######################################################################################################################################################### ####This section will run through how we process ChIPseq replicate data. ### Run Bowtie on the Fastq files. # Requires as input: Fastq file; bowtie reference files # Will generate a bam file. # Bowtie version 1.0.1 # Samtools version 0.1.19 # You will want to run bowtie on both the input and the experimental ChIP files. # You will need to edit this script to specify the path to the bowtie reference files perl getAlignment_Bowtie_SingleEnd.pl FileName.Fastq ### Run MACS on the resultant bamfiles. # Requires: INPUT bowtie_output.bam; CHIP bowtie_output.bam # Script also requires that you specify an identifier name for the output # MACS version 1.4.1 # Output will be a directory containing MACS output for both e-5 and e-10 # Script will also convert wiggle files to FIXED step perl macs_module_multiple.pl CHIP_bowtie_output.bam INPUT_bowtie_output.bam Identifier ### Normalize wiggle files to the median and trim megahits. # This perl script is written to handle TY1072 and TY2205 normalization - Note that it is essential that you include these strain names in the file name. # Requires: e-5 fixed step wiggle files. # Output will be a normalized wiggle file ("_norm_totalcounts_trim.wig" will be appended to the file name) # You will want to run bowtie on both the input and the experimental ChIP files. perl COUNTwig_normalize_bytotalcounts-megahits_forfolder_median.pl MACS141_e5_IDENTIFIER_afterfiting_all_FIXED.wig.gz ### Subtract the input # Requires: CHIP_norm_totalcounts_trim.wig.gz; INPUT_norm_totalcounts_trim.wig.gz # Script also requires that you specify the input ID # Will generate a fixed step wiggle file, subtracting the input value from the chip value at every line. # Example output file name: CHIP_norm_totalcounts_trim_subt_INPUT_ID.wig.gz perl COUNTwig_subtract_SE_folder.pl CHIP_norm_totalcounts_trim.wig.gz INPUT_norm_totalcounts_trim.wig.gz INPUT_ID #####At this point you are done with the individual file normalization. If you have replicates, go on to the next step#### ### Sort the bamfiles # Samtools version 0.1.19 # You will want to sort any replicates of both the ChIPs and the Inputs perl sortBam.pl CHIP_bowtie_output.bam ### Merge the bamfiles # Samtools version 0.1.19 # Requires that the bamfiles be sorted # Requires that you specify an output file name perl mergeMultipleBam.pl MERGED_CHIP_file.bam CHIP_file1_sorted.bam CHIP_file2_sorted.bam ... CHIP_fileN_sorted.bam perl mergeMultipleBam.pl MERGED_INPUT_file.bam INPUT_file1_sorted.bam INPUT_file2_sorted.bam ... INPUT_fileN_sorted.bam ### Run MACS on the merged files # Requires: MERGED_INPUT_file.bam; MERGED_CHIP_file.bam # Script also requires that you specify an identifier name for the output # MACS version 1.4.2 # Output will be a directory containing MACS output for both e-5 and e-10 # Script will also convert wiggle files to FIXED step perl macs_module_replicates.pl MERGED_CHIP_file.bam MERGED_INPUT_file.bam Identifier ### Get the overlapping peaks # Requires: MERGED_MACS_e10_peaks.bed (from the merged MACS) and MACS_e5_peaks.bed (from each of the individual replicated # Bedtools version 2.25.0 # script requires that you BE IN THE SAME DIRECTORY AS THE MERGED MACS OUTPUT FOLDER # Note there is a space separating e10 peak file from the comma separated list of e5 peak files. Note there are no spaces between the comma separated files. # Will produce a peaks_final.bed file in the e10 folder in the merged MACS folder perl getOverlappingPeaks_Replicates.pl path_to_e10_peaks.bed path_to_e5_peaks_rep1.bed,path_to_e5_peaks_rep2.bed,… ### Get the average wiggle file from the input subtracted replicates # Requires the individual input substracted wiggle files # Script also requires that you name the resulting output file # Script will append “_average.wig” to the end of the OutfileName. perl getAverageWig.pl path_to_INPUTSUBT_rep1.wig path_to_INPUTSUBT_rep2.wig path_to_INPUTSUBT_repN.wig OutfileName ######################################################################################################################## ###To generate log2 fixed step wiggle files comparing any two experimental conditions: # We use the MACS e-5 FIXED_STEP wiggle files as input #(1) paste together the files to be compared paste control_MACS_e5_FIXED.wig experimental_MACS_e5_FIXED.wig > ctrl_exp.wig #(2) calculate the log2 value at every step #will append "_log2" to the file name perl getLog2.pl ctrl_exp.wig #(3) separate the data by chromosome #ex: awk '/CHROMOSOME_II /{f=0} f; /CHROMOSOME_I /{f=1}' ctrl_exp_log2.wig > ChrI_ctrl_exp_log2.wig #ex: sed -e '1,/CHROMOSOME_X /d' ctrl_exp_log2.wig > ChrX_ctrl_exp_log2.wig ########################################### SCRIPTS ####################################################################### ######## COUNTwig_normalize_bytotalcounts-megahits_forfolder_median ############## #!/usr/bin/perl #adapted to run on a bunch of files in a folder with only wig files to be normalized. # this also trims the megahits (if one base pair has more than 1/10,000 (0.01%) of the total counts (if flat, 1/100,000,000 expected, thus 10,000 more than expected) # this script generated by the ERCAN LAB at NYU # contact: se71@nyu.edu use Statistics::Descriptive::Discrete; use Cwd; $dir=getcwd; my $wigfile = shift(); #gunzips the file if ($wigfile =~ /gz/) { my $cmd = "gzip -d $wigfile"; chomp ($cmd); system($cmd); $wigfile =~ s/\.gz//g; print STDERR "\nUnzipped: $wigfile\n"; } $Output = $wigfile; $Output =~ s/\.wig/_norm_totalcounts_trim.wig/; print STDERR "\nHere is output filename $Output\n"; open (OUT, ">$Output") || die "Can't open output file $Output\n"; # Custom track names my $desc = $Output; $desc =~ s/\.wig//g; open(aLINE, $wigfile) || die "\nCan't open inputfile $wigfile\n"; my $count = 0; my $totalines = 0; my $totalcounts =0; my $divifactor = 0; my $factor_x = 0; my $trimcutoff = 0; my $norm_value = 0; my $trimmedbases = 0; my $max = 0; my $mtFlag = 0; my $stats = new Statistics::Descriptive::Discrete; while (my $aline = ){ chomp($aline); #my @alineSeg = split(/\s/, $aline); if ($aline =~ /track/) { print STDERR "\nA: $aline\t"; } elsif ($aline =~ /fixedStep/) { if ($aline =~ /CHROMOSOME_MtDNA/) { print STDERR "\nA: Skipping CHROMOSOME_MtDNA...\n"; $mtFlag = 1; } else { $mtFlag = 0; print STDERR "\nA: $aline\t"; } } else { unless ($mtFlag) { unless ($aline eq " ") { $max = $aline if ($max < $aline); $totalcounts= $totalcounts+$aline; $stats->add_data($aline); } $count++; $totalines++; } } if ($count % 1000000 == 0) { print STDERR "$count."; } } close aLINE; $divifactor = $stats->median();# $totalcounts/$totalines; $factor_x = $divifactor/2; $trimcutoff = $totalcounts/10000; print STDERR "\nTotal count is : $totalcounts\n"; print STDERR "\nTotal lines is : $totalines\n"; print STDERR "\nThe old division factor is: ".$totalcounts/$totalines."\n"; print STDERR "\nThe max is : $max\n"; print STDERR "\nThe division factor is : $divifactor\n"; print STDERR "\nThe trim cutoff is : $trimcutoff\n"; print STDERR "Average: ".$stats->mean()."\n"; print STDERR "Median: ".$divifactor."\n"; print STDERR "Median for X: $factor_x\n"; print STDERR "Dividing counts by median counts...\n"; open(aLINE, $wigfile) || die "\nCan't open inputfile $wigfile\n"; $count = 0; my $Xflag = 0; while (my $aline = ){ chomp($aline); #my @alineSeg = split(/\s/, $aline); if ($aline =~ /track/) { print STDERR "\nA: $aline\t"; print OUT "track type=wiggle_0 name=\"$desc\" description=\"$desc\" visibility=full autoScale=off windowingFunction=mean smoothingWindow=8 viewLimits=-2:5\n"; } elsif ($aline =~ /fixedStep/) { if ($aline =~ /CHROMOSOME_X/) { $Xflag = 1; print STDERR "\nX! " if ($wigfile =~ /TY2205/ || $wigfile =~ /TY1072/); } else { $Xflag = 0; } print STDERR "\nA: $aline\t"; print OUT $aline."\n"; } else { if ($aline eq " ") { print OUT " \n"; $count++; } else { if($aline > $trimcutoff) { if ($Xflag && ($wigfile =~ /TY2205/ || $wigfile =~ /TY1072/)) { $norm_value = sprintf "%.2f", ($trimcutoff / $factor_x); } else { $norm_value = sprintf "%.2f", ($trimcutoff / $divifactor); } $trimmedbases++; } else { if ($Xflag && ($wigfile =~ /TY2205/ || $wigfile =~ /TY1072/)) { $norm_value = sprintf "%.2f", ($aline / $factor_x); } else { $norm_value = sprintf "%.2f", ($aline / $divifactor); } } print OUT "$norm_value\n"; $count++; } if($count % 1000000 == 0) { print STDERR "$count."; } } } close aLINE; close OUT; print STDERR "\nTotal trimmed bases : $trimmedbases\n"; # Gzipping $cmd = "gzip $Output"; chomp ($cmd); system($cmd); print STDERR "Gzipped file: $Output\n\n\n"; $r=`rm $wigfile`; print STDERR "Removed $wigfile\n"; sub COUNTwig_median{ # Open file my ($filename) = @_; open(LINE, $filename) || die "Can't open file $filename\n"; # Reading the file print STDERR "Reading a file for median calculation...\n"; my $tbase = 0; my %bCount; #Hash: Key, basecounts species; Value, the number (occurence) of the Key while (my $line = ) { chomp $line; $line =~ s/\r//g; my @lineSeg = split(/\s/, $line); if (not $lineSeg[0] eq 'track' and not $lineSeg[0] eq 'fixedStep' and not $lineSeg[0] eq '') { $bCount{$line}++; $tbase++; if($tbase % 1000000 == 0){print STDERR "$tbase.";} } } print STDERR "\nThe toal number of bp is: $tbase\n"; # Sort hash in the acending order and store keys and values in arrays individually. my (@bCountValue, @bCountKey); foreach $key (sort {$a<=>$b} (keys(%bCount))) { push (@bCountKey, $key); push (@bCountValue, $bCount{$key}); } # Median calculation my $cumlative = 0; my $hashCount = 0; my $stopper = 0; my ($median, $premedian, $postmedian); if ($tbase % 2) { do { $cumlative += $bCountValue[$hashCount]; if($cumlative >= int($tbase/2) +1) { $median = $bCountKey[$hashCount]; $stopper = 1; print STDERR "HashCount: $hashCount\n"; } $hashCount++; } until ($stopper == 1); }else { do { $cumlative += $bCountValue[$hashCount]; if ($cumlative >= $tbase/2) { if($cumlative >= ($tbase/2)+1) { $median = $bCountKey[$hashCount]; $stopper = 1; print STDERR "HashCount: $hashCount\n"; } elsif ($cumlative < ($tbase/2)+1) { $premedian = $bCountKey[$hashCount]; $postmedian = $bCountKey[$hashCount+1]; $median = ($premedian + $postmedian)/2; $stopper = 1; print STDERR "HashCount: $hashCount\n"; print STDERR "BaseCount of median-1: $premedian\n"; print STDERR "BaseCount of median+1: $postmedian\n"; } } $hashCount++; } until ($stopper == 1); } return $median; } sub median{ my @array = @_; my $count = scalar @array; @array = sort { $a <=> $b } @array; print "this is the sorted @array\n"; if ($count % 2) { $median = $array[int($count/2)]; } else { $median = ($array[$count/2] + $array[$count/2 - 1]) / 2; } return $median; } sub log10 { my $n = shift; return log($n)/log(10); } sub select_files { #WORKS my ($search, $numFiles) = @_; if($search eq 'ALL'){$search = ();} my @files = get_files($search); my $num = print_number_array(\@files); unless($numFiles == 1){ $num++; print STDERR "$num. All files\n"; } my $done = $num + 1; my $fSelect = 0; my (@selectFiles, $sFile); while ($fSelect == 0){ if($numFiles == 1){ my $checkFlag = 0; while($checkFlag == 0){ print STDERR "\nSelect a file by number: "; $sFile = ; chomp($sFile); $checkFlag = check_num($sFile, $num); } push(@selectFiles, $files[$sFile - 1]); $fSelect = 1; }else{ print STDERR "\nSelect a file by number or $done when FINISHED: "; $sFile = ; chomp($sFile); if ($sFile == $num){ @selectFiles = @files; $fSelect = 1; }elsif($sFile == $done){ $fSelect = 1; }elsif($sFile > 0 && $sFile < $num){ push(@selectFiles, $files[$sFile - 1]); }else{ print STDERR "$sFile is not a valid option\n"; } } } return @selectFiles; } sub get_files { #WORKS my $search = shift; my @flist; my $path = "." ; opendir(DIR, $path) || die "Can't open $path: $!"; while ( $entry = readdir(DIR) ) { # my $type = ( -d "$path\\$entry" ) ? "dir" : "file"; #Windows my $type = ( -d "$path\/$entry" ) ? "dir" : "file"; # Linux if($type =~ 'file'){ unless($entry =~ '.pl' || $entry =~ '.tsv' || $entry =~ '.txt'){ if (defined($search)){ if ($entry =~ $search){push(@flist, $entry);} }else{push(@flist, $entry);} } } }closedir( DIR ); return @flist; } sub print_number_array { #WORKS my $count = 0; local(*vals) = @_; foreach my $val (@vals){ $count++; print STDERR "$count. $val\n"; } return $count; } sub check_num { #WORKS my ($entry, $bound) = @_; if ( $entry > 0 && $entry <= $bound){ return 1; }else{ print STDERR "\n$entry is not a valid option\n"; return 0; } } sub check_type { my $file = shift; if ( $fType{$file} > 0 && $fType{$file} < ($#fTypes + 2) ){ print STDERR "\n$file is a $fTypes[$fType{$file} - 1] type file\n"; return $fType{$file}; }else{ $type = file_type($file); print STDERR "\n$file is a $fTypes[$type - 1] type file\n"; return $type; } } ##### COUNTwig_subtract_SE_folder ####### #!/usr/bin/perl # this script generated by the ERCAN LAB at NYU # contact: se71@nyu.edu use strict; use warnings; print STDERR "\nThis program returns the difference calculated by sample minus input\n"; my $treat = shift(); my $input = shift(); my $fileB = shift(); if ($treat =~ /gz/) { my $cmd = "gzip -d $treat"; chomp ($cmd); system($cmd); $treat =~ s/\.gz//g; print STDERR "\nUnzipped: $treat\n"; } if ($input =~ /gz/) { my $cmd = "gzip -d $input"; chomp ($cmd); system($cmd); $input =~ s/\.gz//g; print STDERR "\nUnzipped: $input\n"; } my $cmd = "wc -l $treat"; print STDERR "\nThe total line number for sample is: \n"; system($cmd); $cmd = "wc -l $input"; print STDERR "\nThe total line number for input is: \n"; system($cmd); $fileB = '_subt_' . $fileB; my $Output = $treat; $Output =~ s/\.wig/$fileB\.wig/g; open (OUT, ">$Output") || die "Can't open output file $Output\n"; print STDERR "\nHere is output filename $Output\n"; # Custom track names my $desc = $Output; $desc =~ s/\.wig//g; # Caluculation of the difference print STDERR "Calculating difference (A-B)\n"; open(aLINE, $treat) || die "\nCan't open inputfile $treat\n"; open(bLINE, $input) || die "\nCan't open inputfile $input\n"; my $count = 0; while (my $aline = ){ my $bline = ; chomp($aline); chomp($bline); #my @alineSeg = split(/\s/, $aline); #my @blineSeg = split(/\s/, $bline); if ($aline =~ /track/) { print STDERR "\nSample: $aline\t"; print STDERR "Input: $bline\n"; print OUT "track type=wiggle_0 name=\"$desc\" description=\"$desc\" visibility=full autoScale=off windowingFunction=mean smoothingWindow=8 viewLimits=-2:5\n"; } elsif ($aline =~ /fixedStep/) { print STDERR "\nSample: $aline\t"; print STDERR "Input: $bline\n"; print OUT $aline."\n"; } else { if ($aline eq " " || $bline eq " ") { print OUT " \n"; } else { my $diff = ($aline - $bline); print OUT "$diff\n"; } $count++; } if($count % 1000000 == 0) { print STDERR "$count."; } } close aLINE; close bLINE; close OUT; # Gzipping $cmd = "gzip $Output"; chomp ($cmd); system($cmd); $cmd = "gzip $treat"; chomp ($cmd); system($cmd); $cmd = "gzip $input"; chomp ($cmd); #system($cmd); sub COUNTwig_median{ # Open file my ($filename) = @_; open(LINE, $filename) || die "Can't open file $filename\n"; # Reading the file print STDERR "Reading a file for median calculation...