge, col. Smets and Van, 1 female, 1 male,
12°18'09"N, 102°59'20"EPeltonotus nasutusIRSBNCambodia, Pursat Prov., Phnum Samkos Wildlife Sanctuary, IV.16, 2005, light trapping, col. Smets and Van, 3 female, 1 male NEW COUNTRY RECORD
12°18'09"N, 102°59'20"EPeltonotus nasutusIRSBNCambodia, Pursat Prov., Phnum Samkos Wildlife Sanctuary, IV.15, 2005, light trapping, col. Smets and Van, 1 female NEW COUNTRY RECORD
12°51'2"N, 102°36'34"EPeltonotus nasutusDrumontCambodia, Pailin Prov., 270m, V.6-16.2008, col. Murzin, 2 female NEW COUNTRY RECORD
21°17'13"N, 101°10'02"EPeltonotus nasutusIRSBNNW Laos, Louang Namtha Prov., Muang Sing, Houaylong-Kao, VI.2-19.2010, 6 male, 16 female
17°58'0"N, 102°35'59"EPeltonotus nasutusIRSBNLaos, Vientiane Prov., IV.4-1915, 1 female
17°58'0"N, 102°35'59"EPeltonotus nasutusIRSBNLaos, Vientiane Prov., V.18-1915, 1 male
20°09'0"N, 101°19'53"EPeltonotus nasutusLi et al. 2012Laos, Bokeo Prov., Pha Ngam
16°46'30"N, 102°37'10"EPeltonotus nasutusJameson and Wada 2004Thailand, Khorat
14°35'21"N, 98°44'29"EPeltonotus nasutusJameson and Wada 2004Thailand, Pu Nam Long Hot Spring
14°47'53"N, 98°44'29"EPeltonotus nasutusJameson and Wada 2004Thailand, Khao Leam Dam
19°05'47"N, 100°57'09"EPeltonotus nasutusJameson and Wada 2004Thailand, Nan Province
19°21'46"N, 98°59'01"EPeltonotus nasutusJameson and Wada 2004Thailand, Ban Chiang Dao
14°43'02"N, 102°01'23"EPeltonotus nasutusJameson and Wada 2004Thailand, Khorat Prov., Pak Thong Chai
17°57'46"N, 102°36'54"EPeltonotus nasutusJameson and Wada 2004Laos, Vientane
19°36'41"N, 103°43'44"EPeltonotus nasutusJameson and Wada 2004Laos, Xiangkhouang
22°20'59"N, 96°55'00"EPeltonotus nasutusJameson and Wada 2004Myanmar, Gokhteik
21°14'14"N, 106°22'34"EPeltonotus nasutusJameson and Wada 2004Vietnam, Tonkin (north Vietnam)
10°44'57"N, 106°40'43"EPeltonotus nasutusJameson and Wada 2004Vietnam, Cochinchina (southern Vietnam)