Table 4. Species biplot scaling and percent variation explained by the first four axes in the "All Plants" CCA

 

 

Species scores (biplot scaling)

Cumulative fit per species as fraction of variance of species

Name (4)

Group

AX1

AX2

AX3

AX4

Eigenvalue

 

0.7842

0.595

0.4909

0.443

AX1

AX2

AX3

AX4

VAR(y)

% EXPL

Acer pensylvanicum

Tree

–0.6477

–0.0126

–0.224

–0.2932

0.1013

0.1014

0.1135

0.1343

4.14

23.83

Acer saccharum

Tree

–0.6045

–0.0624

0.0168

–0.4523

0.2439

0.2465

0.2467

0.3832

1.5

54.95

Acer spicatum

Tree

0.2383

0.5021

0.4853

–0.257

0.0058

0.0315

0.0555

0.0622

9.81

29.61

Actaea alba

  

–0.4637

0.2921

–0.1342

0.2284

0.0148

0.0207

0.0219

0.0255

14.53

25.69

Actaea rubra

  

0.5377

1.3499

–0.0968

0.2485

0.0214

0.1563

0.157

0.1616

13.51

33.48

Adiantum pedatum

SVP

–0.3871

0.4501

–0.0971

–0.0594

0.0107

0.0251

0.0257

0.026

14.07

20.41

Amphicarpaea bracteata

  

–0.4168

0.2298

–0.475

0.3906

0.0157

0.0204

0.0408

0.0545

11.09

32.03

Anemone virginiana

  

–0.9448

–0.0694

–0.1383

0.4348

0.0347

0.0349

0.0357

0.043

25.69

18.17

Apocynum androsaemifolium

  

–0.8865

–0.4118

0.0369

1.2884

0.0524

0.0637

0.0638

0.1746

14.99

25.22

Aquilegia canadensis

  

–0.7731

–0.8345

0.3009

4.263

0.0178

0.0385

0.0412

0.5821

33.6

75.91

Aralia nudicaulis

  

–0.6696

–0.11

0.0676

0.1693

0.1205

0.1237

0.1249

0.1326

3.72

17.98

Arisaema triphyllum

  

0.191

0.0565

–0.1383

–0.2677

0.012

0.0131

0.0193

0.0429

3.04

20.37

Asarum canadense

  

0.4281

1.1342

–0.131

–0.1059

0.0188

0.1506

0.1523

0.1535

9.76

36.96

Aster acuminatus

Asteraceae

–0.4093

–0.09

0.0078

–0.3071

0.0116

0.0121

0.0121

0.0186

14.49

9.53

Aster cordifolius

Asteraceae

–0.4146

0.5617

–0.0956

1.0412

0.0051

0.0146

0.0149

0.0473

33.4

38.12

Aster macrophyllus

Asteraceae

–0.8502

–0.4524

–0.004

2.4327

0.0576

0.0739

0.0739

0.5455

12.55

57.95

Athyrium filix-femina

SVP

0.9648

–0.2507

0.3542

–0.4498

0.1003

0.107

0.1206

0.1424

9.28

20.09

Betula alleghaniensis

Tree

0.757

–0.2126

1.2175

–0.5845

0.0319

0.0345

0.1171

0.1361

17.94

23.03

Bidens frondosa

Asteraceae

1.2615

–0.3955

–1.399

0.3219

0.1689

0.1855

0.3932

0.4042

9.42

55.05

Botrychium virginianum

SVP

–0.0488

–0.1113

–0.4355

–0.5518

0

0.0002

0.0029

0.0073

69.49

1.98

Brachyelytrum erectum

Poaceae

–0.2238

–0.2353

0.0937

–0.6219

0.0018

0.0039

0.0042

0.0183

27.39

10.29

Cardamine diphylla

  

1.896

–0.4412

1.3784

–0.4994

0.187

0.1972

0.296

0.309

19.22

49.99

Cardamine parviflora

  