\n"; my $tbase = 0; my %bCount; #Hash: Key, basecounts species; Value, the number (occurence) of the Key while (my $line = ) { chomp $line; $line =~ s/\r//g; my @lineSeg = split(/\s/, $line); if (not $lineSeg[0] eq 'track' and not $lineSeg[0] eq 'fixedStep' and not $lineSeg[0] eq '') { $bCount{$line}++; $tbase++; if($tbase % 1000000 == 0){print STDERR "$tbase.";} } } print STDERR "\nThe toal number of bp is: $tbase\n"; # Sort hash in the acending order and store keys and values in arrays individually. my (@bCountValue, @bCountKey); foreach my $key (sort {$a<=>$b} (keys(%bCount))) { push (@bCountKey, $key); push (@bCountValue, $bCount{$key}); } # Median calculation my $cumlative = 0; my $hashCount = 0; my $stopper = 0; my ($median, $premedian, $postmedian); if ($tbase % 2) { do { $cumlative += $bCountValue[$hashCount]; if($cumlative >= int($tbase/2) +1) { $median = $bCountKey[$hashCount]; $stopper = 1; print STDERR "HashCount: $hashCount\n"; } $hashCount++; } until ($stopper == 1); }else { do { $cumlative += $bCountValue[$hashCount]; if ($cumlative >= $tbase/2) { if($cumlative >= ($tbase/2)+1) { $median = $bCountKey[$hashCount]; $stopper = 1; print STDERR "HashCount: $hashCount\n"; } elsif ($cumlative < ($tbase/2)+1) { $premedian = $bCountKey[$hashCount]; $postmedian = $bCountKey[$hashCount+1]; $median = ($premedian + $postmedian)/2; $stopper = 1; print STDERR "HashCount: $hashCount\n"; print STDERR "BaseCount of median-1: $premedian\n"; print STDERR "BaseCount of median+1: $postmedian\n"; } } $hashCount++; } until ($stopper == 1); } return $median; } sub median{ my @array = @_; my $count = scalar @array; @array = sort { $a <=> $b } @array; print "this is the sorted @array\n"; my $median; if ($count % 2) { $median = $array[int($count/2)]; } else { $median = ($array[$count/2] + $array[$count/2 - 1]) / 2; } return $median; } sub log10 { my $n = shift; return log($n)/log(10); } sub select_files { #WORKS my ($search, $numFiles) = @_; if($search eq 'ALL'){$search = ();} my @files = get_files($search); my $num = print_number_array(\@files); unless($numFiles == 1){ $num++; print STDERR "$num. All files\n"; } my $done = $num + 1; my $fSelect = 0; my (@selectFiles, $sFile); while ($fSelect == 0){ if($numFiles == 1){ my $checkFlag = 0; while($checkFlag == 0){ print STDERR "\nSelect a file by number: "; $sFile = ; chomp($sFile); $checkFlag = check_num($sFile, $num); } push(@selectFiles, $files[$sFile - 1]); $fSelect = 1; }else{ print STDERR "\nSelect a file by number or $done when FINISHED: "; $sFile = ; chomp($sFile); if ($sFile == $num){ @selectFiles = @files; $fSelect = 1; }elsif($sFile == $done){ $fSelect = 1; }elsif($sFile > 0 && $sFile < $num){ push(@selectFiles, $files[$sFile - 1]); }else{ print STDERR "$sFile is not a valid option\n"; } } } return @selectFiles; } sub get_files { #WORKS my $search = shift; my @flist; my $path = "." ; opendir(DIR, $path) || die "Can't open $path: $!"; while ( my $entry = readdir(DIR) ) { # my $type = ( -d "$path\\$entry" ) ? "dir" : "file"; #Windows my $type = ( -d "$path\/$entry" ) ? "dir" : "file"; # Linux if($type =~ 'file'){ unless($entry =~ '.pl' || $entry =~ '.tsv' || $entry =~ '.txt'){ if (defined($search)){ if ($entry =~ $search){push(@flist, $entry);} }else{push(@flist, $entry);} } } }closedir( DIR ); return @flist; } sub print_number_array { #WORKS my $count = 0; my $vals = @_; foreach my $val (@{$vals}){ $count++; print STDERR "$count. $val\n"; } return $count; } sub check_num { #WORKS my ($entry, $bound) = @_; if ( $entry > 0 && $entry <= $bound){ return 1; }else{ print STDERR "\n$entry is not a valid option\n"; return 0; } } #sub check_type { # my $file = shift; # if ( $fType{$file} > 0 && $fType{$file} < ($#fTypes + 2) ){ # print STDERR "\n$file is a $fTypes[$fType{$file} - 1] type file\n"; # return $fType{$file}; # }else{ # $type = file_type($file); # print STDERR "\n$file is a $fTypes[$type - 1] type file\n"; # return $type; # } #} ###### getAlignment_Bowtie_SingleEnd ####### # this script aligns single-end reads in fastq format using bowtie # usage # go to the directory where you have the required files # perl getAlignment_Bowtie_SingleEnd.pl # this script generated by the ERCAN LAB at NYU # contact: se71@nyu.edu use Cwd; use strict; use warnings; my $dir = getcwd; my $file = shift(); my @temp = split(/\./, $file); my $id = $temp[0]; $id =~ s/.fastq//; my $log2 = $dir."/Read_Alignment_module_log_".$id.".txt"; open(LOG2,">$log2"); print LOG2 "Input Directory: $dir\n"; print LOG2 "Input Sequenced Reads .fastq file: $file\n\n\n"; my $indexfile = "PATH/TO/BOWTIEREFERENCE"; my $samfile = $file; $samfile =~ s/.fastq/.sam/; my $bamfile = $samfile; $bamfile =~ s/.sam/.bam/; my $bresult = $id."_bresult.txt"; print LOG2 "Index file is: $indexfile\n"; print LOG2 "Output file will be: $bamfile\n"; print LOG2 "Running BOWTIE issuing the command:"; my $bowtie_cmd = "bowtie -q --nomaqround -m 4 -n 2 -e 70 -l 28 --best --chunkmbs 224 -p 8 --seed=123 -S $indexfile $file $samfile"; print LOG2 "$bowtie_cmd\n"; my $start = time(); my $bowtie = `bowtie -q --nomaqround -m 4 -n 2 -e 70 -l 28 --best --chunkmbs 224 -p 8 --seed=123 -S $indexfile $file $samfile 2>> $bresult`; my $time = int((time() - $start)/60); print LOG2 "Finished running BOWTIE ($time min)\n\n\n"; print LOG2 "Converting $samfile to $bamfile using the command:\n"; my $sam_to_bam_cmd = "samtools view -bS $samfile > $bamfile"; print LOG2 "$sam_to_bam_cmd\n"; my $sam_to_bam = `samtools view -bS $samfile > $bamfile`; print LOG2 "Converted .sam file to .bam file\n\n\n"; my $r = `rm $samfile`; print LOG2 "Removed $samfile\n"; print LOG2 "END OF RUN\n\n\n"; print LOG2 "BOWTIE RESULT\n"; my $append_bresult = `cat $bresult >>$log2`; my $rm = `rm $bresult`; #### getAverageWig ####### #!/usr/bin/perl -w # this script generated by the ERCAN LAB at NYU # contact: se71@nyu.edu use strict; my @fHs; # contains all filehandles my $ident = pop(@ARGV); my $desc = $ident."_average"; my $outfile = $desc.".wig"; for (my $i=0; $i < scalar(@ARGV); $i++) { open($fHs[$i], "<", $ARGV[$i]) || die "Can't open inputfile $ARGV[$i]: $!.\n"; } open(OUT, ">", $outfile) || die "Can't open inputfile $outfile: $!.\n"; my $count = 0; while (0 == grep !defined, my @line = map scalar <$_>, @fHs) { map(chomp, @line); if ($line[0] =~ /^track/) { print STDERR "\nA: $line[0]\t"; print OUT "track type=wiggle_0 name=\"$desc\" description=\"$desc\" visibility=full autoScale=off windowingFunction=mean smoothingWindow=8 viewLimits=-2:5\n"; } elsif ($line[0] =~ /^fixedStep/) { foreach my $l (@line) { if ($l ne $line[0]) { print "Problem in line $. for $line[0].\n"; exit(1); } } print STDERR "\nA: $line[0]\t"; print OUT $line[0]."\n"; } elsif ($line[0] =~ /^\#/) { print "next: $. ?\n"; next; } else { my @vals; foreach my $l (@line) { push @vals, $l unless ($l eq " "); } if (scalar(@vals) < 2) { print OUT " \n"; } else { my $average = mean(\@vals); print OUT "$average\n"; } $count++; } if($count % 1000000 == 0) { print STDERR "$count."; } } print "\n"; close (OUT); sub mean { my $vals = shift(); my $sum = 0; foreach my $val (@$vals) { # print "val: $val\n"; $sum += $val; } # print "sum: $sum\n"; my $average = $sum/scalar(@$vals); # print "Average: $average\n"; return(sprintf "%.4f", $average); } ###### getLog2 #!/usr/bin/perl # this script generated by the ERCAN LAB at NYU # contact: se71@nyu.edu use strict; use warnings; use Statistics::Descriptive; my $file = shift(); my $Output = $file; $Output =~ s/.wig/_log2.wig/g; open (OUT, ">$Output") || die "Can't open output file $Output\n"; open(toDIV, $file) || die "Can't open inputfile $file\n"; my $count = 1; #set count scalar to one my $chr; while (my $todiv = ){ #declares todiv. goes through every line in the wiggle file chomp($todiv); my @data = split("\t",$todiv); my $n2 = $data[0]; my $mut = $data[1]; if ($n2 =~ /track/) { #if the line contains the word "track" print OUT $n2."\n"; #print the track name to the output file } elsif ($n2 =~ /fixedStep/) { if ($n2 =~ /chrom=(CHROMOSOME_[IVX]*) /) { print OUT $n2."\n"; #print the line to the output file $count = 1; #and reset the count to 1 $chr=$1; print STDERR "\n Processing $chr."; }} else { if ($n2 eq " ") { #if its blank, dont change it. print OUT " \n"; $count++; } elsif ($mut eq " ") { #if its blank, dont change it. print OUT " \n"; $count++; } else { my $log2 = sprintf "%.2f", (log($mut/$n2))/log(2); #will print the log2 value rounded to two places after the decimal point print OUT "$log2\n"; $count++; } } if($count % 1000000 == 0){print STDERR "$count.";} } close toDIV; close OUT; ##### getOverlappingPeaks_Replicates #!/usr/bin/perl -w # this script generated by the ERCAN LAB at NYU # contact: se71@nyu.edu use strict; my $input = $ARGV[0]; my $files = $ARGV[1]; my $outfile = $input; $outfile =~ s/.bed/_final.bed/; my @files = split(",", $files); my %cutoff = (2 => 2, 3 => 2, 4 => 3, 5 => 3, 6 => 4, 7 => 4); my %peaks; my %results; open (INPUT, "<", $input) || die "Couldn't open file $input: $!.\n"; while () { chomp(); my @line = split("\t", $_); unless ($results{$line[3]}) { $results{$line[3]} = $_; } else { print "Problem with peak $line[3]!\n"; } } close INPUT; my $output = "temp.txt"; foreach my $file (@files) { print "Comparing $input to $file...\n"; system("intersectBed -wa -u -a $input -b $file > $output"); open(INPUT, "<", $output) || die "Couldn't open intersect file: $!.\n"; while () { chomp(); my @line = split("\t", $_); if ($peaks{$line[3]}) { $peaks{$line[3]}++; } else { $peaks{$line[3]} = 1; } } close INPUT; } system("rm $output"); my $n_files = scalar(@files); my $cutoff = $cutoff{$n_files}; print "Number of files: $n_files\n"; print "Cutoff: $cutoff\n"; my $count = 0; open(OUTPUT, ">", $outfile) || die "Couldn't open file $outfile: $!.\n"; foreach my $peak (keys %peaks) { if ($peaks{$peak} >= $cutoff) { print OUTPUT $results{$peak}."\n"; $count++; } } close OUTPUT; print $count." overlapping peaks found in replicates.\n"; ##### macs_module_multiple ####### #!/usr/bin/perl -w # This script takes in treat.bam and its matching input (control.bam) files for multiple samples # performs MACS generating variable step wiggle files, converts it to fixed step start to end wiggle files # performs normalization using wignorm generating variable step normalized wiggle files, converts it to fixed step start to end wiggle files # usage perl macs_module.pl file containing treat.bam control.bam identifier # this script generated by the ERCAN LAB at NYU # contact: se71@nyu.edu use strict; use Cwd; my $mdir = getcwd; my $dir = getcwd; chdir($dir); my $treatbam = shift(); my $controlbam = shift(); my $identifier = shift(); print "Running MACS for $treatbam...\n"; my $log3 = $mdir."/MACS_module_log_$identifier.txt"; open (LOG3,">$log3"); print LOG3 "Input Directory: $mdir\n"; print LOG3 "Input treat.bam file: $treatbam\n"; print LOG3 "Input control.bam file: $controlbam\n"; print LOG3 "MACS output file identifier: $identifier\n\n\n"; my $output_dir = $treatbam; $output_dir =~ s/.bam//; mkdir($output_dir); chdir($output_dir); unless (($treatbam=~/.bam/) && ($controlbam=~/.bam/)) { print LOG3 "Error: Please enter Input files in .bam format\n"; exit(0); } if ($treatbam =~ /gz/) { my $cmd = "gzip -d $treatbam"; chomp($cmd); system($cmd); $treatbam =~ s/\.gz//g; print LOG3 "Unzipped file: $treatbam\n"; } if ($controlbam =~ /gz/) { my $cmd = "gzip -d $controlbam"; chomp($cmd); system($cmd); $controlbam =~ s/\.gz//g; print LOG3 "Unzipped file: $controlbam\n"; } mkdir("MACS_pval1e-10") || print $!; mkdir("MACS_pval1e-5") || print $!; my $macs5folder = $dir."/".$output_dir."/MACS_pval1e-5"; my $macs10folder = $dir."/".$output_dir."/MACS_pval1e-10"; chdir($macs5folder)|| print $!; print LOG3 "In folder $macs5folder to run MACS at p-value 1e-5 and generate treat and control variable step wiggle files \n"; print LOG3 "Running MACS1.4 issuing the command:"; my $macs5_cmd = "macs14 -t $treatbam -c $controlbam -n MACS141_e5_$identifier -g ce -f BAM -w -S --space=1"; print LOG3 "$macs5_cmd\n"; # the following line is the command line running macs e-5 is default, thus is not indicated. my $macs5 = `macs14 -t $dir/$treatbam -c $dir/$controlbam -n MACS141_e5_$identifier -g ce -f BAM -w -S --space=1`; print LOG3 "Finished running MACS1.4 at p-value 1e-5\n\n"; chdir($macs10folder)|| print $!; print LOG3 "In folder $macs10folder to run MACS at p-value 1e-10\n"; print LOG3 "Running MACS1.4 issuing the command:"; my $macs10_cmd = "macs14 -t $treatbam -c $controlbam -n MACS141_e10_$identifier -g ce -f BAM -p 1e-10"; print LOG3 "$macs10_cmd\n"; my $macs10 = `macs14 -t $dir/$treatbam -c $dir/$controlbam -n MACS141_e10_$identifier -g ce -f BAM -p 1e-10`; print LOG3 "Finished running MACS1.4 at p-value 1e-10\n\n\n"; ### wignorm chdir($mdir."/".$output_dir)|| print $!; print LOG3 "In folder $mdir/$output_dir to perform normalization using WIGNORM\n"; print LOG3 "Unzipping the wiggle files from MACS1.4\n"; my $treatwig = $macs5folder."/MACS141_e5_".$identifier."_MACS_wiggle/treat"."/MACS141_e5_".$identifier."_treat_afterfiting_all.wig.gz"; my $cmd1 = "gzip -d $treatwig"; chomp($cmd1); system($cmd1); $treatwig =~ s/\.gz//g; print LOG3 "\nUnzipped treat wiggle file: $treatwig"; #$controlwig=$mdir."/MACS_pval1e-5/MACS5_MACS_wiggle/control/MACS5_control_afterfiting_all.wig.gz"; my $controlwig = $macs5folder."/MACS141_e5_".$identifier."_MACS_wiggle/control"."/MACS141_e5_".$identifier."_control_afterfiting_all.wig.gz"; my $cmd2 = "gzip -d $controlwig"; chomp($cmd2); system($cmd2); $controlwig =~ s/\.gz//g; print LOG3 "\nUnzipped control wiggle file: $controlwig\n"; my $wignorm_cmd = "wignorm -t $treatwig -c $controlwig -n treat_control"; print LOG3 "Running wignorm issuing the command: $wignorm_cmd\n"; my $wignorm = `wignorm -t $treatwig -c $controlwig -n $identifier"_wignorm"`; print LOG3 "Finished running wignorm\n\n\n"; # my $rm_scores_bed = `rm $mdir/$output_dir/treat_control_wignorm_peaks.bed`; ### convert the variable step wiggle files to fixed step wiggle files my $wigfile_t = $macs5folder."/MACS141_e5_".$identifier."_MACS_wiggle/treat"."/MACS141_e5_".$identifier."_treat_afterfiting_all.wig"; my $wigfile_c = $macs5folder."/MACS141_e5_".$identifier."_MACS_wiggle/control"."/MACS141_e5_".$identifier."_control_afterfiting_all.wig"; my $wigfile = $mdir."/".$output_dir."/".$identifier."_wignorm_scores.wig"; my @wfiles = ($wigfile_t,$wigfile_c,$wigfile); my %chr_len = ("CHROMOSOME_I" => 15072423, "CHROMOSOME_II" => 15279345, "CHROMOSOME_III" => 13783700, "CHROMOSOME_IV" => 17493793, "CHROMOSOME_MtDNA" => 13794, "CHROMOSOME_V" => 20924149, "CHROMOSOME_X" => 17718866, "plasmid+rexwithSNPs" => 7763); foreach my $f (@wfiles) { print LOG3 "Converting variable step wiggle file $f to fixed step wiggle file\n"; my $end = 0; my $cnt = 1; my $flag = 0; my $out = $f; $out =~ s/\.wig/\_FIXED\.wig/g; open (OUT,">$out"); my $currchrom = "none"; # print STDERR "\nProcessing wig file $f.