Material examined.

Taxon classification

Animalia

Coleoptera

Lycidae

Bocek

Matej

Bocak

Ladislav

Species limits in polymorphic mimetic

Eniclases

net-winged beetles from New Guinean mountains (

Coleoptera

Lycidae

)

Zookeys265201620165931535

10.3897/zookeys.593.7728

Eniclases

bokondinensis

http://zoobank.org/C10E3541-F9FB-4C01-8570-9F7568B3BBD7

sp. n.

Figs 17, 18, 32

Material examined.

Holotype. Female (BM0095), Indonesia, Irian Jaya, Bokondini, 1900 m, Nov.–Dec. 1991 (LMBC). Paratypes. 2 females (BM0092, 94), the same data as the holotype (LMBC).

Diagnosis.

Eniclases bokondinensis has a characteristic combination of the black pronotum and light yellow elytra with dark colored apex (Figs 17–18). The shape of antennae and the relative size of eyes are similar to those of Eniclases variabilis and Eniclases elelimensis (Table 1, Figs 32, 38, 39). The co-mimics of Eniclases bokondinensis are large-bodied and this species has a larger body than its closest relatives (Fig. 1, Table 1).

Description.

Female. Body length 9.2 mm, head, antennae, thorax, and abdomen dark brown to black, elytra pale yellow in humeral half, gradually infuscate to apex (Figs 17–18), trochanters and basal parts of femora brown, rest of legs black. Head with small, hemispherically prominent eyes, their diameter 0.72–0.82 interocular distance, antennae flat, slender, acutely serrate, apical process of antennomere 3 about half length of its body, further antennomeres similar in shape. Pronotum transverse, with weak bulge in basal third, elytra with weak but regular secondary costae and mostly regular, subquadrate cells (Figs 17–18).

Distribution.

New Guinea, Bokondini region.

Etymology.

The specific name refers to the type locality.

Figures 6–17.

Habitus of Eniclases: 6 Eniclases divaricatus, male 7 Eniclases pseudoapertus sp. n., male 8 Eniclases apertus, male 9 Eniclases tikapurensis sp. n., male 10 Eniclases bicolor sp. n., female 11 Eniclases infuscatus sp. n., female 12 Eniclases brancuccii sp. n., female 13, 14 Eniclases similis, male 15, 16 Eniclases niger sp. n., male 17 Eniclases bokondinensis sp. n., female. Scale bars: 2 mm.

Figures 18–29.

Habitus of Eniclases: 18 Eniclases bokondinensis sp. n., female 19 Eniclases elelimensis, female 20–29 Eniclases variabilis sp. n. Scale bars: 2 mm.

Figures 30–50.

Basal antennomeres of (Figs 30–42): 30 Eniclases apertus, male 31 Eniclases bicolor sp. n., female 32 Eniclases bokondinensis sp. n., female 33 Eniclases brancuccii sp. n., male 34 Eniclases divaricatus, male 35 Eniclases infuscatus sp. n., female 36 Eniclases pseudoluteolus, male 37 Eniclases similis, male 38 Eniclases variabilis sp. n., male 39 Eniclases elelimensis, male 40 Eniclases niger sp. n., female 41 Eniclases tikapurensis, male 42 Eniclases pseudoapertus sp. n., male. Male genitalia, ventral view (Figs 43–50): 43 Eniclases pseudoapertus sp. n. 44 Eniclases divaricatus 45 Eniclases tikapurensis sp. n. 46 Eniclases brancuccii sp. n. 47 Eniclases variabilis sp. n. 48 Eniclases pseudoluteolus sp. n. 49 Eniclases niger sp. n. 50 Eniclases similis. Scale bars: 0.5 mm.

Table 1.

Measurements of Eniclases spp. (all measurements in mm, n.a. – not available).

	Body length	Width humeri	Pronotum		Eye diameter/distance
	Body length	Width humeri	length	width	male	female
Eniclases pseudoapertus	6.3	1.6	0.75	1.2	1.4	n.a.
Eniclases divaricatus	8.8–9.7	2.1–2.3	1.2–1.3	1.7–1.7	0.92–0.96	0.80–0.87
Eniclases pseudoluteolus	9.3	2.3	1.15	1.6	0.9	n.a.
Eniclases apertus		5.8–6.9	1.4–1.7	0.9	1.25	1.15–1.17
Eniclases tikapurensis	9.5–11.1	2.0–2.5	1.1–1.3	1.4–1.7	1.11–1.40	0.92–1.06
Eniclases bicolor	10.3	2.4	1.4	1.7	n.a.	0.71–0.74
Eniclases infuscatus	12.1	2.5	1.25	1.6	n.a.	0.79–0.84
Eniclases brancuccii	7.6–8.0	1.8–1.9	1.0–1.1	7.5–8.0	1.00	0.84–0.91
Eniclases bokondinensis	9.2	2.05	1.0	1.35	n.a.	0.72–0.82
Eniclases elelimensis	6.9–8.1	1.5–1.9	0.9–1.1	1.3–1.4	n.a.	0.78–0.89
Eniclases variabilis	6.6–8.2	1.6–2.0	0.1–1.1	1.1–1.35	0.83–0.95	0.70–0.85
Eniclases niger	9.2–11.6	2.2–2.8	1.3–1.6	9.0–11.5	1.17–1.28	0.89
Eniclases similis	7.5–9.7	1.9–2.3	1.1–1.4	1.8–1.8	1.02–1.15	0.89

Figures 1–2.

1 Phylogenetic tree of Eniclases inferred from the maximum likelihood optimality criterion; a basal part of outgroups omitted. The numbers at branches show bootstrap support greater than 50%, genetic divergence within respective putative species and posterior probabilities inferred from the bPTP model 2 Dated tree produced using Bayesian inference.