–0.733

0.0012

–0.0426

0.6331

0.0286

0.0286

0.0287

0.0501

18.75

16.91

Carex albursina

Carex

–0.6405

–0.1311

0.2497

–0.4923

0.0187

0.0195

0.0223

0.0334

21.95

6.63

Carex appalachica

Carex

–0.8124

0.3067

–0.7954

–0.2415

0.0091

0.0105

0.0192

0.02

72.13

3.01

Carex arctata

Carex

–0.7743

–0.2145

0.0482

–0.387

0.0607

0.0654

0.0656

0.0807

9.88

17.98

Carex backii

Carex

–0.7118

–0.227

0.0532

0.911

0.0432

0.0476

0.0478

0.1186

11.73

22.23

Carex communis

Carex

–0.6489

–0.1039

–0.1373

–0.0643

0.0682

0.0699

0.073

0.0736

6.18

12.04

Carex deweyana

Carex

–0.5378

–0.0648

0.2477

–0.0257

0.0131

0.0133

0.0161

0.0161

22.06

11.95

Carex laxiflora

Carex

–0.5831

0.1716

–0.2447

–0.4924

0.0102

0.0111

0.0129

0.0202

33.29

9.26

Carex leptonervia

Carex

0.7117

–0.2898

–0.6382

–0.5534

0.0344

0.0401

0.0678

0.0886

14.72

27.75

Carex lucorum

Carex

–0.7091

–0.2224

–0.1612

–0.4132

0.0185

0.0204

0.0213

0.0276

27.11

13.49

Carex pedunculata

Carex

–0.7512

0.0468

0.1295

–0.4476

0.0335

0.0336

0.0346

0.0465

16.84

21.11

Carex pensylvanica

Carex

–0.7859

–0.1923

0.1933

0.4009

0.0379

0.0402

0.0424

0.0523

16.3

22.61

Carex plantaginea

Carex

0.324

0.5153

–0.928

–0.2705

0.0037

0.013

0.0433

0.0459

28.45

23.88

Carex platyphylla

Carex

–0.7804

0.0532

–0.4988

0.7798

0.0297

0.0298

0.0419

0.0716

20.52

19.48

Carex rosea

Carex

–0.9414

–0.0036

–0.1723

0.6125

0.0302

0.0302

0.0312

0.044

29.33

14.02

Carex scabrata

Carex

1.4179

–0.3291

–1.4197

–0.3404

0.1586

0.1672

0.3262

0.3353

12.68

47.09

Carex sprengellii

Carex

–0.5461

0.2696

0.8786

0.317

0.0012

0.0015

0.0045

0.0049

253.54

3.28

Carya cordiformis

Tree

–0.4263

–0.2354

–0.2932

–0.8418

0.003

0.0039

0.0053

0.0171

60.48

8.61

Caulophyllum thalictroides

  

0.2844

0.9293

0.3288

–0.1619

0.007

0.0815

0.0908

0.0931

11.59

20.59

Chenopodium gigantospermum

 

–0.5266

0.454

–0.1003

0.3529

0.0291

0.0508

0.0518

0.0649

9.52

20.93

Chrysosplenium americanum

 

1.252

–1.4156

0.029

–0.2191

0.1149

0.2619

0.2619

0.2655

13.64

48.59

Cinna latifolia

Poaceae

0.8317

–0.1498

–1.0705

–0.2528

0.0755

0.0779

0.203

0.2099

9.16

44.21

Circaea alpina

 

0.9193

0.1568

–0.5407

–0.3198

0.0846

0.0871

0.1163

0.1266

9.99

30.67

Circaea lutentiana

 

0.0884

–0.0027

–0.1125

–0.0958

0.0003

0.0003

0.0008

0.0012

25.77

14.07

Clintonia borealis

 

