Material examined.

Taxon classification

Animalia

Coleoptera

Lycidae

Bocek

Matej

Bocak

Ladislav

Species limits in polymorphic mimetic

Eniclases

net-winged beetles from New Guinean mountains (

Coleoptera

Lycidae

)

Zookeys265201620165931535

10.3897/zookeys.593.7728

Eniclases

tikapurensis

http://zoobank.org/CF8E0FB2-3F8F-4D6B-8AA3-3430D281B886

sp. n.

Figs 9, 41, 45

Material examined.

Holotype. Male (BM0039), Indonesia, Irian Jaya, Yiwika, N of Wamena, 2000 m, Nov.–Dec. 1991 (LMBC). Paratypes. 3 males, 2 females (BM0040–44), Indonesia, Irian Jaya, Tikapura village, 2200 m, Nov.–Dec. 1991 (LMBC).

Diagnosis.

Eniclases tikapurensis resembles in general appearance Eniclases papuensis Bocak & Bocakova, 1991 from the Panai Lake area approximately 250 km west of Yiwika. Both species are characteristic in pale hue of the yellow upper part (Fig. 9) but differ in the relative size of eyes when Eniclases tikapurensis has much larger eyes (Table 1).

Description.

Male. Body length 9.5–11.1 mm, slender (Table 1). Head, antennae, thorax, legs except trochanters and bases of femora bark brown to black, pronotum and elytra except posterior edge pale yellow (Fig. 9). Head with large, hemispherically prominent eyes, their diameter 1.11–1.40 times minimum interocular distance. Antennae serrate, antennomere 3 triangular, with pointed apical process, antennomere 4 parallel-sided in most of its length, its process shorter, shape of other antennomeres similar (Fig. 41). Pronotum transverse, lateral margins with weak bulge in basal third and prominent posterior angles, elytra with weak but well developed, straight secondary costae and regular, mostly quadrate, small elytral cells.

Distribution.

New Guinea, Upper Baliem Valley.

Etymology.

The specific name refers to the type locality, the village Tikapura, north of Tagime.

Remark.

The genetically distant population from Bokondini is a sister to Eniclases tikapurensis, but does not differ in any morphological character. As their delimitation would be based only on mtDNA sequence and further information on nuclear markers and geographical distribution is needed for these two putative cryptic species, we postpone the formal description of the population from Bokondini. The sequenced specimens representing the Bokondini population are designated as Eniclases sp. A. in Fig. 1.

Figures 6–17.

Habitus of Eniclases: 6 Eniclases divaricatus, male 7 Eniclases pseudoapertus sp. n., male 8 Eniclases apertus, male 9 Eniclases tikapurensis sp. n., male 10 Eniclases bicolor sp. n., female 11 Eniclases infuscatus sp. n., female 12 Eniclases brancuccii sp. n., female 13, 14 Eniclases similis, male 15, 16 Eniclases niger sp. n., male 17 Eniclases bokondinensis sp. n., female. Scale bars: 2 mm.

Figures 30–50.

Basal antennomeres of (Figs 30–42): 30 Eniclases apertus, male 31 Eniclases bicolor sp. n., female 32 Eniclases bokondinensis sp. n., female 33 Eniclases brancuccii sp. n., male 34 Eniclases divaricatus, male 35 Eniclases infuscatus sp. n., female 36 Eniclases pseudoluteolus, male 37 Eniclases similis, male 38 Eniclases variabilis sp. n., male 39 Eniclases elelimensis, male 40 Eniclases niger sp. n., female 41 Eniclases tikapurensis, male 42 Eniclases pseudoapertus sp. n., male. Male genitalia, ventral view (Figs 43–50): 43 Eniclases pseudoapertus sp. n. 44 Eniclases divaricatus 45 Eniclases tikapurensis sp. n. 46 Eniclases brancuccii sp. n. 47 Eniclases variabilis sp. n. 48 Eniclases pseudoluteolus sp. n. 49 Eniclases niger sp. n. 50 Eniclases similis. Scale bars: 0.5 mm.

Table 1.

Measurements of Eniclases spp. (all measurements in mm, n.a. – not available).

	Body length	Width humeri	Pronotum		Eye diameter/distance
	Body length	Width humeri	length	width	male	female
Eniclases pseudoapertus	6.3	1.6	0.75	1.2	1.4	n.a.
Eniclases divaricatus	8.8–9.7	2.1–2.3	1.2–1.3	1.7–1.7	0.92–0.96	0.80–0.87
Eniclases pseudoluteolus	9.3	2.3	1.15	1.6	0.9	n.a.
Eniclases apertus		5.8–6.9	1.4–1.7	0.9	1.25	1.15–1.17
Eniclases tikapurensis	9.5–11.1	2.0–2.5	1.1–1.3	1.4–1.7	1.11–1.40	0.92–1.06
Eniclases bicolor	10.3	2.4	1.4	1.7	n.a.	0.71–0.74
Eniclases infuscatus	12.1	2.5	1.25	1.6	n.a.	0.79–0.84
Eniclases brancuccii	7.6–8.0	1.8–1.9	1.0–1.1	7.5–8.0	1.00	0.84–0.91
Eniclases bokondinensis	9.2	2.05	1.0	1.35	n.a.	0.72–0.82
Eniclases elelimensis	6.9–8.1	1.5–1.9	0.9–1.1	1.3–1.4	n.a.	0.78–0.89
Eniclases variabilis	6.6–8.2	1.6–2.0	0.1–1.1	1.1–1.35	0.83–0.95	0.70–0.85
Eniclases niger	9.2–11.6	2.2–2.8	1.3–1.6	9.0–11.5	1.17–1.28	0.89
Eniclases similis	7.5–9.7	1.9–2.3	1.1–1.4	1.8–1.8	1.02–1.15	0.89

Figures 1–2.

1 Phylogenetic tree of Eniclases inferred from the maximum likelihood optimality criterion; a basal part of outgroups omitted. The numbers at branches show bootstrap support greater than 50%, genetic divergence within respective putative species and posterior probabilities inferred from the bPTP model 2 Dated tree produced using Bayesian inference.