AnimaliaColeopteraLycidaeBocekMatejBocakLadislavSpecies limits in polymorphic mimetic Eniclases net-winged beetles from New Guinean mountains (Coleoptera, Lycidae)Zookeys26520162016593153510.3897/zookeys.593.7728 Eniclases pseudoluteolus http://zoobank.org/9E3094C1-2958-4E4B-9D02-C58FCEDC7594 sp. n.Figs 36, 48Material examined.

Holotype. Male (BM0084), Indonesia, Irian Jaya, Maibri vill., Arfak Mts., 1600 m, Nov.–Dec. 1991 (LMBC).

Diagnosis.

Eniclases pseudoluteolus belongs to the group of uniformly yellow species which additionally includes Eniclases luteolus Waterhouse, 1878, Eniclases nigriceps Bocak & Bocakova, 1991, Eniclases fuscicornis Bocak & Bocakova, 1991, and Eniclases pallidus Bocak & Bocakova, 1991. Most of them have large eyes (Bocak and Bocakova 1991). Two species, Eniclases robustus Bocak & Bocakova, 1991 and Eniclases proximus Bocak & Bocakova, 1991 have the similar size of eyes with Eniclases pseudoluteolus. These species differ from Eniclases pseudoluteolus in the slender antennomere 3 and light colored thorax (antennomere of Eniclases pseudoluteolus as in Fig. 36). The similar species occur only in Eastern New Guinea in the vicinity of Wau and Mt. Hagen (Fig. 2).

Description.

Male. Body length 9.3 mm, head, apical antennomeres, thorax, and abdomen dark brown to black, pronotum and elytra yellow, trochanters brown, rest of legs black. Head with small, hemispherically prominent eyes, their diameter 0.90 interocular distance, antennae flat, acutely serrate, antennomere 3 triangular, wide, its apical process short (Fig. 36), further antennomeres similar in shape, becoming slenderer to apex of antennae. Pronotum transverse, with weak bulge in basal third, elytra with weak secondary costae and mostly regular, subquadrate cells.

Distribution.

New Guinea, Arfak Mts.

Etymology.

The specific name refers to the similarity with Eniclases luteolus.

Basal antennomeres of (Figs 30–42): 30 Eniclases apertus, male 31 Eniclases bicolor sp. n., female 32 Eniclases bokondinensis sp. n., female 33 Eniclases brancuccii sp. n., male 34 Eniclases divaricatus, male 35 Eniclases infuscatus sp. n., female 36 Eniclases pseudoluteolus, male 37 Eniclases similis, male 38 Eniclases variabilis sp. n., male 39 Eniclases elelimensis, male 40 Eniclases niger sp. n., female 41 Eniclases tikapurensis, male 42 Eniclases pseudoapertus sp. n., male. Male genitalia, ventral view (Figs 43–50): 43 Eniclases pseudoapertus sp. n. 44 Eniclases divaricatus 45 Eniclases tikapurensis sp. n. 46 Eniclases brancuccii sp. n. 47 Eniclases variabilis sp. n. 48 Eniclases pseudoluteolus sp. n. 49 Eniclases niger sp. n. 50 Eniclases similis. Scale bars: 0.5 mm.

1 Phylogenetic tree of Eniclases inferred from the maximum likelihood optimality criterion; a basal part of outgroups omitted. The numbers at branches show bootstrap support greater than 50%, genetic divergence within respective putative species and posterior probabilities inferred from the bPTP model 2 Dated tree produced using Bayesian inference.

 BocakLBocakovaM (1991) Revision of the genus Eniclases Waterhouse, 1879 (Coleoptera, Lycidae, Metriorrhynchinae). Mitteilungen der Münchener Entomologischen Gesellschaft 81: 203–226.