–0.6758

–0.1481

–0.0466

–0.0348

0.0263

0.0276

0.0277

0.0278

17.34

33.33

Cornus alternifolia

Shrub

–0.2744

0.2473

–0.0579

–0.3498

0.0134

0.0243

0.0249

0.0467

5.61

12.9

Cystopteris fragilis

SVP

–0.5706

0.3238

0.026

0.4247

0.013

0.0172

0.0172

0.0244

25.06

21.55

Deparia acrostichoides

SVP

0.8444

0.5971

–0.7727

0.0518

0.1393

0.209

0.3257

0.3262

5.12

46.48

Deschampsia flexuosa

Poaceae

–0.9038

–0.8466

0.5691

1.9922

0.0155

0.0291

0.0352

0.1104

52.77

18.88

Diervilla lonicera

Shrub

–0.618

–0.2346

0.08

3.016

0.0194

0.0222

0.0226

0.4856

19.65

66.73

Dirca palustris

Shrub

–0.4325

0.087

0.0044

0.1066

0.0101

0.0105

0.0105

0.0111

18.6

13.81

Dryopteris carthusiana

SVP

0.9417

–0.9334

2.8351

0.6249

0.0361

0.0716

0.399

0.4149

24.55

62.14

Dryopteris intermedia

SVP

0.1876

0.1333

0.1277

–0.3047

0.0091

0.0136

0.0178

0.0417

3.89

41.88

Dryopteris marginalis

SVP

–0.6595

–0.0167

–0.112

0.6121

0.0766

0.0767

0.0789

0.1448

5.68

37.78

Epilobium glandulosum

 

1.1921

–0.5477

–0.8018

0.1872

0.2156

0.2611

0.3586

0.3639

6.59

53.1

Epipactis helleborine

 

–0.1793

–0.2562

–0.3083

–0.3695

0.0024

0.0073

0.0145

0.0247

13.32

3.68

Equisetum arvense

SVP

0.3741

0.1726

–1.0511

–0.3373

0.0036

0.0043

0.0326

0.0355

39.09

23.34

Eupatorium rugosum

Asteraceae

0.5386

0.7967

0.6213

–0.2195

0.0229

0.073

0.1034

0.1072

12.67

18.84

Fagus grandifolia

Tree

–0.5968

0.0631

–0.1628

–0.3946

0.0759

0.0768

0.0824

0.1156

4.69

23.8

Fraxinus americana

Tree

–0.526

–0.1245

–0.0829

0.0878

0.0843

0.089

0.0911

0.0934

3.28

24.7

Galium palustre

 

1.4952

–1.9569

0.4528

–1.0018

0.1091

0.296

0.3061

0.355

20.49

43.75

Galium triflorum

 

–0.1738

0.0795

–0.4268

–0.2609

0.0044

0.0053

0.0317

0.0415

6.91

26.95

Geranium robertianum

 

–0.378

0.2321

0.0304

0.2088

0.0029

0.004

0.004

0.0049

49.23

14.22

Glyceria melicaria

Poaceae

1.5967

–0.5692

–1.6421

0.3502

0.2229

0.2513

0.487

0.4978

11.44

54.22

Glyceria striata

Poaceae

0.8172

–0.7404

–0.2542

–0.1829

0.043

0.0783

0.0825

0.0846

15.53

27.06

Huperzia lucidula

SVP

0.8426

–0.6346

0.8376

–0.863

0.0542

0.085

0.1386

0.1955

13.09

27.07

Impatiens capensis

 

1.6987

–0.93

–1.2916

0.4538

0.2551

0.3316

0.4791

0.4973

11.31

58.24

Impatiens pallida

 

0.582

2.3625

0.3561

0.8072

0.0105

0.1831

0.187

0.2072

32.33

39.02

Laportea canadensis

 

1.4035

2.5018

0.2967

–0.0046

0.1114

0.4654

0.4703

0.4703

17.68

64.59

Lonicera canadensis

Shrub

–0.6118

–0.07

–0.0178

–0.1365

0.0289

0.0293

0.0293

0.0307

12.95

11.03

Maianthemum canadense

 

–0.7907

–0.282

–0.0307

–0.311

0.0956

0.1077

0.1079

0.1227

6.54

25.92

Smilacina racemosa

 

–0.7993

–0.0758

–0.1815

0.7276

0.0447

0.0451

0.0474

0.0844

14.29

30.38

Matteucia sthruthiopteris

SVP

1.8114

1.2618

0.7749

0.0713

0.227

0.3372

0.3787

0.3791

14.45

51.92

Medeola virginiana

 

0.4793

–0.6269

0.4118

–0.9072

0.007

0.019

0.0241

0.0492

32.84

8.25

Milium effusum

Poaceae

–0.814

–0.0011

–0.0521

–0.294

0.0398

0.0398

0.0399

0.0451

16.66

11.53

Mimulus ringens

 