\nPrinting output to $out.\n"; open (WIG,$f); while () { chomp(); if ($flag == 0) { if ($_ =~ 'track') { print OUT "$_ windowingFunction=mean smoothingWindow=8 viewLimits=-2:5\n"; } if ($_ =~ 'variableStep') { my ($variable,$chrom,$span) = split(" ",$_); $chrom =~ s/chrom\=//g; $span =~ s/span\=//; if($currchrom ne $chrom) { # print STDERR "\tProcessing chrom $chrom\n"; $currchrom = $chrom; $end = $chr_len{$currchrom}; print OUT "fixedStep chrom=$chrom start=1 step=1 span=$span\n"; $flag=1; } } } else { my ($pos,$val) = split(" ",$_); if($pos ne 'variableStep') { if ($cnt <= $pos && $pos <= $end) { for (my $i = $cnt; $i < $pos; $i++) { print OUT " \n"; $cnt++; } if ($pos <= $end) { print OUT $val; print OUT "\n"; $cnt++; } } } else { for (my $i = $cnt; $i <= $end; $i++) { print OUT " \n"; } $cnt= 1; my ($variable,$chrom,$span) = split(" ", $_); $chrom =~ s/chrom\=//g; $span =~ s/span\=//; if ($currchrom ne $chrom) { # print STDERR "\tProcessing chrom $chrom\n"; $currchrom = $chrom; $end = $chr_len{$currchrom}; print OUT "fixedStep chrom=$chrom start=1 step=1 span=$span\n"; $flag = 1; } } } } # end of while for (my $i = $cnt; $i <= $end; $i++) { print OUT " \n"; } print LOG3 "Check fixed step normalized wiggle file: $out\n"; my $cmd = `gzip $f`; print LOG3 "Gzipped file: $f\n"; $cmd = `gzip $out`; print LOG3 "Gzipped file: $out\n\n\n"; } ### Check peak bed file for peak regions smaller than chromosome start or larger than chromosome length and move these to 0/chromsome length my @bedfiles = ($macs5folder."/MACS141_e5_".$identifier."_peaks.bed", $macs10folder."/MACS141_e10_".$identifier."_peaks.bed", $mdir."/".$output_dir."/".$identifier."_wignorm_peaks.bed"); for my $bedfile_cur (@bedfiles) { my $bedfile_new = $bedfile_cur; $bedfile_new =~ s/.bed/_corrected.bed/; print "Total number of peaks in $bedfile_cur:\n"; system("wc -l $bedfile_cur"); print LOG3 "Checking peak bed file $bedfile_cur for annotations outside the chromosome regions.\n"; open (INPUT, "<", $bedfile_cur) || print "$!\n"; open (OUTPUT, ">", $bedfile_new) || print "$!\n"; while () { chomp(); my ($chr, $start, $end, $peak, $value) = split("\t", $_); $start = 0 if ($start < 0); $end = $chr_len{$chr} if ($end > $chr_len{$chr}); if (defined($peak) && defined($value)) { print OUTPUT "$chr\t$start\t$end\t$peak\t$value\n"; } else { print OUTPUT "$chr\t$start\t$end\n"; } } close (INPUT); close (OUTPUT); system("rm $bedfile_cur"); rename($bedfile_new, $bedfile_cur); } ###### macs_module_replicates ####### #!/usr/bin/perl -w # This script takes in treat.bam and its matching input (control.bam) files for multiple samples # performs MACS generating variable step wiggle files, converts it to fixed step start to end wiggle files # performs normalization using wignorm generating variable step normalized wiggle files, converts it to fixed step start to end wiggle files # usage perl macs_module.pl file containing treat.bam control.bam identifier # this script generated by the ERCAN LAB at NYU # contact: se71@nyu.edu use strict; use Cwd; my $mdir = getcwd; my $dir = getcwd; #my $file = shift; #open (INFILE, "<", $file) || die "Couldn't open file $file: $!.\n"; #while () { # chomp(); chdir($dir); # my ($treatbam, $controlbam, $identifier) = split("\t", $_); my $treatbam = shift(); my $controlbam = shift(); my $identifier = shift(); print "Running MACS for $treatbam...\n"; my $log3 = $mdir."/MACS_module_log_$identifier.txt"; open (LOG3,">$log3"); print LOG3 "Input Directory: $mdir\n"; print LOG3 "Input treat.bam file: $treatbam\n"; print LOG3 "Input control.bam file: $controlbam\n"; print LOG3 "MACS output file identifier: $identifier\n\n\n"; my $output_dir = $treatbam; $output_dir =~ s/.bam//; mkdir($output_dir); chdir($output_dir); unless (($treatbam=~/.bam/) && ($controlbam=~/.bam/)) { print LOG3 "Error: Please enter Input files in .bam format\n"; exit(0); } if ($treatbam =~ /gz/) { my $cmd = "gzip -d $treatbam"; chomp($cmd); system($cmd); $treatbam =~ s/\.gz//g; print LOG3 "Unzipped file: $treatbam\n"; } if ($controlbam =~ /gz/) { my $cmd = "gzip -d $controlbam"; chomp($cmd); system($cmd); $controlbam =~ s/\.gz//g; print LOG3 "Unzipped file: $controlbam\n"; } mkdir("MACS_pval1e-10") || print $!; mkdir("MACS_pval1e-5") || print $!; my $macs5folder = $dir."/".$output_dir."/MACS_pval1e-5"; my $macs10folder = $dir."/".$output_dir."/MACS_pval1e-10"; chdir($macs5folder)|| print $!; print LOG3 "In folder $macs5folder to run MACS at p-value 1e-5 and generate treat and control variable step wiggle files \n"; print LOG3 "Running MACS1.4 issuing the command:"; my $macs5_cmd = "macs14 -t $treatbam -c $controlbam -n MACS141_e5_$identifier -g ce -f BAM -w -S --space=1"; print LOG3 "$macs5_cmd\n"; # the following line is the command line running macs e-5 is default, thus is not indicated. my $macs5 = `macs14 -t $dir/$treatbam -c $dir/$controlbam -n MACS141_e5_$identifier -g ce -f BAM -w -S --space=1`; print LOG3 "Finished running MACS1.4 at p-value 1e-5\n\n"; chdir($macs10folder)|| print $!; print LOG3 "In folder $macs10folder to run MACS at p-value 1e-10\n"; print LOG3 "Running MACS1.4 issuing the command:"; my $macs10_cmd = "macs14 -t $treatbam -c $controlbam -n MACS141_e10_$identifier -g ce -f BAM -p 1e-10"; print LOG3 "$macs10_cmd\n"; my $macs10 = `macs14 -t $dir/$treatbam -c $dir/$controlbam -n MACS141_e10_$identifier -g ce -f BAM -p 1e-10`; print LOG3 "Finished running MACS1.4 at p-value 1e-10\n\n\n"; ### wignorm chdir($mdir."/".$output_dir)|| print $!; print LOG3 "In folder $mdir/$output_dir to perform normalization using WIGNORM\n"; print LOG3 "Unzipping the wiggle files from MACS1.4\n"; my $treatwig = $macs5folder."/MACS141_e5_".$identifier."_MACS_wiggle/treat"."/MACS141_e5_".$identifier."_treat_afterfiting_all.wig.gz"; my $cmd1 = "gzip -d $treatwig"; chomp($cmd1); system($cmd1); $treatwig =~ s/\.gz//g; print LOG3 "\nUnzipped treat wiggle file: $treatwig"; #$controlwig=$mdir."/MACS_pval1e-5/MACS5_MACS_wiggle/control/MACS5_control_afterfiting_all.wig.gz"; my $controlwig = $macs5folder."/MACS141_e5_".$identifier."_MACS_wiggle/control"."/MACS141_e5_".$identifier."_control_afterfiting_all.wig.gz"; my $cmd2 = "gzip -d $controlwig"; chomp($cmd2); system($cmd2); $controlwig =~ s/\.gz//g; print LOG3 "\nUnzipped control wiggle file: $controlwig\n"; ### convert the variable step wiggle files to fixed step wiggle files my $wigfile_t = $macs5folder."/MACS141_e5_".$identifier."_MACS_wiggle/treat"."/MACS141_e5_".$identifier."_treat_afterfiting_all.wig"; my $wigfile_c = $macs5folder."/MACS141_e5_".$identifier."_MACS_wiggle/control"."/MACS141_e5_".$identifier."_control_afterfiting_all.wig"; my @wfiles = ($wigfile_t,$wigfile_c); # print "@wfiles"; #my @chr = ("chrI","chrII","chrIII","chrIV","chrM","chrV","chrX"); #my @chr_len = (15072421,15279323,13783681,17493785,13794,20919568,17718854); # (ce6) my %chr_len = ("chrI" => 15072421, "chrII" => 15279323, "chrIII" => 13783681, "chrIV" => 17493785, "chrM" => 13794, "chrV" => 20919568, "chrX" => 17718854); my %chr_len = ("CHROMOSOME_I" => 15072423, "CHROMOSOME_II" => 15279345, "CHROMOSOME_III" => 13783700, "CHROMOSOME_IV" => 17493793, "CHROMOSOME_MtDNA" => 13794, "CHROMOSOME_V" => 20924149, "CHROMOSOME_X" => 17718866, "plasmid+rexwithSNPs" => 7763); foreach my $f (@wfiles) { print LOG3 "Converting variable step wiggle file $f to fixed step wiggle file\n"; my $end = 0; my $cnt = 1; my $flag = 0; my $out = $f; $out =~ s/\.wig/\_FIXED\.wig/g; open (OUT,">$out"); my $currchrom = "none"; # print STDERR "\nProcessing wig file $f.\nPrinting output to $out.\n"; open (WIG,$f); while () { chomp(); if ($flag == 0) { if ($_ =~ 'track') { print OUT "$_ windowingFunction=mean smoothingWindow=8 viewLimits=-2:5\n"; } if ($_ =~ 'variableStep') { my ($variable,$chrom,$span) = split(" ",$_); $chrom =~ s/chrom\=//g; $span =~ s/span\=//; if($currchrom ne $chrom) { # print STDERR "\tProcessing chrom $chrom\n"; $currchrom = $chrom; $end = $chr_len{$currchrom}; print OUT "fixedStep chrom=$chrom start=1 step=1 span=$span\n"; $flag=1; } } } else { my ($pos,$val) = split(" ",$_); #print "$pos\t"; #print "$val\n"; if($pos ne 'variableStep') { if ($cnt <= $pos && $pos <= $end) { for (my $i = $cnt; $i < $pos; $i++) { #print OUT $cnt; #print OUT "\t"; print OUT " \n"; $cnt++; } if ($pos <= $end) { #print OUT $pos; #print OUT "\t"; print OUT $val; print OUT "\n"; $cnt++; } } } else { for (my $i = $cnt; $i <= $end; $i++) { #print OUT $i; #print OUT "\t"; print OUT " \n"; } $cnt= 1; my ($variable,$chrom,$span) = split(" ", $_); $chrom =~ s/chrom\=//g; $span =~ s/span\=//; if ($currchrom ne $chrom) { # print STDERR "\tProcessing chrom $chrom\n"; $currchrom = $chrom; $end = $chr_len{$currchrom}; print OUT "fixedStep chrom=$chrom start=1 step=1 span=$span\n"; $flag = 1; } } } } # end of while for (my $i = $cnt; $i <= $end; $i++) { #print OUT $i; #print OUT "\t"; print OUT " \n"; } print LOG3 "Check fixed step normalized wiggle file: $out\n"; my $cmd = `gzip $f`; print LOG3 "Gzipped file: $f\n"; #uncomment this to delete the macs generated variable step wiggle files and wignorm generated normalized variable step wiggle file #$r=`rm $wigfile_t`; #print LOG3 "Removed file: $wigfile_t\n; #$r=`rm $wigfile_c`; #print LOG3 "Removed file: $wigfile_c\n; #$r=`rm $wigfile`; #print LOG3 "Removed file: $wigfile\n; $cmd = `gzip $out`; print LOG3 "Gzipped file: $out\n\n\n"; } ### Check peak bed file for peak regions smaller than chromosome start or larger than chromosome length and move these to 0/chromsome length my @bedfiles = ($macs5folder."/MACS141_e5_".$identifier."_peaks.bed", $macs10folder."/MACS141_e10_".$identifier."_peaks.bed"); for my $bedfile_cur (@bedfiles) { my $bedfile_new = $bedfile_cur; $bedfile_new =~ s/.bed/_corrected.bed/; print "Total number of peaks in $bedfile_cur:\n"; system("wc -l $bedfile_cur"); print LOG3 "Checking peak bed file $bedfile_cur for annotations outside the chromosome regions.\n"; open (INPUT, "<", $bedfile_cur) || print "$!\n"; open (OUTPUT, ">", $bedfile_new) || print "$!\n"; while () { chomp(); my ($chr, $start, $end, $peak, $value) = split("\t", $_); $start = 0 if ($start < 0); $end = $chr_len{$chr} if ($end > $chr_len{$chr}); if (defined($peak) && defined($value)) { print OUTPUT "$chr\t$start\t$end\t$peak\t$value\n"; } else { print OUTPUT "$chr\t$start\t$end\n"; } } close (INPUT); close (OUTPUT); system("rm $bedfile_cur"); rename($bedfile_new, $bedfile_cur); } #$cmd=`gzip $treatbam`; #print LOG3 "Gzipped file: $treatbam\n"; #$cmd=`gzip $controlbam`; #print LOG3 "Gzipped file: $controlbam\n"; #} #close INFILE; ###### mergeMultipleBam ####### #!/usr/bin/perl -w # this script generated by the ERCAN LAB at NYU # contact: se71@nyu.edu use strict; my $input = shift(); open (INPUT, "<", $input) || die "Couldn't open file $input: $!.\n"; while () { chomp(); print "Running samtools merge for $_...\n"; system("samtools merge $_"); } close INPUT; ###### sortBam ####### #!/usr/bin/perl -w # this script generated by the ERCAN LAB at NYU # contact: se71@nyu.edu use strict; my $file = shift(); my $out = $file; $out =~ s/.bam/_sorted/; print "Sorting file $file...\n"; system("samtools sort $file $out"); ######################################################################################################################################################### ####This section will run through the steps detailing how we do sliding window analysis of DCC binding difference between rex deletion and wild type data. #Slidingwindow_Analysis.R contains both the linux and R commands used to generated the sliding window plots shown in Figure 7A of this manuscript. #This analysis requires the Statistics::Descriptive perl module. #Analysis in R is dependent on two provided annotation files: #(1) Recruitment.Sites.txt #Provides the peak names and coordinates of each of the recruitment sites. The file content is pasted below. #(2) MACS141_e10_DPY27_N2_MxEmb_splitted_summits_final.bed #Provides the coordinates and peak names for the DPY-27 peaks.The file content is pasted below. ###################### ###################### #### GET LOG2 DATA ### ###################### ###################### ### DO THE FOLLOWING ON THE COMMAND LINE ### Required as input: the MACS e-5 FIXED_STEP wiggle files ### Requires running the perl script: getLog2.pl The script is pasted below. # This perl script requires the latest version of perl, is dependent on the Statistics::Descriptive module # 1. Paste together the N2 wiggle file and a file to compare it to. ## Every line will have two entries: (1) the N2 (wt control) value (2) the experimental strain value ## Example: # paste MACS141_e5_DPY27_N2_MxEmb_treat_afterfiting_all_FIXED.wig MACS141_e5_MUT_emb_DPY27_treat_afterfiting_all_FIXED.wig > DPY27_N2_MUT_emb.wig # 2. Calculate the log2 (exp/N2) value at every step. ## Will generate a new wiggle file, appending _log2.wig to the end of the parent file name. ## Example: # perl getLog2.pl DPY27_N2_MUT_emb.wig # 3. Separate log2 data by chromosome. ## 7A. Use awk to pull out autosomal data. ## Example: # awk '/CHROMOSOME_II /{f=0} f; /CHROMOSOME_I /{f=1}' DPY27_N2_MUT_emb.wig > ChrI_DPY27_N2_MUT_emb.wig # awk '/CHROMOSOME_III /{f=0} f; /CHROMOSOME_II /{f=1}' DPY27_N2_MUT_emb.wig > ChrII_DPY27_N2_MUT_emb.wig # awk '/CHROMOSOME_IV /{f=0} f; /CHROMOSOME_III /{f=1}' DPY27_N2_MUT_emb.wig > ChrIII_DPY27_N2_MUT_emb.wig # awk '/CHROMOSOME_V /{f=0} f; /CHROMOSOME_IV /{f=1}' DPY27_N2_MUT_emb.wig > ChrIV_DPY27_N2_MUT_emb.wig # awk '/CHROMOSOME_X /{f=0} f; /CHROMOSOME_V /{f=1}' DPY27_N2_MUT_emb.wig > ChrV_DPY27_N2_MUT_emb.wig ## 7B. Use sed to pull out X chromosome data. ## Example: # sed -e '1,/CHROMOSOME_X /d' DPY27_N2_MUT_emb.wig > ChrX_DPY27_N2_MUT_emb.wig ######################## ######################## #### WINDOW ANALYSIS ### ######################## ######################## ### DO THE FOLLOWING in R # 1. Read in annotation files rex<-read.table(file=‘Recruitment.Sites.txt’,header=T,stringsAsFactors=F) DPY27.summits<-read.table(file='MACS141_e10_DPY27_N2_MxEmb_splitted_summits_final.bed',header=F) colnames(DPY27.summits)<-c('chr','start','end','peak') # 2. Read in data files for each chromosome and assemble chromosome data frames # Example: ### Chromosome X ### N2_MUT1_DPY27_ChrX<-as.data.frame(read.delim(file='ChrX_DPY27_N2_MUT1_emb_log2.wig',header=F,stringsAsFactors=F)) N2_MUT2_DPY27_ChrX<-as.data.frame(read.delim(file=‘ChrX_DPY27_N2_MUT2_emb_log2.wig’,header=F,stringsAsFactors=F)) N2_MUT3_DPY27_ChrX<-as.data.frame(read.delim(file=‘ChrX_DPY27_N2_MUT3_emb_log2.wig’,header=F,stringsAsFactors=F)) chrX<-data.frame(N2_MUT1_DPY27=N2_MUT1_DPY27_ChrX, N2_MUT2_DPY27=N2_MUT2_DPY27_ChrX, N2_MUT3_DPY27=N2_MUT3_DPY27_ChrX, ) colnames(chrX)<-c('N2_MUT1_DPY27','N2_MUT2_DPY27','N2_MUT3_DPY27') # 3. Get the mean value (na.rm) for a 400bp window centered on each DPY-27 