AnimaliaColeopteraLycidaeBocekMatejBocakLadislavSpecies limits in polymorphic mimetic Eniclases net-winged beetles from New Guinean mountains (Coleoptera, Lycidae)Zookeys26520162016593153510.3897/zookeys.593.7728 Eniclases divaricatus (Pic, 1921)Figs 6, 34 Trichalus (Trichalolus) divaricatusPic, 1921: 10. Material examined.

Lectotype. Female, New Guinea, Humboldt Bay, Doherty lgt., coll. Pic (MHNP). Other material examined. 4 males, 2 females (BM0001–2, 9, 15–17), Indonesia, Irian Jaya, Sentani, Cyclops Mts., 300 m, Nov.–Dec. 1991; female (BM0057), Indonesia, Irian Jaya, Elelim, path to Apalapsili, 600 m, Nov.–Dec. 1991 (LMBC).

Diagnosis.

Eniclases divaricatus is the only Papuan species with the bicolored elytra and flabellate male antennae (Figs 6, 34). Additionally, this species has the characteristic pattern of bright humeri and dark colored suture, which is shared only with some specimens of Eniclases similis from the same locality (Figs 6, 13). The similarly colored females cannot be distinguished as their relative size of eyes is similar (Table 1).

Redescription.

Male. Body length 8.8–9.7 mm. Head, thorax, legs, and humeri yellow to orange, antennae except basal part, abdomen, and most of elytra dark colored. Head with moderately large, hemispherically prominent eyes, their diameter 0.92–0.96 times minimum interocular distance Antennae flabellate, lamella of antennomere 3 slightly longer than the body of antennomere, other lamellae similar (Fig. 34). Pronotum transverse, with almost straight, slightly elevated lateral margins, elytra with quite strong straight secondary costae and regular dense cells (Fig. 6).

Distribution.

Central North New Guinea.

Habitus of Eniclases: 6 Eniclases divaricatus, male 7 Eniclases pseudoapertus sp. n., male 8 Eniclases apertus, male 9 Eniclases tikapurensis sp. n., male 10 Eniclases bicolor sp. n., female 11 Eniclases infuscatus sp. n., female 12 Eniclases brancuccii sp. n., female 13, 14 Eniclases similis, male 15, 16 Eniclases niger sp. n., male 17 Eniclases bokondinensis sp. n., female. Scale bars: 2 mm.

Basal antennomeres of (Figs 30–42): 30 Eniclases apertus, male 31 Eniclases bicolor sp. n., female 32 Eniclases bokondinensis sp. n., female 33 Eniclases brancuccii sp. n., male 34 Eniclases divaricatus, male 35 Eniclases infuscatus sp. n., female 36 Eniclases pseudoluteolus, male 37 Eniclases similis, male 38 Eniclases variabilis sp. n., male 39 Eniclases elelimensis, male 40 Eniclases niger sp. n., female 41 Eniclases tikapurensis, male 42 Eniclases pseudoapertus sp. n., male. Male genitalia, ventral view (Figs 43–50): 43 Eniclases pseudoapertus sp. n. 44 Eniclases divaricatus 45 Eniclases tikapurensis sp. n. 46 Eniclases brancuccii sp. n. 47 Eniclases variabilis sp. n. 48 Eniclases pseudoluteolus sp. n. 49 Eniclases niger sp. n. 50 Eniclases similis. Scale bars: 0.5 mm.

Measurements of Eniclases spp. (all measurements in mm, n.a. – not available).

Body lengthWidth humeriPronotumEye diameter/distance
lengthwidthmalefemale
Eniclases pseudoapertus 6.31.60.751.21.4n.a.
Eniclases divaricatus 8.8–9.72.1–2.31.2–1.31.7–1.70.92–0.960.80–0.87
Eniclases pseudoluteolus 9.32.31.151.60.9n.a.
Eniclases apertus 5.8–6.91.4–1.70.91.251.15–1.17
Eniclases tikapurensis 9.5–11.12.0–2.51.1–1.31.4–1.71.11–1.400.92–1.06
Eniclases bicolor 10.32.41.41.7n.a.0.71–0.74
Eniclases infuscatus 12.12.51.251.6n.a.0.79–0.84
Eniclases brancuccii 7.6–8.01.8–1.91.0–1.17.5–8.01.000.84–0.91
Eniclases bokondinensis 9.22.051.01.35n.a.0.72–0.82
Eniclases elelimensis 6.9–8.11.5–1.90.9–1.11.3–1.4n.a.0.78–0.89
Eniclases variabilis 6.6–8.21.6–2.00.1–1.11.1–1.350.83–0.950.70–0.85
Eniclases niger 9.2–11.62.2–2.81.3–1.69.0–11.51.17–1.280.89
Eniclases similis 7.5–9.71.9–2.31.1–1.41.8–1.81.02–1.150.89