1.518

–1.8962

0.3724

0.0281

0.0719

0.184

0.1883

0.1883

32.07

44.77

Onoclea sensibilis

SVP

1.2867

–1.2485

1.4706

–0.2237

0.2031

0.3942

0.6595

0.6656

8.15

67.94

Oryzopsis asperifolia

Poaceae

–0.722

–0.3097

0.0142

–0.191

0.0586

0.0694

0.0695

0.0736

8.89

21.43

Oryzopsis racemosa

Poaceae

–0.5241

0.3209

–0.2067

0.8029

0.0188

0.0259

0.0288

0.0729

14.61

40.57

Osmorhiza claytonii

 

–0.0959

0.1639

–0.1186

–0.2102

0.0007

0.0027

0.0038

0.0071

13.28

11.98

Osmunda cinnamomea

SVP

1.4722

–2.1361

0.9892

–0.7106

0.1341

0.4163

0.4769

0.5081

16.17

67.48

Ostrya virginiana

Tree

–0.6096

–0.0986

–0.045

–0.3311

0.0877

0.09

0.0905

0.1164

4.24

28.1

Parthenocissus quinquefolia

  

0.3367

–0.0051

1.6206

0.3602

0.0038

0.0038

0.0919

0.0963

29.8

30.18

Phegopteris connectilis

SVP

2.1008

0.7278

1.529

–0.0639

0.1262

0.1413

0.2081

0.2083

34.98

37.81

Pilea pumila

  

1.1535

–0.3196

–1.2636

–0.5197

0.0738

0.0795

0.1681

0.1831

18.02

36.21

Poa compressa

Poaceae

–0.6289

–0.2246

–0.6036

1.5723

0.0111

0.0126

0.0228

0.0925

35.48

17.57

Poa saltuensis

Poaceae

–0.562

–0.0103

–0.2667

–0.5333

0.0121

0.0121

0.0148

0.0256

26.18

16.88

Polygonatum pubescens

   

–0.6007

–0.3019

–0.0067

–0.0811

0.0829

0.1038

0.1038

0.1053

4.35

35.26

Polygonum cilinode

  

–0.5381

–0.111

0.0745

–0.5665

0.0504

0.0526

0.0535

0.1095

5.74

23.04

Polypodium appalachianum

SVP

–0.7785

–0.113

–0.045

0.2144

0.0492

0.0502

0.0504

0.0541

12.32

39.69

Polystichum acrostichoides

SVP

–0.6819

0.2032

–0.5804

–0.5483

0.0248

0.027

0.045

0.061

18.75

11.15

Prunus pensylvanica

Tree

–0.7573

–1.0111

0.31

4.7952

0.0132

0.0366

0.0388

0.5666

43.57

75.99

Prunus serotina

Tree

–0.3502

–0.0097

0.3142

–0.3697

0.0041

0.0041

0.0075

0.0121

29.59

4.61

Prunus virginiana

Tree

–0.6407

–0.4162

0.4073

2.4656

0.025

0.0356

0.0457

0.416

16.42

62.37

Pteridium aquilinum

SVP

–0.6529

–0.4278

–0.5882

0.265

0.0067

0.0096

0.0151

0.0162

63.38

12.6

Pyrola elliptica

  

1.1223

–0.5093

0.0575

–0.6652

0.0346

0.0417

0.0418

0.0539

36.43

10.31

Quercus rubra

Tree

–0.8026

–0.1454

–0.1636

0.5152

0.0813

0.084

0.0874

0.1209

7.92

31.62

Ranunculus abortivus

  