summit. ChrX.DPY27.summit.data<-as.data.frame(cbind(DPY27.summits[which(DPY27.summits$chr=='CHROMOSOME_X'),],t(apply(DPY27.summits[which(DPY27.summits$chr=='CHROMOSOME_X'),],1,function(x){colMeans(chrX[(as.numeric(x[2])-200) :(as.numeric(x[3])+200) ,],na.rm=T)})))) colnames(ChrX.DPY27.summit.data)<-c('chr','start','end','peak','N2_MUT1','N2_MUT2’,’N2_MUT3’) # 4. Sliding window using only the 400bp averaged summit data. ### Note, if you have deleted/added a recruitment site, you need to remove this site from the analysis. # Example: recsitetoremove.ind<-which( ChrX.DPY27.summit.data$peak == rex$dpy27.peak[which(rex$meyer.name == ‘(recruitment site to remove)’)]) MUT1.DPY27.X.data<-ChrX.DPY27.summit.data[-recsitetoremove.ind , c(1,2,3,4,5)] # The following will return window data across the X chromosome: window size = 5kb, step size = 1kb, data=summit means get.window.data.from.summits<-function(window,step,df,data.column){ X.length<-17718866 i<-1 j<-1 pos <- matrix(0, ncol=2, nrow=ceiling(X.length/step)) while ((i+window) < X.length) { pos[j, ] <- c(i, i+window) i <- i + step j <- j+1 } while(j <= nrow(pos)){ pos[j, ] <- c(i, X.length) i <- i + step j <- j+1 } ranges<-as.data.frame(pos) colnames(ranges)<-c('start','end') n.summits<-c() mean.summits<-c() for(i in 1:nrow(ranges)){ start<-ranges$start[i] end<-ranges$end[i] temp<-df[(which((df$start > start & df$start < end ) | (df$end > start & df$end < end))),] n.summits[i]<-nrow(temp) mean.summits[i]<-mean(temp[,as.numeric(data.column)],na.rm=T) } ranges$n.summits<-n.summits ranges$mean.summits<-mean.summits return(ranges) } # Decide which recruitment site to use for reference. rex8.dpy27.peak<-rex$dpy27.peak[which(rex$meyer.name=='rex-8')] #strongest recruitment site/ 3 motifs: 10.9.5,8.25 rex32.dpy27.peak<-rex$dpy27.peak[which(rex$meyer.name=='rex-32')] #left of the chromosome (3Mb)/2ndstrongest recruitment site/3motifs: 10,9.5,9.5 rex33.dpy27.peak<-rex$dpy27.peak[which(rex$meyer.name=='rex-33')] #middle of the chromosome (6.3Mb)/3rd strongest recruitment site/3motifs: 10,7.5,7.5 get.pvalue.summits<-function(window,df,data.column,ref){ pval.summits<-c() #start.coord<-c() #end.coord<-c() #dist<-c() n.summits<-c() mean.summits<-c() anti.pval<-c() for(i in 1:nrow(df)){ start<-df$start[i] - window/2 end<-df$end[i] + window/2 if(start <= 0){start<-1} if(end > max(df$end)){end<-max(df$end)} ind<-which((df$start > start & df$start < end) | (df$end > start & df$end < end)) n.summits[i]<-length(ind) if(length(ind) > 0){ in.range<-df[ind,data.column] all<-df[,data.column] pval.summits[i]<-t.test(in.range,all,alternative='less')$p.value anti.pval[i]<-t.test(in.range,all,alternative='greater')$p.value #start.coord[i]<-df$start[start] #end.coord[i]<-df$end[end] #dist[i]<-end.coord[i]-start.coord[i] mean.summits[i]<-mean(in.range,na.rm=T) } else{ pval.summits[i]<-1 anti.pval[i]<-1 mean.summits[i]<-df[i,data.column] } } #df$start.coord<-start.coord #df$end.coord<-end.coord #df$dist<-dist df$mean.summits<-mean.summits df$pval.summits<-pval.summits df$anti.pval<-anti.pval df$n.summits<-n.summits if(ref=='rex-8'){rexref<-as.character(rex8.dpy27.peak)} if(ref=='rex-32'){rexref <-as.character(rex32.dpy27.peak)} if(ref=='rex-33'){rexref <-as.character(rex33.dpy27.peak)} RF<-df$mean.summits[which(df$peak==rexref)] df$dev<-(df$mean.summits - RF )/abs(RF) return(df) } # Example: MUT1_DPY27_summit_pval_Xchr_1Mb<-get.pvalue.summits(1.0e6,MUT1.DPY27.X.data,5,'rex-32') plot(MUT1_DPY27_summit_pval_Xchr_1Mb$start, MUT1_DPY27_summit_pval_Xchr_1Mb $dev,type='l',lwd=5, ylim=c(-1.1,0.25)) points(MUT1_DPY27_summit_pval_Xchr_1Mb$start[which(MUT1_DPY27_summit_pval_Xchr_1Mb$pval.summits<0.01)], MUT1_DPY27_summit_pval_Xchr_1Mb$dev[which(MUT1_DPY27_summit_pval_Xchr_1Mb$pval.summits<0.01)],col='red',pch=16) points(MUT1_DPY27_summit_pval_Xchr_1Mb$start[which(MUT1_DPY27_summit_pval_Xchr_1Mb$anti.pval<0.01)], MUT1_DPY27_summit_pval_Xchr_1Mb$dev[which(MUT1_DPY27_summit_pval_Xchr_1Mb$anti.pval<0.01)],col='purple',pch=16) # 5. Get window data along entire chromosome get.window.data.across.chr<-function(window,step,df){ start<-seq(1,nrow(df)-window, step) end<-seq((window + 1),nrow(df), step) ranges<-data.frame(start=start,end=end) old <- Sys.time() temp<-as.data.frame(t(as.matrix(apply(ranges,1,function(x){colMeans(df[x[1]:x[2],],na.rm=T)})))) new <- Sys.time() - old print(new) temp<-as.data.frame(cbind(ranges,temp)) return(temp) } chrX.window.data<-get.window.data.across.chr(50000,5000,chrX) plot(chrX.window.data$start,chrX.window.data$N2_MUT1_DPY27,type='l',bty='n',xlim=c(0,22e6),col='purple4',xaxt='n',xlab='coordinate (Mb)') axis(1,at=c(5e6,10e6,15e6,20e5),c('5','10','15','20')) ######################################################################################################################################################### ####getLog2.pl #!/usr/bin/perl # this script generated by the ERCAN LAB at NYU # contact: se71@nyu.edu use strict; #insists that all variables are declared use warnings; #finds problems in your code and tells you about them use Statistics::Descriptive; #uses that perl package my $file = shift(); my $Output = $file; $Output =~ s/.wig/_log2.wig/g; #renames the output file open (OUT, ">$Output") || die "Can't open output file $Output\n"; #prints stdout to the output file open(toDIV, $file) || die "Can't open inputfile $file\n"; #opens the file and now refers to it as toDIV my $count = 1; #set count scalar to one my $chr; while (my $todiv = ){ #declares todiv. goes through every line in the wiggle file chomp($todiv); #get rid of new line my @data = split("\t",$todiv); my $n2 = $data[0]; my $mut = $data[1]; if ($n2 =~ /track/) { #if the line contains the word "track" print OUT $n2."\n"; #print the track name to the output file } elsif ($n2 =~ /fixedStep/) { #if the line contains "fixedStep" if ($n2 =~ /chrom=(CHROMOSOME_[IVX]*) /) { #if it also contains chrom = CHROMOSOME_I through X print OUT $n2."\n"; #print the line to the output file $count = 1; #and reset the count to 1 $chr=$1; print STDERR "\n Processing $chr."; }} else { if ($n2 eq " ") { #if its blank, dont change it. print OUT " \n"; $count++; } elsif ($mut eq " ") { #if its blank, dont change it. print OUT " \n"; $count++; } else { my $log2 = sprintf "%.2f", (log($mut/$n2))/log(2); #will print the log2 value rounded to two places after the decimal point print OUT "$log2\n"; $count++; } } if($count % 1000000 == 0){print STDERR "$count.";} } close toDIV; close OUT; ######################################################################################################################################################### ####Recruitment.Sites.txt chr start end peak meyer.name rank sdc.sort k.clust k.order foci n.m12 max12.mot m12.score sdc2.peak dpy27.peak hot trna nc.rna CHROMOSOME_X 11093935 11094336 MACS_peak_959.1 rex-8 1 4 1 1 11094135 3 10 10*9.5*8.25 MACS_peak_959.1 MACS_peak_1926.5 275 NA NA CHROMOSOME_X 4209060 4209461 MACS_peak_792.2 rex-23 8 3 1 2 4209260 2 10 10*9.5 MACS_peak_792.2 MACS_peak_776.2 252 NA NA CHROMOSOME_X 13700596 13700997 MACS_peak_1039 rex-43 10 8 1 3 13700796 0 0 NA MACS_peak_1039 MACS_peak_2360.3 281 NA NA CHROMOSOME_X 6296315 6296716 MACS_peak_857.2 rex-33 3 6 1 4 6296515 3 10 7.25*7.25*10 MACS_peak_857.2 MACS_peak_1146.5 258 NA NA CHROMOSOME_X 806476 806877 MACS_peak_690.2 rex-40 4 1 1 5 806676 3 9.25 7.25*9.25*8.75 MACS_peak_690.2 MACS_peak_142.2 241 NA NA CHROMOSOME_X 2996890 2997291 MACS_peak_770 rex-32 2 2 1 6 2997090 3 10 9.5*10*10 MACS_peak_770 MACS_peak_560.2 249 NA NA CHROMOSOME_X 5429305 5429706 MACS_peak_829 rex-34 5 13 1 7 5429505 2 8.5 7.5*8.5 MACS_peak_829 MACS_peak_986.3 253 NA NA CHROMOSOME_X 14525613 14526014 MACS_peak_1069 Prex-28 6 7 1 8 14525813 2 10 10*9.5 MACS_peak_1069 MACS_peak_2509.6 282 NA NA CHROMOSOME_X 11937263 11937664 MACS_peak_982.2 rex-16 12 9 1 9 11937463 1 8.25 8.25 MACS_peak_982.2 MACS_peak_2066.5 277 NA NA CHROMOSOME_X 16681821 16682222 MACS_peak_1132 rex-35 7 15 1 10 16682021 2 10 7.25*10 MACS_peak_1132 MACS_peak_2921.1 292 NA NA CHROMOSOME_X 8036178 8036579 MACS_peak_891.1 rex-14 11 10 1 11 8036378 2 10 10*10 MACS_peak_891.1 MACS_peak_1428.8 266 358 5714 CHROMOSOME_X 17544270 17544671 MACS_peak_1156.1 rex-41 18 12 1 12 17544470 2 8.25 8.25*5.25 MACS_peak_1156.1 MACS_peak_3062.2 NA NA NA CHROMOSOME_X 9465602 9466003 MACS_peak_923.1 rex-47 13 11 1 13 9465802 1 6.75 6.75 MACS_peak_923.1 MACS_peak_1648.2 269 NA NA CHROMOSOME_X 1626915 1627316 MACS_peak_734 Prex-7 9 5 1 14 1627115 3 9.5 6.25*9.5*5.25 MACS_peak_734 MACS_peak_302 246 NA NA CHROMOSOME_X 16056339 16056740 MACS_peak_1113 Prex-30 16 16 1 15 16056539 0 0 NA MACS_peak_1113 MACS_peak_2789.5 290 NA NA CHROMOSOME_X 1344804 1345205 MACS_peak_723 rex-45 14 17 1 16 1345004 1 7.25 7.25 MACS_peak_723 MACS_peak_256.2 243 NA NA CHROMOSOME_X 12362906 12363307 MACS_peak_994 rex-6 15 14 1 17 12363106 1 5.75 5.75 MACS_peak_994 MACS_peak_2135.1 278 NA NA CHROMOSOME_X 1909369 1909770 MACS_peak_740.2 rex-2 21 19 2 18 1909569 0 0 NA MACS_peak_740.2 MACS_peak_343.3 NA NA NA CHROMOSOME_X 7181565 7181966 MACS_peak_868 rex-24 17 22 2 19 7181765 1 6.25 6.25 MACS_peak_868 MACS_peak_1291.2 NA NA NA CHROMOSOME_X 17545444 17545845 MACS_peak_1156.2 NA 28 21 2 20 17545644 2 8.25 5.25*8.25 MACS_peak_1156.2 MACS_peak_3062.3 NA NA NA CHROMOSOME_X 17181178 17181579 MACS_peak_1145 rex-42 19 18 2 21 17181378 1 7.75 7.75 MACS_peak_1145 MACS_peak_3000 NA NA NA CHROMOSOME_X 13514312 13514713 MACS_peak_1027.2 Prex-22 26 24 2 22 13514512 1 10 10 MACS_peak_1027.2 MACS_peak_2324.2 NA NA NA CHROMOSOME_X 4395419 4395820 MACS_peak_797 rex-1 20 23 2 23 4395619 1 6.5 6.5 MACS_peak_797 MACS_peak_808.2 NA NA NA CHROMOSOME_X 16205731 16206132 MACS_peak_1119 Prex-31 23 20 2 24 16205931 0 0 NA MACS_peak_1119 MACS_peak_2822.3 NA NA NA CHROMOSOME_X 1322970 1323371 MACS_peak_721 rex-44 22 25 2 25 1323170 0 0 NA MACS_peak_721 MACS_peak_252 NA NA NA CHROMOSOME_X 409988 410389 MACS_peak_680 Prex-1 25 27 2 26 410188 0 0 NA MACS_peak_680 MACS_peak_76 238 515 NA CHROMOSOME_X 1389368 1389769 MACS_peak_726.4 Prex-6 30 30 2 27 1389568 0 0 NA MACS_peak_726.4 MACS_peak_262.4 NA NA NA CHROMOSOME_X 14813290 14813691 MACS_peak_1076 rex-39 29 28 2 28 14813490 0 0 NA MACS_peak_1076 MACS_peak_2563.2 NA NA NA CHROMOSOME_X 1888396 1888797 MACS_peak_739 rex-21 31 26 2 29 1888596 0 0 NA MACS_peak_739 MACS_peak_340 NA NA NA CHROMOSOME_X 15736382 15736783 MACS_peak_1103 rex-46 24 29 2 30 15736582 2 7.75 7.75*5.25 MACS_peak_1103 MACS_peak_2741.3 NA NA NA CHROMOSOME_X 16209616 16210017 MACS_peak_1120 NA 36 31 2 31 16209816 0 0 NA MACS_peak_1120 MACS_peak_2822.5 NA 195*196*197*198 5213 CHROMOSOME_X 11924078 11924479 MACS_peak_981 rex-7 33 32 2 32 11924278 0 0 NA MACS_peak_981 MACS_peak_2065.1 NA NA NA CHROMOSOME_X 11898995 11899396 MACS_peak_979 rex-36 27 33 2 33 11899195 2 8 7*8 MACS_peak_979 MACS_peak_2061 NA NA NA CHROMOSOME_X 1858579 1858980 MACS_peak_737.2 NA 39 36 3 34 1858779 0 0 NA MACS_peak_737.2 MACS_peak_335.4 NA 549 NA CHROMOSOME_X 10667802 10668203 MACS_peak_952 rex-28 32 35 3 35 10668002 1 8 8 MACS_peak_952 MACS_peak_1857.2 NA 82*83 4618 CHROMOSOME_X 2364702 2365103 MACS_peak_753 NA 37 37 3 36 2364902 1 7.75 7.75 MACS_peak_753 MACS_peak_427.2 NA NA NA CHROMOSOME_X 1379471 1379872 MACS_peak_725 rex-18 40 39 3 37 1379671 0 0 NA MACS_peak_725 MACS_peak_260 NA 473 NA CHROMOSOME_X 1437084 1437485 MACS_peak_727.1 NA 38 34 3 38 1437284 1 8.75 8.75 MACS_peak_727.1 MACS_peak_272.2 NA NA NA CHROMOSOME_X 11095380 11095781 MACS_peak_959.2 NA 44 47 3 39 11095580 0 0 NA MACS_peak_959.2 MACS_peak_1926.6 NA 237 NA CHROMOSOME_X 4208516 4208917 MACS_peak_792.1 NA 34 38 3 40 4208716 2 8.75 8.75*6.5 MACS_peak_792.1 MACS_peak_776.2 NA NA NA CHROMOSOME_X 13442367 13442768 MACS_peak_1025 NA 41 41 3 41 13442567 0 0 NA MACS_peak_1025 MACS_peak_2309 NA 51 NA CHROMOSOME_X 12730000 12730401 MACS_peak_1006 rex-31 35 40 3 42 12730200 2 9.5 9.5*7.25 MACS_peak_1006 MACS_peak_2195 NA NA NA CHROMOSOME_X 16327193 16327594 MACS_peak_1122 NA 43 44 3 43 16327393 1 8.75 8.75 MACS_peak_1122 MACS_peak_2844 NA NA NA CHROMOSOME_X 16202218 16202619 MACS_peak_1118 NA 42 46 3 44 16202418 1 7.5 7.5 MACS_peak_1118 MACS_peak_2822.1 NA NA NA CHROMOSOME_X 1908787 1909188 MACS_peak_740.1 NA 45 48 3 45 1908987 0 0 NA MACS_peak_740.1 MACS_peak_343.2 NA NA NA CHROMOSOME_X 785349 785750 MACS_peak_688 NA 49 42 3 46 785549 0 0 NA MACS_peak_688 MACS_peak_139.1 NA NA NA CHROMOSOME_X 9467549 9467950 MACS_peak_923.2 NA 54 57 3 47 9467749 0 0 NA MACS_peak_923.2 MACS_peak_1648.4 NA 314 4784 CHROMOSOME_X 13435041 13435442 MACS_peak_1024 NA 50 43 3 48 13435241 0 0 NA MACS_peak_1024 MACS_peak_2308 NA NA NA CHROMOSOME_X 15741776 15742177 MACS_peak_1104 NA 55 54 3 49 15741976 0 0 NA MACS_peak_1104 MACS_peak_2742 NA 309 5111 CHROMOSOME_X 11934867 11935268 MACS_peak_982.1 NA 58 61 3 50 11935067 0 0 NA MACS_peak_982.1 MACS_peak_2066.2 NA 148*149 5602 CHROMOSOME_X 1338889 1339290 MACS_peak_722 NA 46 49 3 51 1339089 1 8 8 MACS_peak_722 MACS_peak_255.1 NA NA NA CHROMOSOME_X 13665321 13665722 MACS_peak_1033 NA 59 59 3 52 13665521 0 0 NA MACS_peak_1033 MACS_peak_2356 NA 1 NA CHROMOSOME_X 13604836 13605237 MACS_peak_1031 NA 53 45 3 53 13605036 0 0 NA MACS_peak_1031 MACS_peak_2341 NA NA NA CHROMOSOME_X 1386915 1387316 MACS_peak_726.3 NA 52 52 3 54 1387115 0 0 NA MACS_peak_726.3 MACS_peak_262.3 NA NA NA CHROMOSOME_X 10756106 10756507 MACS_peak_954 rex-29 47 50 3 55 10756306 1 7.25 7.25 MACS_peak_954 MACS_peak_1873 NA NA NA CHROMOSOME_X 1880680 1881081 MACS_peak_738 NA 64 62 3 56 1880880 0 0 NA MACS_peak_738 MACS_peak_339 NA 578 4939 CHROMOSOME_X 8048595 8048996 MACS_peak_892 rex-17 51 53 3 57 8048795 1 8.25 8.25 MACS_peak_892 MACS_peak_1430.1 NA NA NA CHROMOSOME_X 8404208 8404609 MACS_peak_904 rex-25 60 64 3 58 8404408 0 0 NA MACS_peak_904 MACS_peak_1487 267 450 NA CHROMOSOME_X 1438901 1439302 MACS_peak_727.2 NA 57 51 3 59 1439101 0 0 NA MACS_peak_727.2 MACS_peak_272.3 NA NA NA CHROMOSOME_X 2613032 2613433 MACS_peak_761 NA 62 56 3 60 2613232 0 0 NA MACS_peak_761 MACS_peak_481 NA 526 NA CHROMOSOME_X 1492175 1492576 MACS_peak_730 rex-19 48 58 3 61 1492375 2 8 8*6.75 MACS_peak_730 MACS_peak_279 NA NA NA CHROMOSOME_X 489661 490062 MACS_peak_681 NA 61 63 3 62 489861 1 7.75 7.75 MACS_peak_681 MACS_peak_91 NA 445 5321 CHROMOSOME_X 16695099 16695500 MACS_peak_1133 NA 63 55 3 63 16695299 0 0 NA MACS_peak_1133 MACS_peak_2924 NA NA NA CHROMOSOME_X 10353798 10354199 MACS_peak_942 rex-26 56 60 3 64 10353998 1 8.75 8.75 MACS_peak_942 MACS_peak_1807 NA NA NA ######################################################################################################################################################### ####MACS141_e10_DPY27_N2_MxEmb_splitted_summits_final.bed CHROMOSOME_I 9788714 9788715 MACS_peak_6 CHROMOSOME_II 5543574 5543575 MACS_peak_8 CHROMOSOME_II 6853194 6853195 MACS_peak_11 CHROMOSOME_II 11759373 11759374 MACS_peak_13 CHROMOSOME_II 11967677 11967678 MACS_peak_14 CHROMOSOME_II 14804714 14804715 MACS_peak_17 CHROMOSOME_III 3867570 3867571 MACS_peak_19 CHROMOSOME_III 5031594 5031595 MACS_peak_22 CHROMOSOME_III 5211352 5211353 MACS_peak_23 CHROMOSOME_III 7854644 7854645 MACS_peak_24 CHROMOSOME_III 8043906 8043907 