- Current Issue
- Archive
- News & Multimedia
- For Authors
- About PNAS
- Collected Articles
- 100th Anniversary
- Colloquium Papers
- Commentaries
- Core Concepts
- Cozzarelli Prize
- Editorials
- Feature Articles
- Front Matter
- Inaugural Articles
- In This Issue
- Inner Workings
- Letters and Replies
- News Features
- Opinions
- Perspectives
- PNAS Classics
- PNAS Plus
- PNAS Portals
- Profiles
- QnAs
- Retrospectives
- Science and Culture
- Significance Statements
- Special Features
- Sustainability Science
- Browse by topic
- Anthropology
- Applied Biological Sciences
- Applied Mathematics
- Applied Physical Sciences
- Astronomy
- Biochemistry
- Biophysics and Computational Biology
- Cell Biology
- Chemistry
- Computer Sciences
- Developmental Biology
- Earth, Atmospheric, and Planetary Sciences
- Ecology
- Economic Sciences
- Engineering
- Environmental Sciences
- Evolution
- Genetics
- Geology
- Geophysics
- Immunology and Inflammation
- Mathematics
- Medical Sciences
- Microbiology
- Molecular Biology
- Neuroscience
- Pharmacology
- Physics
- Physiology
- Plant Biology
- Political Sciences
- Population Biology
- Psychological and Cognitive Sciences
- Social Sciences
- Statistics
- Sustainability Science
- Systems Biology
- Early Edition
- All Early Edition
- Highlights from Early Edition