0.4105

0.2092

–1.0408

–0.3138

0.0065

0.0081

0.0497

0.0535

26.07

20.27

Ribes cynosbati

Shrub

–0.8227

–0.0216

–0.1226

0.3834

0.0305

0.0305

0.0312

0.0378

22.22

18.57

Ribes glandulosum

Shrub

0.9245

–1.4818

2.9883

1.347

0.0273

0.0975

0.383

0.441

31.28

73.2

Rubus alleghaniensis

Shrub

–0.6155

0.0863

–0.2016

–0.2621

0.0424

0.0432

0.0477

0.0554

8.94

9.15

Rubus idaeus

Shrub

–0.508

0.1322

–0.0107

0.2805

0.0077

0.0083

0.0083

0.0106

33.37

13.64

Rubus occidentalis

Shrub

–0.1109

0.4003

–0.6456

–0.4603

0.0004

0.0062

0.0213

0.0289

27.69

23.17

Rubus odoratus

Shrub

–0.2775

0.3562

0.2413

–0.0287

0.0047

0.0125

0.0161

0.0162

16.27

35

Rubus pubescens

Shrub

0.9447

–0.3766

2.4928

–0.2256

0.0384

0.0445

0.312

0.3142

23.23

34.22

Sambucus racemosa

Shrub

–0.2975

0.1322

0.1413

–0.1685

0.0073

0.0088

0.0104

0.0128

12.07

18.78

Sanguinaria canadensis

  

–0.3645

0.3412

0.0356

–0.188

0.0086

0.0161

0.0162

0.0185

15.47

11.09

Saxifraga virginiensis

  

–0.7063

–0.0516

–0.4029

1.366

0.0219

0.0221

0.0292

0.1113

22.74

23.27

Scutellaria lateriflora

  

1.6314

–1.1055

–1.0578

0.2207

0.2756

0.4021

0.518

0.523

9.66

76.93

Solidago canadensis var scabra

Asteraceae

0.2438

0.3118

–0.5752

1.1275

0.0021

0.0056

0.0174

0.0628

28

35.24

Solidago caesia

Asteraceae

–0.6973

0.0368

–0.1076

0.225

0.1162

0.1165

0.1193

0.1314

4.18

33.77

Solidago flexicaulis

Asteraceae

–0.1975

0.2734

–0.3988

–0.1863

0.0023

0.0067

0.016

0.0181

17.02

16.75

Solidago rugosa

Asteraceae

0.9802

–1.153

3.4421

1.2313

0.0196

0.0467

0.2886

0.3195

49

67.9

Streptopus roseus

  

–0.7079

–0.1138

–0.0227

0.1898

0.0211

0.0216

0.0216

0.0231

23.8

27.06

Taraxacum officinale

Asteraceae

–0.0803

–0.0216

–0.2969

–0.1456

0.0013

0.0014

0.0197

0.0241

4.83

9.07

Thalictrum dioicum

  

–0.4928

0.3148

–0.2572

0.4995

0.0208

0.0293

0.035

0.0564

11.65

49.25

Tiarella cordifolia

  

1.093

–0.2577

0.4911

–0.5184

0.2248

0.2373

0.2827

0.3333

5.31

47.53

Tilia americana

Tree

–0.6273

0.176

–0.0779

0.1577

0.0412

0.0444

0.0451

0.0477

9.56

38.09

Trientalis borealis

  

0.383

–1.0412

0.7095

–0.2382

0.0165

0.1386

0.1953

0.2016

8.88

29.85

Ulmus rubra

Tree

0.3118

0.7424

–0.8277

–0.3891

0.0035

0.0232

0.0478

0.0532

27.9

29.37

Uvularia grandiflora

  

–0.4267

0.3337

–0.2468

–0.1114

0.0175

0.0281

0.034

0.0352

10.43

13.44

Vaccinium myrtilloides

Shrub

–0.8338

–0.9413

0.3588

4.1979

0.0216

0.0492

0.0532

0.6014

32.14

76.99

Valeriana officinalis

  

0.0657

0.3699

–0.9137

–0.4277

0.0001

0.0033

0.0228

0.027

42.85

21.11

Viburnum acerifolium

Shrub

–0.9036

–0.2398

0.0373

0.3705

0.0712

0.0762

0.0763

0.0883

11.47

30.39

Viburnum alnifolium

Shrub

0.4009

0.0145

–0.2986

–0.5713

0.0212

0.0212

0.033

0.0761

7.58

26.67

Viola canadensis

  

0.0235

0.3157

0.1706

–0.1447

0

0.0036

0.0046

0.0053

28.1

8.76

Viola pubescens

  

–0.3451

0.0346

–0.0044

–0.3826

0.0052

0.0053

0.0053

0.0117

22.85

18.84

†Plant phylogenetic or functional group. Species without a group were included in the analysis of "All plants" only.