MACS_peak_25 CHROMOSOME_III 10759129 10759130 MACS_peak_26 CHROMOSOME_IV 654907 654908 MACS_peak_28 CHROMOSOME_IV 4390648 4390649 MACS_peak_30 CHROMOSOME_IV 11026168 11026169 MACS_peak_32 CHROMOSOME_V 8235483 8235484 MACS_peak_35 CHROMOSOME_V 15873963 15873964 MACS_peak_39 CHROMOSOME_V 19643160 19643161 MACS_peak_40 CHROMOSOME_X 21167 21168 MACS_peak_43 CHROMOSOME_X 47796 47797 MACS_peak_45 CHROMOSOME_X 104889 104890 MACS_peak_47 CHROMOSOME_X 117289 117290 MACS_peak_48 CHROMOSOME_X 163662 163663 MACS_peak_50 CHROMOSOME_X 191129 191130 MACS_peak_52 CHROMOSOME_X 199204 199205 MACS_peak_54 CHROMOSOME_X 230136 230137 MACS_peak_55 CHROMOSOME_X 239550 239551 MACS_peak_56 CHROMOSOME_X 258689 258690 MACS_peak_57 CHROMOSOME_X 290280 290281 MACS_peak_58 CHROMOSOME_X 295823 295824 MACS_peak_59 CHROMOSOME_X 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CHROMOSOME_X 1654539 1654540 MACS_peak_306 CHROMOSOME_X 1682761 1682762 MACS_peak_310 CHROMOSOME_X 1688804 1688805 MACS_peak_311 CHROMOSOME_X 1698390 1698391 MACS_peak_313 CHROMOSOME_X 1708927 1708928 MACS_peak_315 CHROMOSOME_X 1710519 1710520 MACS_peak_316 CHROMOSOME_X 1720428 1720429 MACS_peak_318 CHROMOSOME_X 1722373 1722374 MACS_peak_319 CHROMOSOME_X 1730351 1730352 MACS_peak_320 CHROMOSOME_X 1746793 1746794 MACS_peak_322 CHROMOSOME_X 1791336 1791337 MACS_peak_323 CHROMOSOME_X 1793889 1793890 MACS_peak_324 CHROMOSOME_X 1796280 1796281 MACS_peak_325 CHROMOSOME_X 1798699 1798700 MACS_peak_326 CHROMOSOME_X 1801170 1801171 MACS_peak_327 CHROMOSOME_X 1824371 1824372 MACS_peak_328 CHROMOSOME_X 1828850 1828851 MACS_peak_329 CHROMOSOME_X 1837902 1837903 MACS_peak_330 CHROMOSOME_X 1845620 1845621 MACS_peak_332 CHROMOSOME_X 1849578 1849579 MACS_peak_334 CHROMOSOME_X 1853788 1853789 MACS_peak_335.1 CHROMOSOME_X 1856375 1856376 MACS_peak_335.2 CHROMOSOME_X 1858072 1858073 MACS_peak_335.3 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2116765 MACS_peak_376 CHROMOSOME_X 2131809 2131810 MACS_peak_379 CHROMOSOME_X 2135004 2135005 MACS_peak_380 CHROMOSOME_X 2136992 2136993 MACS_peak_381 CHROMOSOME_X 2139138 2139139 MACS_peak_382 CHROMOSOME_X 2140871 2140872 MACS_peak_383 CHROMOSOME_X 2145776 2145777 MACS_peak_385 CHROMOSOME_X 2157959 2157960 MACS_peak_388 CHROMOSOME_X 2176645 2176646 MACS_peak_389 CHROMOSOME_X 2178610 2178611 MACS_peak_390 CHROMOSOME_X 2189097 2189098 MACS_peak_392 CHROMOSOME_X 2202768 2202769 MACS_peak_394 CHROMOSOME_X 2206659 2206660 MACS_peak_395 CHROMOSOME_X 2216823 2216824 MACS_peak_396 CHROMOSOME_X 2227263 2227264 MACS_peak_398 CHROMOSOME_X 2234132 2234133 MACS_peak_400 CHROMOSOME_X 2237626 2237627 MACS_peak_401 CHROMOSOME_X 2243013 2243014 MACS_peak_402 CHROMOSOME_X 2247138 2247139 MACS_peak_404 CHROMOSOME_X 2258638 2258639 MACS_peak_407 CHROMOSOME_X 2261132 2261133 MACS_peak_408 CHROMOSOME_X 2268265 2268266 MACS_peak_409 CHROMOSOME_X 2276121 2276122 MACS_peak_410 CHROMOSOME_X 2282936 2282937 MACS_peak_411 CHROMOSOME_X 2288134 2288135 MACS_peak_412 CHROMOSOME_X 2291334 2291335 MACS_peak_413 CHROMOSOME_X 2294376 2294377 MACS_peak_414 CHROMOSOME_X 2299404 2299405 MACS_peak_415 CHROMOSOME_X 2312800 2312801 MACS_peak_419 CHROMOSOME_X 2316353 2316354 MACS_peak_420 CHROMOSOME_X 2321859 2321860 MACS_peak_421 CHROMOSOME_X 2323763 2323764 MACS_peak_422 CHROMOSOME_X 2330703 2330704 MACS_peak_423 CHROMOSOME_X 2334109 2334110 MACS_peak_424 CHROMOSOME_X 2348113 2348114 MACS_peak_425 CHROMOSOME_X 2363055 2363056 MACS_peak_427.1 CHROMOSOME_X 2364904 2364905 MACS_peak_427.2 CHROMOSOME_X 2368679 2368680 MACS_peak_428 CHROMOSOME_X 2372219 2372220 MACS_peak_429 CHROMOSOME_X 2375219 2375220 MACS_peak_430 CHROMOSOME_X 2382441 2382442 MACS_peak_431 CHROMOSOME_X 2393892 2393893 MACS_peak_432 CHROMOSOME_X 2395400 2395401 MACS_peak_433 CHROMOSOME_X 2402130 2402131 MACS_peak_434 CHROMOSOME_X 2407537 2407538 MACS_peak_435 CHROMOSOME_X 2409490 2409491 MACS_peak_436 CHROMOSOME_X 2414439 2414440 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6020323 6020324 MACS_peak_1092 CHROMOSOME_X 6022538 6022539 MACS_peak_1093 CHROMOSOME_X 6028841 6028842 MACS_peak_1094 CHROMOSOME_X 6030624 6030625 MACS_peak_1095 CHROMOSOME_X 6034968 6034969 MACS_peak_1096 CHROMOSOME_X 6057914 6057915 MACS_peak_1097 CHROMOSOME_X 6064549 6064550 MACS_peak_1098 CHROMOSOME_X 6066861 6066862 MACS_peak_1099 CHROMOSOME_X 6084613 6084614 MACS_peak_1104 CHROMOSOME_X 6092099 6092100 MACS_peak_1105 CHROMOSOME_X 6097691 6097692 MACS_peak_1106 CHROMOSOME_X 6100987 6100988 MACS_peak_1107 CHROMOSOME_X 6107271 6107272 MACS_peak_1108 CHROMOSOME_X 6109491 6109492 MACS_peak_1109 CHROMOSOME_X 6113031 6113032 MACS_peak_1110 CHROMOSOME_X 6118295 6118296 MACS_peak_1111 CHROMOSOME_X 6121916 6121917 MACS_peak_1113 CHROMOSOME_X 6128419 6128420 MACS_peak_1114 CHROMOSOME_X 6134703 6134704 MACS_peak_1115 CHROMOSOME_X 6136637 6136638 MACS_peak_1116 CHROMOSOME_X 6146119 6146120 MACS_peak_1118 CHROMOSOME_X 6147695 6147696 MACS_peak_1119 CHROMOSOME_X 6151284 6151285 MACS_peak_1120 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MACS_peak_1183 CHROMOSOME_X 6515512 6515513 MACS_peak_1184 CHROMOSOME_X 6520132 6520133 MACS_peak_1185 CHROMOSOME_X 6551273 6551274 MACS_peak_1188 CHROMOSOME_X 6588108 6588109 MACS_peak_1194 CHROMOSOME_X 6591932 6591933 MACS_peak_1195 CHROMOSOME_X 6593539 6593540 MACS_peak_1196 CHROMOSOME_X 6597247 6597248 MACS_peak_1197 CHROMOSOME_X 6609410 6609411 MACS_peak_1200 CHROMOSOME_X 6611666 6611667 MACS_peak_1201 CHROMOSOME_X 6613728 6613729 MACS_peak_1202 CHROMOSOME_X 6617199 6617200 MACS_peak_1203.1 CHROMOSOME_X 6618218 6618219 MACS_peak_1203.2 CHROMOSOME_X 6619784 6619785 MACS_peak_1203.3 CHROMOSOME_X 6622994 6622995 MACS_peak_1203.4 CHROMOSOME_X 6625410 6625411 MACS_peak_1204 CHROMOSOME_X 6639157 6639158 MACS_peak_1206 CHROMOSOME_X 6647063 6647064 MACS_peak_1207 CHROMOSOME_X 6653280 6653281 MACS_peak_1209 CHROMOSOME_X 6657526 6657527 MACS_peak_1211 CHROMOSOME_X 6664101 6664102 MACS_peak_1212 CHROMOSOME_X 6669218 6669219 MACS_peak_1213 CHROMOSOME_X 6686847 6686848 MACS_peak_1215 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CHROMOSOME_X 7475154 7475155 MACS_peak_1334 CHROMOSOME_X 7478596 7478597 MACS_peak_1335 CHROMOSOME_X 7485888 7485889 MACS_peak_1336 CHROMOSOME_X 7495473 7495474 MACS_peak_1337 CHROMOSOME_X 7501138 7501139 MACS_peak_1338 CHROMOSOME_X 7502564 7502565 MACS_peak_1339 CHROMOSOME_X 7507700 7507701 MACS_peak_1340 CHROMOSOME_X 7518932 7518933 MACS_peak_1342 CHROMOSOME_X 7520891 7520892 MACS_peak_1343 CHROMOSOME_X 7522684 7522685 MACS_peak_1344 CHROMOSOME_X 7524126 7524127 MACS_peak_1345 CHROMOSOME_X 7528118 7528119 MACS_peak_1346 CHROMOSOME_X 7536258 7536259 MACS_peak_1347 CHROMOSOME_X 7545079 7545080 MACS_peak_1348 CHROMOSOME_X 7546514 7546515 MACS_peak_1349 CHROMOSOME_X 7554787 7554788 MACS_peak_1350 CHROMOSOME_X 7560193 7560194 MACS_peak_1351 CHROMOSOME_X 7562743 7562744 MACS_peak_1352.1 CHROMOSOME_X 7565152 7565153 MACS_peak_1352.2 CHROMOSOME_X 7569858 7569859 MACS_peak_1353 CHROMOSOME_X 7573711 7573712 MACS_peak_1354 CHROMOSOME_X 7577645 7577646 MACS_peak_1356 CHROMOSOME_X 7595549 7595550 MACS_peak_1357 CHROMOSOME_X 7598475 7598476 MACS_peak_1358 CHROMOSOME_X 7608752 7608753 MACS_peak_1360 CHROMOSOME_X 7634044 7634045 MACS_peak_1364 CHROMOSOME_X 7636226 7636227 MACS_peak_1365 CHROMOSOME_X 7648428 7648429 MACS_peak_1368 CHROMOSOME_X 7651214 7651215 MACS_peak_1369.1 CHROMOSOME_X 7654179 7654180 MACS_peak_1369.2 CHROMOSOME_X 7655619 7655620 MACS_peak_1369.3 CHROMOSOME_X 7661325 7661326 MACS_peak_1370.1 CHROMOSOME_X 7661778 7661779 MACS_peak_1370.2 CHROMOSOME_X 7662699 7662700 MACS_peak_1370.3 CHROMOSOME_X 7668744 7668745 MACS_peak_1372 CHROMOSOME_X 7678571 7678572 MACS_peak_1373 CHROMOSOME_X 7682119 7682120 MACS_peak_1374.1 CHROMOSOME_X 7682496 7682497 MACS_peak_1374.2 CHROMOSOME_X 7689723 7689724 MACS_peak_1375 CHROMOSOME_X 7710221 7710222 MACS_peak_1377 CHROMOSOME_X 7713017 7713018 MACS_peak_1378 CHROMOSOME_X 7716028 7716029 MACS_peak_1379 CHROMOSOME_X 7720998 7720999 MACS_peak_1380 CHROMOSOME_X 7745482 7745483 MACS_peak_1382 CHROMOSOME_X 7747178 7747179 MACS_peak_1383 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MACS_peak_1507 CHROMOSOME_X 8529007 8529008 MACS_peak_1508 CHROMOSOME_X 8546178 8546179 MACS_peak_1510 CHROMOSOME_X 8550311 8550312 MACS_peak_1511 CHROMOSOME_X 8551857 8551858 MACS_peak_1512 CHROMOSOME_X 8556802 8556803 MACS_peak_1513 CHROMOSOME_X 8564213 8564214 MACS_peak_1514 CHROMOSOME_X 8566210 8566211 MACS_peak_1515 CHROMOSOME_X 8568466 8568467 MACS_peak_1516.1 CHROMOSOME_X 8569308 8569309 MACS_peak_1516.2 CHROMOSOME_X 8599926 8599927 MACS_peak_1520 CHROMOSOME_X 8605199 8605200 MACS_peak_1521 CHROMOSOME_X 8608901 8608902 MACS_peak_1522 CHROMOSOME_X 8624219 8624220 MACS_peak_1523 CHROMOSOME_X 8627809 8627810 MACS_peak_1524 CHROMOSOME_X 8638956 8638957 MACS_peak_1525 CHROMOSOME_X 8646422 8646423 MACS_peak_1528 CHROMOSOME_X 8651249 8651250 MACS_peak_1529 CHROMOSOME_X 8654561 8654562 MACS_peak_1530 CHROMOSOME_X 8662599 8662600 MACS_peak_1531 CHROMOSOME_X 8667517 8667518 MACS_peak_1532 CHROMOSOME_X 8673245 8673246 MACS_peak_1534 CHROMOSOME_X 8674792 8674793 MACS_peak_1535 CHROMOSOME_X 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MACS_peak_1565 CHROMOSOME_X 8889090 8889091 MACS_peak_1566 CHROMOSOME_X 8900280 8900281 MACS_peak_1569 CHROMOSOME_X 8911230 8911231 MACS_peak_1571 CHROMOSOME_X 8925555 8925556 MACS_peak_1572 CHROMOSOME_X 8931156 8931157 MACS_peak_1573 CHROMOSOME_X 8933654 8933655 MACS_peak_1574 CHROMOSOME_X 8940178 8940179 MACS_peak_1575 CHROMOSOME_X 8944254 8944255 MACS_peak_1576 CHROMOSOME_X 8952600 8952601 MACS_peak_1578 CHROMOSOME_X 8964875 8964876 MACS_peak_1579 CHROMOSOME_X 8969512 8969513 MACS_peak_1580 CHROMOSOME_X 8971538 8971539 MACS_peak_1581 CHROMOSOME_X 8986613 8986614 MACS_peak_1582 CHROMOSOME_X 8995760 8995761 MACS_peak_1585 CHROMOSOME_X 9024426 9024427 MACS_peak_1587 CHROMOSOME_X 9028989 9028990 MACS_peak_1588 CHROMOSOME_X 9048815 9048816 MACS_peak_1591 CHROMOSOME_X 9056555 9056556 MACS_peak_1592 CHROMOSOME_X 9067379 9067380 MACS_peak_1594.1 CHROMOSOME_X 9068256 9068257 MACS_peak_1594.2 CHROMOSOME_X 9072811 9072812 MACS_peak_1595 CHROMOSOME_X 9078329 9078330 MACS_peak_1596 CHROMOSOME_X 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MACS_peak_1627 CHROMOSOME_X 9347703 9347704 MACS_peak_1628 CHROMOSOME_X 9369125 9369126 MACS_peak_1630 CHROMOSOME_X 9373160 9373161 MACS_peak_1631 CHROMOSOME_X 9387248 9387249 MACS_peak_1632 CHROMOSOME_X 9396624 9396625 MACS_peak_1635 CHROMOSOME_X 9402090 9402091 MACS_peak_1636 CHROMOSOME_X 9410206 9410207 MACS_peak_1638 CHROMOSOME_X 9416476 9416477 MACS_peak_1639 CHROMOSOME_X 9422885 9422886 MACS_peak_1640 CHROMOSOME_X 9431186 9431187 MACS_peak_1641 CHROMOSOME_X 9437053 9437054 MACS_peak_1642 CHROMOSOME_X 9441634 9441635 MACS_peak_1643 CHROMOSOME_X 9444028 9444029 MACS_peak_1644 CHROMOSOME_X 9446228 9446229 MACS_peak_1645 CHROMOSOME_X 9452691 9452692 MACS_peak_1647 CHROMOSOME_X 9464645 9464646 MACS_peak_1648.1 CHROMOSOME_X 9465798 9465799 MACS_peak_1648.2 CHROMOSOME_X 9467232 9467233 MACS_peak_1648.3 CHROMOSOME_X 9467755 9467756 MACS_peak_1648.4 CHROMOSOME_X 9469150 9469151 MACS_peak_1648.5 CHROMOSOME_X 9477628 9477629 MACS_peak_1650 CHROMOSOME_X 9479099 9479100 MACS_peak_1651 CHROMOSOME_X 9483909 9483910 MACS_peak_1652 CHROMOSOME_X 9487153 9487154 MACS_peak_1653 CHROMOSOME_X 9490530 9490531 MACS_peak_1654 CHROMOSOME_X 9494868 9494869 MACS_peak_1655.1 CHROMOSOME_X 9500045 9500046 MACS_peak_1655.2 CHROMOSOME_X 9522873 9522874 MACS_peak_1656 CHROMOSOME_X 9528160 9528161 MACS_peak_1657 CHROMOSOME_X 9537743 9537744 MACS_peak_1658 CHROMOSOME_X 9539518 9539519 MACS_peak_1659 CHROMOSOME_X 9543408 9543409 MACS_peak_1660 CHROMOSOME_X 9551198 9551199 MACS_peak_1661.1 CHROMOSOME_X 9552570 9552571 MACS_peak_1661.2 CHROMOSOME_X 9561038 9561039 MACS_peak_1663 CHROMOSOME_X 9564074 9564075 MACS_peak_1664 CHROMOSOME_X 9582922 9582923 MACS_peak_1666 CHROMOSOME_X 9584523 9584524 MACS_peak_1667 CHROMOSOME_X 9599238 9599239 MACS_peak_1670 CHROMOSOME_X 9602078 9602079 MACS_peak_1671 CHROMOSOME_X 9610155 9610156 MACS_peak_1672 CHROMOSOME_X 9622844 9622845 MACS_peak_1673 CHROMOSOME_X 9637282 9637283 MACS_peak_1675.1 CHROMOSOME_X 9637873 9637874 MACS_peak_1675.2 CHROMOSOME_X 9648213 9648214 MACS_peak_1678 CHROMOSOME_X 9667682 9667683 MACS_peak_1683.1 CHROMOSOME_X 9668695 9668696 MACS_peak_1683.2 CHROMOSOME_X 9673190 9673191 MACS_peak_1684 CHROMOSOME_X 9680490 9680491 MACS_peak_1686 CHROMOSOME_X 9697770 9697771 MACS_peak_1688 CHROMOSOME_X 9702939 9702940 MACS_peak_1690 CHROMOSOME_X 9704644 9704645 MACS_peak_1691 CHROMOSOME_X 9734501 9734502 MACS_peak_1695 CHROMOSOME_X 9737944 9737945 MACS_peak_1696 CHROMOSOME_X 9743522 9743523 MACS_peak_1697 CHROMOSOME_X 9756975 9756976 MACS_peak_1699 CHROMOSOME_X 9774415 9774416 MACS_peak_1702.1 CHROMOSOME_X 9774897 9774898 MACS_peak_1702.2 CHROMOSOME_X 9785973 9785974 MACS_peak_1703 CHROMOSOME_X 9791273 9791274 MACS_peak_1704 CHROMOSOME_X 9796052 9796053 MACS_peak_1705 CHROMOSOME_X 9801195 9801196 MACS_peak_1706 CHROMOSOME_X 9804783 9804784 MACS_peak_1707 CHROMOSOME_X 9807507 9807508 MACS_peak_1708 CHROMOSOME_X 9822736 9822737 MACS_peak_1709 CHROMOSOME_X 9827544 9827545 MACS_peak_1710 CHROMOSOME_X 9830290 9830291 MACS_peak_1711.1 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MACS_peak_1767.3 CHROMOSOME_X 10154536 10154537 MACS_peak_1768 CHROMOSOME_X 10158864 10158865 MACS_peak_1769 CHROMOSOME_X 10163318 10163319 MACS_peak_1770 CHROMOSOME_X 10168287 10168288 MACS_peak_1771 CHROMOSOME_X 10170826 10170827 MACS_peak_1772 CHROMOSOME_X 10176577 10176578 MACS_peak_1773 CHROMOSOME_X 10192327 10192328 MACS_peak_1774 CHROMOSOME_X 10194555 10194556 MACS_peak_1775 CHROMOSOME_X 10197380 10197381 MACS_peak_1776 CHROMOSOME_X 10203614 10203615 MACS_peak_1777.1 CHROMOSOME_X 10204418 10204419 MACS_peak_1777.2 CHROMOSOME_X 10205686 10205687 MACS_peak_1777.3 CHROMOSOME_X 10215566 10215567 MACS_peak_1779 CHROMOSOME_X 10218730 10218731 MACS_peak_1780 CHROMOSOME_X 10224431 10224432 MACS_peak_1781 CHROMOSOME_X 10226063 10226064 MACS_peak_1782 CHROMOSOME_X 10228441 10228442 MACS_peak_1783 CHROMOSOME_X 10231023 10231024 MACS_peak_1784 CHROMOSOME_X 10232481 10232482 MACS_peak_1785 CHROMOSOME_X 10238318 10238319 MACS_peak_1786 CHROMOSOME_X 10244775 10244776 MACS_peak_1789 CHROMOSOME_X 10249854 10249855 MACS_peak_1790 CHROMOSOME_X 10271548 10271549 MACS_peak_1792 CHROMOSOME_X 10327824 10327825 MACS_peak_1800 CHROMOSOME_X 10332393 10332394 MACS_peak_1801 CHROMOSOME_X 10338552 10338553 MACS_peak_1802 CHROMOSOME_X 10345844 10345845 MACS_peak_1804 CHROMOSOME_X 10348494 10348495 MACS_peak_1805 CHROMOSOME_X 10353987 10353988 MACS_peak_1807 CHROMOSOME_X 10358248 10358249 MACS_peak_1809 CHROMOSOME_X 10372644 10372645 MACS_peak_1810 CHROMOSOME_X 10377832 10377833 MACS_peak_1811 CHROMOSOME_X 10391336 10391337 MACS_peak_1813.1 CHROMOSOME_X 10394398 10394399 MACS_peak_1813.2 CHROMOSOME_X 10395855 10395856 MACS_peak_1813.3 CHROMOSOME_X 10396673 10396674 MACS_peak_1813.4 CHROMOSOME_X 10410875 10410876 MACS_peak_1814 CHROMOSOME_X 10414387 10414388 MACS_peak_1815 CHROMOSOME_X 10419844 10419845 MACS_peak_1816 CHROMOSOME_X 10429300 10429301 MACS_peak_1817 CHROMOSOME_X 10431064 10431065 MACS_peak_1818 CHROMOSOME_X 10432845 10432846 MACS_peak_1819 CHROMOSOME_X 10435666 10435667 MACS_peak_1820 CHROMOSOME_X 10439179 10439180 MACS_peak_1821 CHROMOSOME_X 10482543 10482544 MACS_peak_1827 CHROMOSOME_X 10505281 10505282 MACS_peak_1830 CHROMOSOME_X 10508007 10508008 MACS_peak_1831 CHROMOSOME_X 10512452 10512453 MACS_peak_1832 CHROMOSOME_X 10520331 10520332 MACS_peak_1833 CHROMOSOME_X 10524616 10524617 MACS_peak_1834 CHROMOSOME_X 10529173 10529174 MACS_peak_1835 CHROMOSOME_X 10555626 10555627 MACS_peak_1838 CHROMOSOME_X 10559358 10559359 MACS_peak_1839 CHROMOSOME_X 10566892 10566893 MACS_peak_1840.1 CHROMOSOME_X 10568552 10568553 MACS_peak_1840.2 CHROMOSOME_X 10576566 10576567 MACS_peak_1840.3 CHROMOSOME_X 10576981 10576982 MACS_peak_1840.4 CHROMOSOME_X 10580228 10580229 MACS_peak_1841 CHROMOSOME_X 10590902 10590903 MACS_peak_1843.1 CHROMOSOME_X 10591445 10591446 MACS_peak_1843.2 CHROMOSOME_X 10593940 10593941 MACS_peak_1843.3 CHROMOSOME_X 10596646 10596647 MACS_peak_1843.4 CHROMOSOME_X 10597678 10597679 MACS_peak_1843.5 CHROMOSOME_X 10600031 10600032 MACS_peak_1843.6 CHROMOSOME_X 10618880 10618881 MACS_peak_1844 CHROMOSOME_X 10623255 10623256 MACS_peak_1845.1 CHROMOSOME_X 10624329 10624330 MACS_peak_1845.2 CHROMOSOME_X 10629316 10629317 MACS_peak_1846 CHROMOSOME_X 10637628 10637629 MACS_peak_1848 CHROMOSOME_X 10640760 10640761 MACS_peak_1850 CHROMOSOME_X 10647680 10647681 MACS_peak_1852 CHROMOSOME_X 10656714 10656715 MACS_peak_1855 CHROMOSOME_X 10660345 10660346 MACS_peak_1856 CHROMOSOME_X 10667588 10667589 MACS_peak_1857.1 CHROMOSOME_X 10667999 10668000 MACS_peak_1857.2 CHROMOSOME_X 10670272 10670273 MACS_peak_1858 CHROMOSOME_X 10677681 10677682 MACS_peak_1860.1 CHROMOSOME_X 10678340 10678341 MACS_peak_1860.2 CHROMOSOME_X 10679354 10679355 MACS_peak_1860.3 CHROMOSOME_X 10681766 10681767 MACS_peak_1861 CHROMOSOME_X 10684616 10684617 MACS_peak_1862 CHROMOSOME_X 10686625 10686626 MACS_peak_1863 CHROMOSOME_X 10695113 10695114 MACS_peak_1864 CHROMOSOME_X 10705082 10705083 MACS_peak_1865 CHROMOSOME_X 10709841 10709842 MACS_peak_1866 CHROMOSOME_X 10711191 10711192 MACS_peak_1867 CHROMOSOME_X 10725520 10725521 MACS_peak_1869 CHROMOSOME_X 10731916 10731917 MACS_peak_1870 CHROMOSOME_X 10746468 10746469 MACS_peak_1872 CHROMOSOME_X 10756307 10756308 MACS_peak_1873 CHROMOSOME_X 10805705 10805706 MACS_peak_1878 CHROMOSOME_X 10810416 10810417 MACS_peak_1879 CHROMOSOME_X 10821328 10821329 MACS_peak_1881 CHROMOSOME_X 10826771 10826772 MACS_peak_1882 CHROMOSOME_X 10834610 10834611 MACS_peak_1883 CHROMOSOME_X 10840727 10840728 MACS_peak_1885 CHROMOSOME_X 10843783 10843784 MACS_peak_1886.1 CHROMOSOME_X 10846334 10846335 MACS_peak_1886.2 CHROMOSOME_X 10852075 10852076 MACS_peak_1887 CHROMOSOME_X 10870475 10870476 MACS_peak_1890 CHROMOSOME_X 10877337 10877338 MACS_peak_1892 CHROMOSOME_X 10900230 10900231 MACS_peak_1896.1 CHROMOSOME_X 10901115 10901116 MACS_peak_1896.2 CHROMOSOME_X 10910795 10910796 MACS_peak_1899 CHROMOSOME_X 10923648 10923649 MACS_peak_1901 CHROMOSOME_X 10932261 10932262 MACS_peak_1902 CHROMOSOME_X 10937580 10937581 MACS_peak_1903 CHROMOSOME_X 10949890 10949891 MACS_peak_1904 CHROMOSOME_X 10963747 10963748 MACS_peak_1905 CHROMOSOME_X 10965693 10965694 MACS_peak_1906 CHROMOSOME_X 10979550 10979551 MACS_peak_1908 CHROMOSOME_X 10990751 10990752 MACS_peak_1909.1 CHROMOSOME_X 10992288 10992289 MACS_peak_1909.2 CHROMOSOME_X 10994475 10994476 MACS_peak_1910 CHROMOSOME_X 10998265 10998266 MACS_peak_1911 CHROMOSOME_X 11002999 11003000 MACS_peak_1912 CHROMOSOME_X 11011198 11011199 MACS_peak_1913 CHROMOSOME_X 11014952 11014953 MACS_peak_1914 CHROMOSOME_X 11024160 11024161 MACS_peak_1915.1 CHROMOSOME_X 11024954 11024955 MACS_peak_1915.2 CHROMOSOME_X 11057262 11057263 MACS_peak_1919.1 CHROMOSOME_X 11059846 11059847 MACS_peak_1919.2 CHROMOSOME_X 11062072 11062073 MACS_peak_1919.3 CHROMOSOME_X 11064955 11064956 MACS_peak_1920 CHROMOSOME_X 11068684 11068685 MACS_peak_1921 CHROMOSOME_X 11072831 11072832 MACS_peak_1922 CHROMOSOME_X 11080491 11080492 MACS_peak_1923 CHROMOSOME_X 11089177 11089178 MACS_peak_1926.1 CHROMOSOME_X 11091453 11091454 MACS_peak_1926.2 CHROMOSOME_X 11091776 11091777 MACS_peak_1926.3 CHROMOSOME_X 11093034 11093035 MACS_peak_1926.4 CHROMOSOME_X 11094125 11094126 MACS_peak_1926.5 CHROMOSOME_X 11095570 11095571 MACS_peak_1926.6 CHROMOSOME_X 11103245 11103246 MACS_peak_1927 CHROMOSOME_X 11120950 11120951 MACS_peak_1932 CHROMOSOME_X 11124040 11124041 MACS_peak_1933 CHROMOSOME_X 11127808 11127809 MACS_peak_1934 CHROMOSOME_X 11140800 11140801 MACS_peak_1938 CHROMOSOME_X 11155449 11155450 MACS_peak_1939 CHROMOSOME_X 11159778 11159779 MACS_peak_1940 CHROMOSOME_X 11163377 11163378 MACS_peak_1941 CHROMOSOME_X 11174470 11174471 MACS_peak_1942 CHROMOSOME_X 11184214 11184215 MACS_peak_1943 CHROMOSOME_X 11190581 11190582 MACS_peak_1945 CHROMOSOME_X 11207101 11207102 MACS_peak_1947.1 CHROMOSOME_X 11210544 11210545 MACS_peak_1947.2 CHROMOSOME_X 11215622 11215623 MACS_peak_1948 CHROMOSOME_X 11223020 11223021 MACS_peak_1950.1 CHROMOSOME_X 11223477 11223478 MACS_peak_1950.2 CHROMOSOME_X 11235253 11235254 MACS_peak_1952 CHROMOSOME_X 11238365 11238366 MACS_peak_1953 CHROMOSOME_X 11245209 11245210 MACS_peak_1954 CHROMOSOME_X 11248217 11248218 MACS_peak_1956 CHROMOSOME_X 11252533 11252534 MACS_peak_1957 CHROMOSOME_X 11257234 11257235 MACS_peak_1958 CHROMOSOME_X 11262427 11262428 MACS_peak_1959 CHROMOSOME_X 11266985 11266986 MACS_peak_1960 CHROMOSOME_X 11289008 11289009 MACS_peak_1964.1 CHROMOSOME_X 11290651 11290652 MACS_peak_1964.2 CHROMOSOME_X 11297093 11297094 MACS_peak_1965.1 CHROMOSOME_X 11299711 11299712 MACS_peak_1965.2 CHROMOSOME_X 11306717 11306718 MACS_peak_1966 CHROMOSOME_X 11310390 11310391 MACS_peak_1967 CHROMOSOME_X 11328927 11328928 MACS_peak_1971 CHROMOSOME_X 11339385 11339386 MACS_peak_1973.1 CHROMOSOME_X 11345096 11345097 MACS_peak_1973.2 CHROMOSOME_X 11361287 11361288 MACS_peak_1976 CHROMOSOME_X 11366043 11366044 MACS_peak_1977 CHROMOSOME_X 11384765 11384766 MACS_peak_1978 CHROMOSOME_X 11393323 11393324 MACS_peak_1979 CHROMOSOME_X 11400928 11400929 MACS_peak_1980 CHROMOSOME_X 11407992 11407993 MACS_peak_1982 CHROMOSOME_X 11414990 11414991 MACS_peak_1983.1 CHROMOSOME_X 11415273 11415274 MACS_peak_1983.2 CHROMOSOME_X 11420067 11420068 MACS_peak_1984 CHROMOSOME_X 11434097 11434098 MACS_peak_1986 CHROMOSOME_X 11437770 11437771 MACS_peak_1987 CHROMOSOME_X 11439066 11439067 MACS_peak_1988 CHROMOSOME_X 11448371 11448372 MACS_peak_1990 CHROMOSOME_X 11452469 11452470 MACS_peak_1992 CHROMOSOME_X 11454316 11454317 MACS_peak_1993 CHROMOSOME_X 11461748 11461749 MACS_peak_1994.1 CHROMOSOME_X 11465407 11465408 MACS_peak_1994.2 CHROMOSOME_X 11467395 11467396 MACS_peak_1994.3 CHROMOSOME_X 11511345 11511346 MACS_peak_1996 CHROMOSOME_X 11521589 11521590 MACS_peak_1997.1 CHROMOSOME_X 11522230 11522231 MACS_peak_1997.2 CHROMOSOME_X 11546728 11546729 MACS_peak_1998.1 CHROMOSOME_X 11548846 11548847 MACS_peak_1998.2 CHROMOSOME_X 11552986 11552987 MACS_peak_1999 CHROMOSOME_X 11554637 11554638 MACS_peak_2000 CHROMOSOME_X 11558362 11558363 MACS_peak_2001 CHROMOSOME_X 11560137 11560138 MACS_peak_2002 CHROMOSOME_X 11563793 11563794 MACS_peak_2003 CHROMOSOME_X 11571869 11571870 MACS_peak_2004 CHROMOSOME_X 11574366 11574367 MACS_peak_2005 CHROMOSOME_X 11589314 11589315 MACS_peak_2007 CHROMOSOME_X 11592900 11592901 MACS_peak_2008 CHROMOSOME_X 11595115 11595116 MACS_peak_2009 CHROMOSOME_X 11600606 11600607 MACS_peak_2010 CHROMOSOME_X 11603484 11603485 MACS_peak_2011.1 CHROMOSOME_X 11605815 11605816 MACS_peak_2011.2 CHROMOSOME_X 11608217 11608218 MACS_peak_2011.3 CHROMOSOME_X 11618059 11618060 MACS_peak_2013 CHROMOSOME_X 11639183 11639184 MACS_peak_2016 CHROMOSOME_X 11640712 11640713 MACS_peak_2017 CHROMOSOME_X 11643755 11643756 MACS_peak_2018.1 CHROMOSOME_X 11644062 11644063 MACS_peak_2018.2 CHROMOSOME_X 11652863 11652864 MACS_peak_2021 CHROMOSOME_X 11656805 11656806 MACS_peak_2022 CHROMOSOME_X 11662695 11662696 MACS_peak_2023.1 CHROMOSOME_X 11666725 11666726 MACS_peak_2023.2 CHROMOSOME_X 11673486 11673487 MACS_peak_2024 CHROMOSOME_X 11680969 11680970 MACS_peak_2026 CHROMOSOME_X 11693178 11693179 MACS_peak_2028 CHROMOSOME_X 11710389 11710390 MACS_peak_2029 CHROMOSOME_X 11718951 11718952 MACS_peak_2032 CHROMOSOME_X 11724818 11724819 MACS_peak_2033 CHROMOSOME_X 11740276 11740277 MACS_peak_2035 CHROMOSOME_X 11744574 11744575 MACS_peak_2036 CHROMOSOME_X 11757851 11757852 MACS_peak_2038 CHROMOSOME_X 11766548 11766549 MACS_peak_2039 CHROMOSOME_X 11782611 11782612 MACS_peak_2042 CHROMOSOME_X 11798148 11798149 MACS_peak_2043 CHROMOSOME_X 11801371 11801372 MACS_peak_2044 CHROMOSOME_X 11806308 11806309 MACS_peak_2045 CHROMOSOME_X 11813742 11813743 MACS_peak_2046 CHROMOSOME_X 11816243 11816244 MACS_peak_2047 CHROMOSOME_X 11819902 11819903 MACS_peak_2048 CHROMOSOME_X 11821357 11821358 MACS_peak_2049 CHROMOSOME_X 11822756 11822757 MACS_peak_2050 CHROMOSOME_X 11838211 11838212 MACS_peak_2053 CHROMOSOME_X 11857158 11857159 MACS_peak_2055 CHROMOSOME_X 11870475 11870476 MACS_peak_2057 CHROMOSOME_X 11872399 11872400 MACS_peak_2058 CHROMOSOME_X 11899185 11899186 MACS_peak_2061 CHROMOSOME_X 11915824 11915825 MACS_peak_2063 CHROMOSOME_X 11920761 11920762 MACS_peak_2064 CHROMOSOME_X 11924292 11924293 MACS_peak_2065.1 CHROMOSOME_X 11926412 11926413 MACS_peak_2065.2 CHROMOSOME_X 11932632 11932633 MACS_peak_2066.1 CHROMOSOME_X 11935088 11935089 MACS_peak_2066.2 CHROMOSOME_X 11936078 11936079 MACS_peak_2066.3 CHROMOSOME_X 11936583 11936584 MACS_peak_2066.4 CHROMOSOME_X 11937462 11937463 MACS_peak_2066.5 CHROMOSOME_X 11938768 11938769 MACS_peak_2066.6 CHROMOSOME_X 11947235 11947236 MACS_peak_2067.1 CHROMOSOME_X 11948324 11948325 MACS_peak_2067.2 CHROMOSOME_X 11957100 11957101 MACS_peak_2069 CHROMOSOME_X 11963778 11963779 MACS_peak_2070 CHROMOSOME_X 11965791 11965792 MACS_peak_2071 CHROMOSOME_X 11974858 11974859 MACS_peak_2072 CHROMOSOME_X 11983764 11983765 MACS_peak_2073 CHROMOSOME_X 11988027 11988028 MACS_peak_2074 CHROMOSOME_X 11992344 11992345 MACS_peak_2075 CHROMOSOME_X 11995964 11995965 MACS_peak_2076 CHROMOSOME_X 12000257 12000258 MACS_peak_2077 CHROMOSOME_X 12005948 12005949 MACS_peak_2078 CHROMOSOME_X 12009898 12009899 MACS_peak_2079 CHROMOSOME_X 12012983 12012984 MACS_peak_2080 CHROMOSOME_X 12018091 12018092 MACS_peak_2081 CHROMOSOME_X 12021551 12021552 MACS_peak_2082 CHROMOSOME_X 12027428 12027429 MACS_peak_2083 CHROMOSOME_X 12034301 12034302 MACS_peak_2085 CHROMOSOME_X 12036065 12036066 MACS_peak_2086 CHROMOSOME_X 12044278 12044279 MACS_peak_2087 CHROMOSOME_X 12050279 12050280 MACS_peak_2088 CHROMOSOME_X 12051490 12051491 MACS_peak_2089 CHROMOSOME_X 12061432 12061433 MACS_peak_2090 CHROMOSOME_X 12072022 12072023 MACS_peak_2092 CHROMOSOME_X 12080864 12080865 MACS_peak_2093 CHROMOSOME_X 12095870 12095871 MACS_peak_2094 CHROMOSOME_X 12098156 12098157 MACS_peak_2095 CHROMOSOME_X 12099861 12099862 MACS_peak_2096 CHROMOSOME_X 12106160 12106161 MACS_peak_2097 CHROMOSOME_X 12142467 12142468 MACS_peak_2101 CHROMOSOME_X 12146077 12146078 MACS_peak_2103 CHROMOSOME_X 12149790 12149791 MACS_peak_2104 CHROMOSOME_X 12154236 12154237 MACS_peak_2105 CHROMOSOME_X 12158699 12158700 MACS_peak_2106.1 CHROMOSOME_X 12160410 12160411 MACS_peak_2106.2 CHROMOSOME_X 12162746 12162747 MACS_peak_2107 CHROMOSOME_X 12168145 12168146 MACS_peak_2109.1 CHROMOSOME_X 12168853 12168854 MACS_peak_2109.2 CHROMOSOME_X 12169441 12169442 MACS_peak_2109.3 CHROMOSOME_X 12172055 12172056 MACS_peak_2109.4 CHROMOSOME_X 12175814 12175815 MACS_peak_2109.5 CHROMOSOME_X 12183119 12183120 MACS_peak_2109.6 CHROMOSOME_X 12183337 12183338 MACS_peak_2109.7 CHROMOSOME_X 12184641 12184642 MACS_peak_2109.8 CHROMOSOME_X 12189507 12189508 MACS_peak_2110 CHROMOSOME_X 12191772 12191773 MACS_peak_2111.1 CHROMOSOME_X 12195286 12195287 MACS_peak_2111.2 CHROMOSOME_X 12205465 12205466 MACS_peak_2112.1 CHROMOSOME_X 12205852 12205853 MACS_peak_2112.2 CHROMOSOME_X 12208370 12208371 MACS_peak_2113 CHROMOSOME_X 12230782 12230783 MACS_peak_2116 CHROMOSOME_X 12235484 12235485 MACS_peak_2117 CHROMOSOME_X 12248294 12248295 MACS_peak_2118 CHROMOSOME_X 12249482 12249483 MACS_peak_2119 CHROMOSOME_X 12251139 12251140 MACS_peak_2120 CHROMOSOME_X 12256646 12256647 MACS_peak_2121 CHROMOSOME_X 12260485 12260486 MACS_peak_2122 CHROMOSOME_X 12282992 12282993 MACS_peak_2124 CHROMOSOME_X 12293771 12293772 MACS_peak_2125.1 CHROMOSOME_X 12294367 12294368 MACS_peak_2125.2 CHROMOSOME_X 12304409 12304410 MACS_peak_2126 CHROMOSOME_X 12320808 12320809 MACS_peak_2127 CHROMOSOME_X 12337948 12337949 MACS_peak_2128 CHROMOSOME_X 12348031 12348032 MACS_peak_2131 CHROMOSOME_X 12350296 12350297 MACS_peak_2132 CHROMOSOME_X 12355524 12355525 MACS_peak_2134 CHROMOSOME_X 12363158 12363159 MACS_peak_2135.1 CHROMOSOME_X 12365460 12365461 MACS_peak_2135.2 CHROMOSOME_X 12367650 12367651 MACS_peak_2135.3 CHROMOSOME_X 12378471 12378472 MACS_peak_2138 CHROMOSOME_X 12392404 12392405 MACS_peak_2141 CHROMOSOME_X 12393909 12393910 MACS_peak_2142 CHROMOSOME_X 12398608 12398609 MACS_peak_2143 CHROMOSOME_X 12407392 12407393 MACS_peak_2144 CHROMOSOME_X 12410264 12410265 MACS_peak_2145 CHROMOSOME_X 12414619 12414620 MACS_peak_2146 CHROMOSOME_X 12415995 12415996 MACS_peak_2147 CHROMOSOME_X 12420404 12420405 MACS_peak_2149 CHROMOSOME_X 12425607 12425608 MACS_peak_2150 CHROMOSOME_X 12440488 12440489 MACS_peak_2152 CHROMOSOME_X 12445514 12445515 MACS_peak_2153 CHROMOSOME_X 12449319 12449320 MACS_peak_2154 CHROMOSOME_X 12452256 12452257 MACS_peak_2155.1 CHROMOSOME_X 12454429 12454430 MACS_peak_2155.2 CHROMOSOME_X 12458178 12458179 MACS_peak_2156 CHROMOSOME_X 12480548 12480549 MACS_peak_2157 CHROMOSOME_X 12491095 12491096 MACS_peak_2158 CHROMOSOME_X 12500868 12500869 MACS_peak_2159 CHROMOSOME_X 12503632 12503633 MACS_peak_2160.1 CHROMOSOME_X 12509392 12509393 MACS_peak_2160.2 CHROMOSOME_X 12510148 12510149 MACS_peak_2160.3 CHROMOSOME_X 12515779 12515780 MACS_peak_2160.4 CHROMOSOME_X 12519046 12519047 MACS_peak_2160.5 CHROMOSOME_X 12522950 12522951 MACS_peak_2161 CHROMOSOME_X 12526555 12526556 MACS_peak_2162 CHROMOSOME_X 12533803 12533804 MACS_peak_2163 CHROMOSOME_X 12552579 12552580 MACS_peak_2166 CHROMOSOME_X 12556748 12556749 MACS_peak_2167 CHROMOSOME_X 12563717 12563718 MACS_peak_2168 CHROMOSOME_X 12579620 12579621 MACS_peak_2169 CHROMOSOME_X 12589442 12589443 MACS_peak_2172 CHROMOSOME_X 12618572 12618573 MACS_peak_2177 CHROMOSOME_X 12637662 12637663 MACS_peak_2178 CHROMOSOME_X 12645922 12645923 MACS_peak_2179 CHROMOSOME_X 12656670 12656671 MACS_peak_2180 CHROMOSOME_X 12661113 12661114 MACS_peak_2181.1 CHROMOSOME_X 12665150 12665151 MACS_peak_2181.2 CHROMOSOME_X 12670475 12670476 MACS_peak_2183 CHROMOSOME_X 12675818 12675819 MACS_peak_2185 CHROMOSOME_X 12682689 12682690 MACS_peak_2187 CHROMOSOME_X 12686550 12686551 MACS_peak_2188.1 CHROMOSOME_X 12688446 12688447 MACS_peak_2188.2 CHROMOSOME_X 12689640 12689641 MACS_peak_2188.3 CHROMOSOME_X 12690522 12690523 MACS_peak_2188.4 CHROMOSOME_X 12692027 12692028 MACS_peak_2188.5 CHROMOSOME_X 12693260 12693261 MACS_peak_2188.6 CHROMOSOME_X 12713104 12713105 MACS_peak_2190 CHROMOSOME_X 12717935 12717936 MACS_peak_2191 CHROMOSOME_X 12721734 12721735 MACS_peak_2192 CHROMOSOME_X 12725040 12725041 MACS_peak_2193 CHROMOSOME_X 12730202 12730203 MACS_peak_2195 CHROMOSOME_X 12749972 12749973 MACS_peak_2196 CHROMOSOME_X 12759782 12759783 MACS_peak_2197 CHROMOSOME_X 12781960 12781961 MACS_peak_2199 CHROMOSOME_X 12787277 12787278 MACS_peak_2200 CHROMOSOME_X 12798168 12798169 MACS_peak_2202 CHROMOSOME_X 12823149 12823150 MACS_peak_2206 CHROMOSOME_X 12843191 12843192 MACS_peak_2210.1 CHROMOSOME_X 12843617 12843618 MACS_peak_2210.2 CHROMOSOME_X 12853018 12853019 MACS_peak_2211.1 CHROMOSOME_X 12853610 12853611 MACS_peak_2211.2 CHROMOSOME_X 12854251 12854252 MACS_peak_2211.3 CHROMOSOME_X 12857618 12857619 MACS_peak_2212 CHROMOSOME_X 12860809 12860810 MACS_peak_2213 CHROMOSOME_X 12870074 12870075 MACS_peak_2214 CHROMOSOME_X 12872474 12872475 MACS_peak_2215 CHROMOSOME_X 12877044 12877045 MACS_peak_2216 CHROMOSOME_X 12889869 12889870 MACS_peak_2217.1 CHROMOSOME_X 12890249 12890250 MACS_peak_2217.2 CHROMOSOME_X 12892892 12892893 MACS_peak_2218 CHROMOSOME_X 12899251 12899252 MACS_peak_2219 CHROMOSOME_X 12902425 12902426 MACS_peak_2220 CHROMOSOME_X 12903899 12903900 MACS_peak_2221 CHROMOSOME_X 12928455 12928456 MACS_peak_2222 CHROMOSOME_X 12934841 12934842 MACS_peak_2223 CHROMOSOME_X 12942481 12942482 MACS_peak_2224 CHROMOSOME_X 12944580 12944581 MACS_peak_2225 CHROMOSOME_X 12958205 12958206 MACS_peak_2226 CHROMOSOME_X 12972710 12972711 MACS_peak_2228 CHROMOSOME_X 12975754 12975755 MACS_peak_2229 CHROMOSOME_X 12989340 12989341 MACS_peak_2231.1 CHROMOSOME_X 12990912 12990913 MACS_peak_2231.2 CHROMOSOME_X 12994457 12994458 MACS_peak_2232 CHROMOSOME_X 13003244 13003245 MACS_peak_2233 CHROMOSOME_X 13004541 13004542 MACS_peak_2234 CHROMOSOME_X 13008523 13008524 MACS_peak_2235.1 CHROMOSOME_X 13009811 13009812 MACS_peak_2235.2 CHROMOSOME_X 13012677 13012678 MACS_peak_2235.3 CHROMOSOME_X 13017721 13017722 MACS_peak_2237 CHROMOSOME_X 13021556 13021557 MACS_peak_2238 CHROMOSOME_X 13036864 13036865 MACS_peak_2240 CHROMOSOME_X 13038869 13038870 MACS_peak_2241 CHROMOSOME_X 13041991 13041992 MACS_peak_2242 CHROMOSOME_X 13046421 13046422 MACS_peak_2243 CHROMOSOME_X 13058901 13058902 MACS_peak_2244 CHROMOSOME_X 13068501 13068502 MACS_peak_2247 CHROMOSOME_X 13071100 13071101 MACS_peak_2248 CHROMOSOME_X 13105067 13105068 MACS_peak_2250 CHROMOSOME_X 13108917 13108918 MACS_peak_2251 CHROMOSOME_X 13117196 13117197 MACS_peak_2252 CHROMOSOME_X 13119924 13119925 MACS_peak_2253.1 CHROMOSOME_X 13121177 13121178 MACS_peak_2253.2 CHROMOSOME_X 13122372 13122373 MACS_peak_2253.3 CHROMOSOME_X 13125106 13125107 MACS_peak_2254 CHROMOSOME_X 13141544 13141545 MACS_peak_2255.1 CHROMOSOME_X 13143385 13143386 MACS_peak_2255.2 CHROMOSOME_X 13144239 13144240 MACS_peak_2255.3 CHROMOSOME_X 13145327 13145328 MACS_peak_2255.4 CHROMOSOME_X 13146128 13146129 MACS_peak_2255.5 CHROMOSOME_X 13197139 13197140 MACS_peak_2261 CHROMOSOME_X 13206227 13206228 MACS_peak_2262 CHROMOSOME_X 13209064 13209065 MACS_peak_2263 CHROMOSOME_X 13213598 13213599 MACS_peak_2264 CHROMOSOME_X 13221460 13221461 MACS_peak_2265.1 CHROMOSOME_X 13225188 13225189 MACS_peak_2265.2 CHROMOSOME_X 13225607 13225608 MACS_peak_2265.3 CHROMOSOME_X 13226912 13226913 MACS_peak_2265.4 CHROMOSOME_X 13238502 13238503 MACS_peak_2266 CHROMOSOME_X 13245283 13245284 MACS_peak_2267 CHROMOSOME_X 13251561 13251562 MACS_peak_2269 CHROMOSOME_X 13262031 13262032 MACS_peak_2271 CHROMOSOME_X 13264705 13264706 MACS_peak_2272 CHROMOSOME_X 13279498 13279499 MACS_peak_2274 CHROMOSOME_X 13283559 13283560 MACS_peak_2275 CHROMOSOME_X 13306949 13306950 MACS_peak_2278 CHROMOSOME_X 13314221 13314222 MACS_peak_2280 CHROMOSOME_X 13318586 13318587 MACS_peak_2281 CHROMOSOME_X 13320687 13320688 MACS_peak_2282 CHROMOSOME_X 13328995 13328996 MACS_peak_2284 CHROMOSOME_X 13331673 13331674 MACS_peak_2285 CHROMOSOME_X 13337172 13337173 MACS_peak_2286 CHROMOSOME_X 13341157 13341158 MACS_peak_2287 CHROMOSOME_X 13351043 13351044 MACS_peak_2288 CHROMOSOME_X 13353715 13353716 MACS_peak_2289 CHROMOSOME_X 13358715 13358716 MACS_peak_2291 CHROMOSOME_X 13364888 13364889 MACS_peak_2292 CHROMOSOME_X 13369270 13369271 MACS_peak_2293 CHROMOSOME_X 13377578 13377579 MACS_peak_2294 CHROMOSOME_X 13381032 13381033 MACS_peak_2295.1 CHROMOSOME_X 13381787 13381788 MACS_peak_2295.2 CHROMOSOME_X 13383482 13383483 MACS_peak_2296 CHROMOSOME_X 13386284 13386285 MACS_peak_2297 CHROMOSOME_X 13389018 13389019 MACS_peak_2298 CHROMOSOME_X 13396685 13396686 MACS_peak_2299 CHROMOSOME_X 13401488 13401489 MACS_peak_2300 CHROMOSOME_X 13409627 13409628 MACS_peak_2301 CHROMOSOME_X 13415128 13415129 MACS_peak_2303 CHROMOSOME_X 13419964 13419965 MACS_peak_2304 CHROMOSOME_X 13435217 13435218 MACS_peak_2308 CHROMOSOME_X 13442568 13442569 MACS_peak_2309 CHROMOSOME_X 13448894 13448895 MACS_peak_2310 CHROMOSOME_X 13451850 13451851 MACS_peak_2311 CHROMOSOME_X 13472003 13472004 MACS_peak_2313 CHROMOSOME_X 13477594 13477595 MACS_peak_2314 CHROMOSOME_X 13479623 13479624 MACS_peak_2315 CHROMOSOME_X 13489857 13489858 MACS_peak_2318 CHROMOSOME_X 13493776 13493777 MACS_peak_2319 CHROMOSOME_X 13500793 13500794 MACS_peak_2320 CHROMOSOME_X 13502473 13502474 MACS_peak_2321 CHROMOSOME_X 13503991 13503992 MACS_peak_2322 CHROMOSOME_X 13506111 13506112 MACS_peak_2323 CHROMOSOME_X 13511978 13511979 MACS_peak_2324.1 CHROMOSOME_X 13514506 13514507 MACS_peak_2324.2 CHROMOSOME_X 13519189 13519190 MACS_peak_2324.3 CHROMOSOME_X 13525221 13525222 MACS_peak_2324.4 CHROMOSOME_X 13526592 13526593 MACS_peak_2324.5 CHROMOSOME_X 13527434 13527435 MACS_peak_2324.6 CHROMOSOME_X 13532765 13532766 MACS_peak_2325 CHROMOSOME_X 13541374 13541375 MACS_peak_2328 CHROMOSOME_X 13542914 13542915 MACS_peak_2329 CHROMOSOME_X 13555225 13555226 MACS_peak_2332 CHROMOSOME_X 13558752 13558753 MACS_peak_2333 CHROMOSOME_X 13562095 13562096 MACS_peak_2334 CHROMOSOME_X 13569289 13569290 MACS_peak_2337 CHROMOSOME_X 13574188 13574189 MACS_peak_2338 CHROMOSOME_X 13596114 13596115 MACS_peak_2339 CHROMOSOME_X 13598181 13598182 MACS_peak_2340 CHROMOSOME_X 13605071 13605072 MACS_peak_2341 CHROMOSOME_X 13611658 13611659 MACS_peak_2343 CHROMOSOME_X 13615235 13615236 MACS_peak_2345 CHROMOSOME_X 13619694 13619695 MACS_peak_2347 CHROMOSOME_X 13624564 13624565 MACS_peak_2348 CHROMOSOME_X 13632461 13632462 MACS_peak_2349 CHROMOSOME_X 13638481 13638482 MACS_peak_2350 CHROMOSOME_X 13646798 13646799 MACS_peak_2352 CHROMOSOME_X 13653135 13653136 MACS_peak_2353.1 CHROMOSOME_X 13653654 13653655 MACS_peak_2353.2 CHROMOSOME_X 13657482 13657483 MACS_peak_2354 CHROMOSOME_X 13665522 13665523 MACS_peak_2356 CHROMOSOME_X 13668790 13668791 MACS_peak_2357.1 CHROMOSOME_X 13669766 13669767 MACS_peak_2357.2 CHROMOSOME_X 13675501 13675502 MACS_peak_2358 CHROMOSOME_X 13687177 13687178 MACS_peak_2359 CHROMOSOME_X 13696362 13696363 MACS_peak_2360.1 CHROMOSOME_X 13697665 13697666 MACS_peak_2360.2 CHROMOSOME_X 13700806 13700807 MACS_peak_2360.3 CHROMOSOME_X 13701762 13701763 MACS_peak_2360.4 CHROMOSOME_X 13702929 13702930 MACS_peak_2360.5 CHROMOSOME_X 13716403 13716404 MACS_peak_2361 CHROMOSOME_X 13727017 13727018 MACS_peak_2362 CHROMOSOME_X 13730692 13730693 MACS_peak_2363 CHROMOSOME_X 13739495 13739496 MACS_peak_2365 CHROMOSOME_X 13743503 13743504 MACS_peak_2366.1 CHROMOSOME_X 13746800 13746801 MACS_peak_2366.2 CHROMOSOME_X 13752927 13752928 MACS_peak_2367.1 CHROMOSOME_X 13754685 13754686 MACS_peak_2367.2 CHROMOSOME_X 13760270 13760271 MACS_peak_2368 CHROMOSOME_X 13765599 13765600 MACS_peak_2369 CHROMOSOME_X 13767581 13767582 MACS_peak_2370 CHROMOSOME_X 13768776 13768777 MACS_peak_2371 CHROMOSOME_X 13770856 13770857 MACS_peak_2372 CHROMOSOME_X 13774114 13774115 MACS_peak_2373 CHROMOSOME_X 13778665 13778666 MACS_peak_2374 CHROMOSOME_X 13781544 13781545 MACS_peak_2375 CHROMOSOME_X 13803396 13803397 MACS_peak_2378 CHROMOSOME_X 13812003 13812004 MACS_peak_2380.1 CHROMOSOME_X 13813018 13813019 MACS_peak_2380.2 CHROMOSOME_X 13831064 13831065 MACS_peak_2383 CHROMOSOME_X 13833590 13833591 MACS_peak_2384 CHROMOSOME_X 13841429 13841430 MACS_peak_2385 CHROMOSOME_X 13846142 13846143 MACS_peak_2387 CHROMOSOME_X 13849332 13849333 MACS_peak_2388 CHROMOSOME_X 13889228 13889229 MACS_peak_2390 CHROMOSOME_X 13909860 13909861 MACS_peak_2392 CHROMOSOME_X 13914484 13914485 MACS_peak_2393 CHROMOSOME_X 13917488 13917489 MACS_peak_2394.1 CHROMOSOME_X 13918252 13918253 MACS_peak_2394.2 CHROMOSOME_X 13920530 13920531 MACS_peak_2395 CHROMOSOME_X 13926200 13926201 MACS_peak_2396 CHROMOSOME_X 13937041 13937042 MACS_peak_2398 CHROMOSOME_X 13941404 13941405 MACS_peak_2399 CHROMOSOME_X 13944321 13944322 MACS_peak_2400.1 CHROMOSOME_X 13945640 13945641 MACS_peak_2400.2 CHROMOSOME_X 13948288 13948289 MACS_peak_2401 CHROMOSOME_X 13950716 13950717 MACS_peak_2402 CHROMOSOME_X 13959879 13959880 MACS_peak_2403 CHROMOSOME_X 13969679 13969680 MACS_peak_2405 CHROMOSOME_X 13973959 13973960 MACS_peak_2406 CHROMOSOME_X 13977576 13977577 MACS_peak_2407 CHROMOSOME_X 13995903 13995904 MACS_peak_2409 CHROMOSOME_X 14000535 14000536 MACS_peak_2410.1 CHROMOSOME_X 14001593 14001594 MACS_peak_2410.2 CHROMOSOME_X 14009611 14009612 MACS_peak_2412 CHROMOSOME_X 14015869 14015870 MACS_peak_2413.1 CHROMOSOME_X 14016855 14016856 MACS_peak_2413.2 CHROMOSOME_X 14024598 14024599 MACS_peak_2416 CHROMOSOME_X 14027058 14027059 MACS_peak_2417 CHROMOSOME_X 14028829 14028830 MACS_peak_2418 CHROMOSOME_X 14034572 14034573 MACS_peak_2419 CHROMOSOME_X 14039723 14039724 MACS_peak_2420 CHROMOSOME_X 14044444 14044445 MACS_peak_2421 CHROMOSOME_X 14047744 14047745 MACS_peak_2422 CHROMOSOME_X 14054898 14054899 MACS_peak_2423 CHROMOSOME_X 14063953 14063954 MACS_peak_2425 CHROMOSOME_X 14068746 14068747 MACS_peak_2426 CHROMOSOME_X 14084381 14084382 MACS_peak_2429 CHROMOSOME_X 14090303 14090304 MACS_peak_2430 CHROMOSOME_X 14098791 14098792 MACS_peak_2431 CHROMOSOME_X 14102115 14102116 MACS_peak_2432.1 CHROMOSOME_X 14104654 14104655 MACS_peak_2432.2 CHROMOSOME_X 14110330 14110331 MACS_peak_2434 CHROMOSOME_X 14115626 14115627 MACS_peak_2435 CHROMOSOME_X 14125981 14125982 MACS_peak_2436 CHROMOSOME_X 14127405 14127406 MACS_peak_2437 CHROMOSOME_X 14135057 14135058 MACS_peak_2438 CHROMOSOME_X 14137640 14137641 MACS_peak_2439 CHROMOSOME_X 14141549 14141550 MACS_peak_2440 CHROMOSOME_X 14146900 14146901 MACS_peak_2441 CHROMOSOME_X 14162684 14162685 MACS_peak_2444 CHROMOSOME_X 14169808 14169809 MACS_peak_2445 CHROMOSOME_X 14173983 14173984 MACS_peak_2446 CHROMOSOME_X 14175693 14175694 MACS_peak_2447 CHROMOSOME_X 14186616 14186617 MACS_peak_2449 CHROMOSOME_X 14194747 14194748 MACS_peak_2452 CHROMOSOME_X 14201143 14201144 MACS_peak_2453 CHROMOSOME_X 14224103 14224104 MACS_peak_2458 CHROMOSOME_X 14225487 14225488 MACS_peak_2459 CHROMOSOME_X 14228864 14228865 MACS_peak_2460 CHROMOSOME_X 14238405 14238406 MACS_peak_2462 CHROMOSOME_X 14243687 14243688 MACS_peak_2463.1 CHROMOSOME_X 14244617 14244618 MACS_peak_2463.2 CHROMOSOME_X 14287947 14287948 MACS_peak_2469 CHROMOSOME_X 14292842 14292843 MACS_peak_2470 CHROMOSOME_X 14297316 14297317 MACS_peak_2471 CHROMOSOME_X 14307604 14307605 MACS_peak_2473 CHROMOSOME_X 14312552 14312553 MACS_peak_2474 CHROMOSOME_X 14321339 14321340 MACS_peak_2475 CHROMOSOME_X 14324511 14324512 MACS_peak_2476 CHROMOSOME_X 14327573 14327574 MACS_peak_2477 CHROMOSOME_X 14329503 14329504 MACS_peak_2478 CHROMOSOME_X 14330861 14330862 MACS_peak_2479 CHROMOSOME_X 14334335 14334336 MACS_peak_2480 CHROMOSOME_X 14339069 14339070 MACS_peak_2481 CHROMOSOME_X 14341199 14341200 MACS_peak_2482 CHROMOSOME_X 14349798 14349799 MACS_peak_2483 CHROMOSOME_X 14409742 14409743 MACS_peak_2489 CHROMOSOME_X 14415850 14415851 MACS_peak_2490 CHROMOSOME_X 14421780 14421781 MACS_peak_2491 CHROMOSOME_X 14432440 14432441 MACS_peak_2492 CHROMOSOME_X 14436162 14436163 MACS_peak_2493 CHROMOSOME_X 14438052 14438053 MACS_peak_2494 CHROMOSOME_X 14447701 14447702 MACS_peak_2495 CHROMOSOME_X 14459078 14459079 MACS_peak_2498 CHROMOSOME_X 14480335 14480336 MACS_peak_2500 CHROMOSOME_X 14486577 14486578 MACS_peak_2502 CHROMOSOME_X 14489980 14489981 MACS_peak_2503.1 CHROMOSOME_X 14491298 14491299 MACS_peak_2503.2 CHROMOSOME_X 14495087 14495088 MACS_peak_2504 CHROMOSOME_X 14504391 14504392 MACS_peak_2507.1 CHROMOSOME_X 14506123 14506124 MACS_peak_2507.2 CHROMOSOME_X 14508995 14508996 MACS_peak_2508 CHROMOSOME_X 14513616 14513617 MACS_peak_2509.1 CHROMOSOME_X 14518002 14518003 MACS_peak_2509.2 CHROMOSOME_X 14523425 14523426 MACS_peak_2509.3 CHROMOSOME_X 14524085 14524086 MACS_peak_2509.4 CHROMOSOME_X 14524624 14524625 MACS_peak_2509.5 CHROMOSOME_X 14525862 14525863 MACS_peak_2509.6 CHROMOSOME_X 14527792 14527793 MACS_peak_2509.7 CHROMOSOME_X 14528578 14528579 MACS_peak_2509.8 CHROMOSOME_X 14528925 14528926 MACS_peak_2509.9 CHROMOSOME_X 14536125 14536126 MACS_peak_2510 CHROMOSOME_X 14540409 14540410 MACS_peak_2511 CHROMOSOME_X 14549785 14549786 MACS_peak_2513 CHROMOSOME_X 14557549 14557550 MACS_peak_2515.1 CHROMOSOME_X 14560332 14560333 MACS_peak_2515.2 CHROMOSOME_X 14561668 14561669 MACS_peak_2515.3 CHROMOSOME_X 14566291 14566292 MACS_peak_2516 CHROMOSOME_X 14568521 14568522 MACS_peak_2517 CHROMOSOME_X 14589189 14589190 MACS_peak_2518.1 CHROMOSOME_X 14589577 14589578 MACS_peak_2518.2 CHROMOSOME_X 14602800 14602801 MACS_peak_2520 CHROMOSOME_X 14610502 14610503 MACS_peak_2521 CHROMOSOME_X 14633964 14633965 MACS_peak_2524 CHROMOSOME_X 14636780 14636781 MACS_peak_2525.1 CHROMOSOME_X 14640055 14640056 MACS_peak_2525.2 CHROMOSOME_X 14649347 14649348 MACS_peak_2528 CHROMOSOME_X 14653607 14653608 MACS_peak_2529 CHROMOSOME_X 14656774 14656775 MACS_peak_2530 CHROMOSOME_X 14659493 14659494 MACS_peak_2531 CHROMOSOME_X 14671547 14671548 MACS_peak_2532 CHROMOSOME_X 14673943 14673944 MACS_peak_2533 CHROMOSOME_X 14679103 14679104 MACS_peak_2534 CHROMOSOME_X 14684545 14684546 MACS_peak_2535 CHROMOSOME_X 14688695 14688696 MACS_peak_2536 CHROMOSOME_X 14691411 14691412 MACS_peak_2537 CHROMOSOME_X 14693173 14693174 MACS_peak_2538 CHROMOSOME_X 14694735 14694736 MACS_peak_2539 CHROMOSOME_X 14711265 14711266 MACS_peak_2540 CHROMOSOME_X 14718329 14718330 MACS_peak_2541 CHROMOSOME_X 14720040 14720041 MACS_peak_2542 CHROMOSOME_X 14729513 14729514 MACS_peak_2544 CHROMOSOME_X 14740063 14740064 MACS_peak_2547 CHROMOSOME_X 14744652 14744653 MACS_peak_2548 CHROMOSOME_X 14747087 14747088 MACS_peak_2549 CHROMOSOME_X 14752182 14752183 MACS_peak_2550 CHROMOSOME_X 14753640 14753641 MACS_peak_2551 CHROMOSOME_X 14758084 14758085 MACS_peak_2552 CHROMOSOME_X 14761083 14761084 MACS_peak_2553 CHROMOSOME_X 14776448 14776449 MACS_peak_2556 CHROMOSOME_X 14780463 14780464 MACS_peak_2557 CHROMOSOME_X 14787021 14787022 MACS_peak_2559 CHROMOSOME_X 14790465 14790466 MACS_peak_2560 CHROMOSOME_X 14800114 14800115 MACS_peak_2561 CHROMOSOME_X 14804804 14804805 MACS_peak_2562 CHROMOSOME_X 14810862 14810863 MACS_peak_2563.1 CHROMOSOME_X 14813510 14813511 MACS_peak_2563.2 CHROMOSOME_X 14818818 14818819 MACS_peak_2563.3 CHROMOSOME_X 14823234 14823235 MACS_peak_2564 CHROMOSOME_X 14824832 14824833 MACS_peak_2565 CHROMOSOME_X 14828025 14828026 MACS_peak_2566 CHROMOSOME_X 14829705 14829706 MACS_peak_2567 CHROMOSOME_X 14841169 14841170 MACS_peak_2569 CHROMOSOME_X 14843010 14843011 MACS_peak_2570 CHROMOSOME_X 14853941 14853942 MACS_peak_2571 CHROMOSOME_X 14862996 14862997 MACS_peak_2574 CHROMOSOME_X 14875832 14875833 MACS_peak_2575 CHROMOSOME_X 14883739 14883740 MACS_peak_2576 CHROMOSOME_X 14886141 14886142 MACS_peak_2577 CHROMOSOME_X 14892678 14892679 MACS_peak_2578 CHROMOSOME_X 14915643 14915644 MACS_peak_2581 CHROMOSOME_X 14932167 14932168 MACS_peak_2584 CHROMOSOME_X 14938308 14938309 MACS_peak_2585 CHROMOSOME_X 14946818 14946819 MACS_peak_2586 CHROMOSOME_X 14955070 14955071 MACS_peak_2588 CHROMOSOME_X 14959583 14959584 MACS_peak_2589 CHROMOSOME_X 14962805 14962806 MACS_peak_2590 CHROMOSOME_X 14970475 14970476 MACS_peak_2591 CHROMOSOME_X 14975764 14975765 MACS_peak_2592.1 CHROMOSOME_X 14978804 14978805 MACS_peak_2592.2 CHROMOSOME_X 14983405 14983406 MACS_peak_2593 CHROMOSOME_X 14987866 14987867 MACS_peak_2594 CHROMOSOME_X 14992084 14992085 MACS_peak_2595 CHROMOSOME_X 15006951 15006952 MACS_peak_2599 CHROMOSOME_X 15011117 15011118 MACS_peak_2600 CHROMOSOME_X 15014042 15014043 MACS_peak_2601 CHROMOSOME_X 15019377 15019378 MACS_peak_2602 CHROMOSOME_X 15022202 15022203 MACS_peak_2603 CHROMOSOME_X 15032539 15032540 MACS_peak_2605 CHROMOSOME_X 15040833 15040834 MACS_peak_2606 CHROMOSOME_X 15045531 15045532 MACS_peak_2607 CHROMOSOME_X 15070263 15070264 MACS_peak_2609 CHROMOSOME_X 15076259 15076260 MACS_peak_2611 CHROMOSOME_X 15077726 15077727 MACS_peak_2612 CHROMOSOME_X 15080418 15080419 MACS_peak_2613 CHROMOSOME_X 15083921 15083922 MACS_peak_2614 CHROMOSOME_X 15100682 15100683 MACS_peak_2618 CHROMOSOME_X 15105194 15105195 MACS_peak_2619 CHROMOSOME_X 15108112 15108113 MACS_peak_2620 CHROMOSOME_X 15114578 15114579 MACS_peak_2621 CHROMOSOME_X 15128322 15128323 MACS_peak_2623 CHROMOSOME_X 15132210 15132211 MACS_peak_2624 CHROMOSOME_X 15142585 15142586 MACS_peak_2625 CHROMOSOME_X 15143872 15143873 MACS_peak_2626 CHROMOSOME_X 15146486 15146487 MACS_peak_2627 CHROMOSOME_X 15148919 15148920 MACS_peak_2628 CHROMOSOME_X 15151365 15151366 MACS_peak_2629 CHROMOSOME_X 15158855 15158856 MACS_peak_2630 CHROMOSOME_X 15173607 15173608 MACS_peak_2631 CHROMOSOME_X 15181570 15181571 MACS_peak_2632 CHROMOSOME_X 15189332 15189333 MACS_peak_2633 CHROMOSOME_X 15193949 15193950 MACS_peak_2634 CHROMOSOME_X 15200887 15200888 MACS_peak_2636 CHROMOSOME_X 15213552 15213553 MACS_peak_2638 CHROMOSOME_X 15215783 15215784 MACS_peak_2639 CHROMOSOME_X 15217568 15217569 MACS_peak_2640 CHROMOSOME_X 15222014 15222015 MACS_peak_2641 CHROMOSOME_X 15225407 15225408 MACS_peak_2642 CHROMOSOME_X 15232239 15232240 MACS_peak_2643 CHROMOSOME_X 15236158 15236159 MACS_peak_2645 CHROMOSOME_X 15246679 15246680 MACS_peak_2648 CHROMOSOME_X 15297483 15297484 MACS_peak_2655 CHROMOSOME_X 15307754 15307755 MACS_peak_2657 CHROMOSOME_X 15310409 15310410 MACS_peak_2658 CHROMOSOME_X 15312795 15312796 MACS_peak_2659 CHROMOSOME_X 15327350 15327351 MACS_peak_2660 CHROMOSOME_X 15340945 15340946 MACS_peak_2664 CHROMOSOME_X 15345869 15345870 MACS_peak_2665 CHROMOSOME_X 15347822 15347823 MACS_peak_2666 CHROMOSOME_X 15354028 15354029 MACS_peak_2667 CHROMOSOME_X 15365368 15365369 MACS_peak_2669 CHROMOSOME_X 15373867 15373868 MACS_peak_2670 CHROMOSOME_X 15376722 15376723 MACS_peak_2671 CHROMOSOME_X 15380548 15380549 MACS_peak_2672 CHROMOSOME_X 15385928 15385929 MACS_peak_2673 CHROMOSOME_X 15395874 15395875 MACS_peak_2675 CHROMOSOME_X 15405894 15405895 MACS_peak_2676 CHROMOSOME_X 15407598 15407599 MACS_peak_2677 CHROMOSOME_X 15420740 15420741 MACS_peak_2678 CHROMOSOME_X 15424157 15424158 MACS_peak_2679 CHROMOSOME_X 15431096 15431097 MACS_peak_2681 CHROMOSOME_X 15438923 15438924 MACS_peak_2682 CHROMOSOME_X 15441419 15441420 MACS_peak_2683 CHROMOSOME_X 15459901 15459902 MACS_peak_2685 CHROMOSOME_X 15470138 15470139 MACS_peak_2687 CHROMOSOME_X 15487178 15487179 MACS_peak_2690 CHROMOSOME_X 15492837 15492838 MACS_peak_2691 CHROMOSOME_X 15497045 15497046 MACS_peak_2692 CHROMOSOME_X 15508109 15508110 MACS_peak_2693 CHROMOSOME_X 15511732 15511733 MACS_peak_2695 CHROMOSOME_X 15515790 15515791 MACS_peak_2696 CHROMOSOME_X 15528562 15528563 MACS_peak_2699 CHROMOSOME_X 15535381 15535382 MACS_peak_2700 CHROMOSOME_X 15553127 15553128 MACS_peak_2701 CHROMOSOME_X 15555487 15555488 MACS_peak_2702 CHROMOSOME_X 15556873 15556874 MACS_peak_2703 CHROMOSOME_X 15566739 15566740 MACS_peak_2705.1 CHROMOSOME_X 15568092 15568093 MACS_peak_2705.2 CHROMOSOME_X 15568482 15568483 MACS_peak_2705.3 CHROMOSOME_X 15575536 15575537 MACS_peak_2706 CHROMOSOME_X 15577295 15577296 MACS_peak_2707 CHROMOSOME_X 15579594 15579595 MACS_peak_2708 CHROMOSOME_X 15590664 15590665 MACS_peak_2710 CHROMOSOME_X 15600669 15600670 MACS_peak_2712 CHROMOSOME_X 15607001 15607002 MACS_peak_2713.1 CHROMOSOME_X 15607637 15607638 MACS_peak_2713.2 CHROMOSOME_X 15613926 15613927 MACS_peak_2713.3 CHROMOSOME_X 15616538 15616539 MACS_peak_2714.1 CHROMOSOME_X 15617295 15617296 MACS_peak_2714.2 CHROMOSOME_X 15619442 15619443 MACS_peak_2715 CHROMOSOME_X 15626475 15626476 MACS_peak_2717 CHROMOSOME_X 15630858 15630859 MACS_peak_2718 CHROMOSOME_X 15634012 15634013 MACS_peak_2719 CHROMOSOME_X 15645844 15645845 MACS_peak_2720 CHROMOSOME_X 15647399 15647400 MACS_peak_2721 CHROMOSOME_X 15653305 15653306 MACS_peak_2722 CHROMOSOME_X 15655070 15655071 MACS_peak_2723 CHROMOSOME_X 15659620 15659621 MACS_peak_2724 CHROMOSOME_X 15664482 15664483 MACS_peak_2725 CHROMOSOME_X 15666272 15666273 MACS_peak_2726 CHROMOSOME_X 15668866 15668867 MACS_peak_2727 CHROMOSOME_X 15678640 15678641 MACS_peak_2729 CHROMOSOME_X 15680230 15680231 MACS_peak_2730 CHROMOSOME_X 15689346 15689347 MACS_peak_2732 CHROMOSOME_X 15696487 15696488 MACS_peak_2733 CHROMOSOME_X 15700570 15700571 MACS_peak_2734 CHROMOSOME_X 15704654 15704655 MACS_peak_2735 CHROMOSOME_X 15707506 15707507 MACS_peak_2736 CHROMOSOME_X 15709723 15709724 MACS_peak_2737 CHROMOSOME_X 15714831 15714832 MACS_peak_2738 CHROMOSOME_X 15721192 15721193 MACS_peak_2740 CHROMOSOME_X 15725089 15725090 MACS_peak_2741.1 CHROMOSOME_X 15725699 15725700 MACS_peak_2741.2 CHROMOSOME_X 15736585 15736586 MACS_peak_2741.3 CHROMOSOME_X 15741972 15741973 MACS_peak_2742 CHROMOSOME_X 15747957 15747958 MACS_peak_2743 CHROMOSOME_X 15754991 15754992 MACS_peak_2744 CHROMOSOME_X 15759325 15759326 MACS_peak_2745 CHROMOSOME_X 15766452 15766453 MACS_peak_2746 CHROMOSOME_X 15772839 15772840 MACS_peak_2748 CHROMOSOME_X 15779269 15779270 MACS_peak_2749 CHROMOSOME_X 15790602 15790603 MACS_peak_2750.1 CHROMOSOME_X 15791969 15791970 MACS_peak_2750.2 CHROMOSOME_X 15796632 15796633 MACS_peak_2751 CHROMOSOME_X 15798658 15798659 MACS_peak_2752 CHROMOSOME_X 15811473 15811474 MACS_peak_2753 CHROMOSOME_X 15841215 15841216 MACS_peak_2754 CHROMOSOME_X 15849445 15849446 MACS_peak_2755 CHROMOSOME_X 15860660 15860661 MACS_peak_2756 CHROMOSOME_X 15866286 15866287 MACS_peak_2757 CHROMOSOME_X 15871984 15871985 MACS_peak_2758 CHROMOSOME_X 15876994 15876995 MACS_peak_2759 CHROMOSOME_X 15878779 15878780 MACS_peak_2760 CHROMOSOME_X 15881305 15881306 MACS_peak_2761 CHROMOSOME_X 15906719 15906720 MACS_peak_2765 CHROMOSOME_X 15923081 15923082 MACS_peak_2767 CHROMOSOME_X 15950289 15950290 MACS_peak_2771 CHROMOSOME_X 15953727 15953728 MACS_peak_2772 CHROMOSOME_X 15983418 15983419 MACS_peak_2774 CHROMOSOME_X 15987023 15987024 MACS_peak_2775 CHROMOSOME_X 15993615 15993616 MACS_peak_2776 CHROMOSOME_X 15998434 15998435 MACS_peak_2777 CHROMOSOME_X 16002942 16002943 MACS_peak_2778 CHROMOSOME_X 16005815 16005816 MACS_peak_2779 CHROMOSOME_X 16012799 16012800 MACS_peak_2782 CHROMOSOME_X 16024639 16024640 MACS_peak_2784 CHROMOSOME_X 16033524 16033525 MACS_peak_2786 CHROMOSOME_X 16040341 16040342 MACS_peak_2787 CHROMOSOME_X 16044438 16044439 MACS_peak_2788 CHROMOSOME_X 16051843 16051844 MACS_peak_2789.1 CHROMOSOME_X 16052472 16052473 MACS_peak_2789.2 CHROMOSOME_X 16054407 16054408 MACS_peak_2789.3 CHROMOSOME_X 16055468 16055469 MACS_peak_2789.4 CHROMOSOME_X 16056516 16056517 MACS_peak_2789.5 CHROMOSOME_X 16057278 16057279 MACS_peak_2789.6 CHROMOSOME_X 16057611 16057612 MACS_peak_2789.7 CHROMOSOME_X 16059178 16059179 MACS_peak_2789.8 CHROMOSOME_X 16061283 16061284 MACS_peak_2790 CHROMOSOME_X 16069636 16069637 MACS_peak_2792 CHROMOSOME_X 16085206 16085207 MACS_peak_2795 CHROMOSOME_X 16086813 16086814 MACS_peak_2796 CHROMOSOME_X 16088022 16088023 MACS_peak_2797 CHROMOSOME_X 16095987 16095988 MACS_peak_2798 CHROMOSOME_X 16098249 16098250 MACS_peak_2799 CHROMOSOME_X 16105690 16105691 MACS_peak_2800 CHROMOSOME_X 16113785 16113786 MACS_peak_2801 CHROMOSOME_X 16118772 16118773 MACS_peak_2802 CHROMOSOME_X 16123510 16123511 MACS_peak_2803 CHROMOSOME_X 16127191 16127192 MACS_peak_2804 CHROMOSOME_X 16128946 16128947 MACS_peak_2805 CHROMOSOME_X 16131157 16131158 MACS_peak_2806 CHROMOSOME_X 16133927 16133928 MACS_peak_2807 CHROMOSOME_X 16144058 16144059 MACS_peak_2808.1 CHROMOSOME_X 16147800 16147801 MACS_peak_2808.2 CHROMOSOME_X 16150369 16150370 MACS_peak_2808.3 CHROMOSOME_X 16153428 16153429 MACS_peak_2809 CHROMOSOME_X 16155517 16155518 MACS_peak_2810.1 CHROMOSOME_X 16157020 16157021 MACS_peak_2810.2 CHROMOSOME_X 16159389 16159390 MACS_peak_2811 CHROMOSOME_X 16167097 16167098 MACS_peak_2812 CHROMOSOME_X 16173004 16173005 MACS_peak_2813 CHROMOSOME_X 16178157 16178158 MACS_peak_2814 CHROMOSOME_X 16180046 16180047 MACS_peak_2815 CHROMOSOME_X 16184145 16184146 MACS_peak_2817 CHROMOSOME_X 16189443 16189444 MACS_peak_2818 CHROMOSOME_X 16192714 16192715 MACS_peak_2819 CHROMOSOME_X 16200075 16200076 MACS_peak_2821 CHROMOSOME_X 16202396 16202397 MACS_peak_2822.1 CHROMOSOME_X 16203696 16203697 MACS_peak_2822.2 CHROMOSOME_X 16205916 16205917 MACS_peak_2822.3 CHROMOSOME_X 16208843 16208844 MACS_peak_2822.4 CHROMOSOME_X 16209795 16209796 MACS_peak_2822.5 CHROMOSOME_X 16218396 16218397 MACS_peak_2823 CHROMOSOME_X 16240863 16240864 MACS_peak_2824 CHROMOSOME_X 16252575 16252576 MACS_peak_2825 CHROMOSOME_X 16255100 16255101 MACS_peak_2826 CHROMOSOME_X 16263400 16263401 MACS_peak_2829 CHROMOSOME_X 16265953 16265954 MACS_peak_2830 CHROMOSOME_X 16271304 16271305 MACS_peak_2831 CHROMOSOME_X 16275360 16275361 MACS_peak_2832 CHROMOSOME_X 16284872 16284873 MACS_peak_2834 CHROMOSOME_X 16289415 16289416 MACS_peak_2835 CHROMOSOME_X 16294604 16294605 MACS_peak_2836 CHROMOSOME_X 16297202 16297203 MACS_peak_2837.1 CHROMOSOME_X 16298250 16298251 MACS_peak_2837.2 CHROMOSOME_X 16299378 16299379 MACS_peak_2837.3 CHROMOSOME_X 16300567 16300568 MACS_peak_2837.4 CHROMOSOME_X 16318391 16318392 MACS_peak_2840 CHROMOSOME_X 16319799 16319800 MACS_peak_2841 CHROMOSOME_X 16322014 16322015 MACS_peak_2842 CHROMOSOME_X 16324348 16324349 MACS_peak_2843 CHROMOSOME_X 16327399 16327400 MACS_peak_2844 CHROMOSOME_X 16331562 16331563 MACS_peak_2845.1 CHROMOSOME_X 16332036 16332037 MACS_peak_2845.2 CHROMOSOME_X 16337277 16337278 MACS_peak_2846 CHROMOSOME_X 16339837 16339838 MACS_peak_2847 CHROMOSOME_X 16342367 16342368 MACS_peak_2848 CHROMOSOME_X 16344166 16344167 MACS_peak_2849 CHROMOSOME_X 16349123 16349124 MACS_peak_2850 CHROMOSOME_X 16352613 16352614 MACS_peak_2851 CHROMOSOME_X 16354403 16354404 MACS_peak_2852 CHROMOSOME_X 16359914 16359915 MACS_peak_2853 CHROMOSOME_X 16363787 16363788 MACS_peak_2854 CHROMOSOME_X 16365238 16365239 MACS_peak_2855 CHROMOSOME_X 16376108 16376109 MACS_peak_2857 CHROMOSOME_X 16402156 16402157 MACS_peak_2858 CHROMOSOME_X 16410693 16410694 MACS_peak_2859 CHROMOSOME_X 16414724 16414725 MACS_peak_2860.1 CHROMOSOME_X 16418478 16418479 MACS_peak_2860.2 CHROMOSOME_X 16422385 16422386 MACS_peak_2861 CHROMOSOME_X 16424045 16424046 MACS_peak_2862 CHROMOSOME_X 16426023 16426024 MACS_peak_2863 CHROMOSOME_X 16435660 16435661 MACS_peak_2864 CHROMOSOME_X 16446024 16446025 MACS_peak_2868.1 CHROMOSOME_X 16447152 16447153 MACS_peak_2868.2 CHROMOSOME_X 16450969 16450970 MACS_peak_2869 CHROMOSOME_X 16457118 16457119 MACS_peak_2870 CHROMOSOME_X 16462912 16462913 MACS_peak_2871 CHROMOSOME_X 16484825 16484826 MACS_peak_2873 CHROMOSOME_X 16503142 16503143 MACS_peak_2877 CHROMOSOME_X 16509681 16509682 MACS_peak_2879 CHROMOSOME_X 16513741 16513742 MACS_peak_2880 CHROMOSOME_X 16521824 16521825 MACS_peak_2882 CHROMOSOME_X 16525754 16525755 MACS_peak_2883 CHROMOSOME_X 16527892 16527893 MACS_peak_2884 CHROMOSOME_X 16534671 16534672 MACS_peak_2886.1 CHROMOSOME_X 16536394 16536395 MACS_peak_2886.2 CHROMOSOME_X 16541279 16541280 MACS_peak_2887 CHROMOSOME_X 16551413 16551414 MACS_peak_2889 CHROMOSOME_X 16554020 16554021 MACS_peak_2890 CHROMOSOME_X 16555937 16555938 MACS_peak_2891 CHROMOSOME_X 16558854 16558855 MACS_peak_2892 CHROMOSOME_X 16564650 16564651 MACS_peak_2893 CHROMOSOME_X 16576618 16576619 MACS_peak_2896 CHROMOSOME_X 16579264 16579265 MACS_peak_2897 CHROMOSOME_X 16590934 16590935 MACS_peak_2899 CHROMOSOME_X 16597186 16597187 MACS_peak_2900 CHROMOSOME_X 16602563 16602564 MACS_peak_2902 CHROMOSOME_X 16604714 16604715 MACS_peak_2903 CHROMOSOME_X 16611476 16611477 MACS_peak_2904.1 CHROMOSOME_X 16612452 16612453 MACS_peak_2904.2 CHROMOSOME_X 16617808 16617809 MACS_peak_2904.3 CHROMOSOME_X 16625503 16625504 MACS_peak_2905 CHROMOSOME_X 16626997 16626998 MACS_peak_2906 CHROMOSOME_X 16628662 16628663 MACS_peak_2907 CHROMOSOME_X 16631998 16631999 MACS_peak_2908 CHROMOSOME_X 16634834 16634835 MACS_peak_2909 CHROMOSOME_X 16641824 16641825 MACS_peak_2910 CHROMOSOME_X 16646412 16646413 MACS_peak_2911 CHROMOSOME_X 16648951 16648952 MACS_peak_2912 CHROMOSOME_X 16655434 16655435 MACS_peak_2913 CHROMOSOME_X 16663250 16663251 MACS_peak_2915 CHROMOSOME_X 16669532 16669533 MACS_peak_2917 CHROMOSOME_X 16671386 16671387 MACS_peak_2918 CHROMOSOME_X 16676025 16676026 MACS_peak_2920 CHROMOSOME_X 16682035 16682036 MACS_peak_2921.1 CHROMOSOME_X 16683192 16683193 MACS_peak_2921.2 CHROMOSOME_X 16684679 16684680 MACS_peak_2921.3 CHROMOSOME_X 16687468 16687469 MACS_peak_2922 CHROMOSOME_X 16691705 16691706 MACS_peak_2923 CHROMOSOME_X 16695330 16695331 MACS_peak_2924 CHROMOSOME_X 16704237 16704238 MACS_peak_2925 CHROMOSOME_X 16715835 16715836 MACS_peak_2927 CHROMOSOME_X 16722068 16722069 MACS_peak_2928 CHROMOSOME_X 16726947 16726948 MACS_peak_2930 CHROMOSOME_X 16733564 16733565 MACS_peak_2931 CHROMOSOME_X 16754383 16754384 MACS_peak_2933 CHROMOSOME_X 16770981 16770982 MACS_peak_2935 CHROMOSOME_X 16775720 16775721 MACS_peak_2936 CHROMOSOME_X 16778613 16778614 MACS_peak_2937 CHROMOSOME_X 16786136 16786137 MACS_peak_2938 CHROMOSOME_X 16793552 16793553 MACS_peak_2939 CHROMOSOME_X 16796892 16796893 MACS_peak_2940 CHROMOSOME_X 16799236 16799237 MACS_peak_2941 CHROMOSOME_X 16814116 16814117 MACS_peak_2943.1 CHROMOSOME_X 16815532 16815533 MACS_peak_2943.2 CHROMOSOME_X 16818900 16818901 MACS_peak_2944 CHROMOSOME_X 16821765 16821766 MACS_peak_2945 CHROMOSOME_X 16848750 16848751 MACS_peak_2947 CHROMOSOME_X 16869183 16869184 MACS_peak_2950 CHROMOSOME_X 16872557 16872558 MACS_peak_2951 CHROMOSOME_X 16882694 16882695 MACS_peak_2952 CHROMOSOME_X 16886730 16886731 MACS_peak_2953 CHROMOSOME_X 16890589 16890590 MACS_peak_2954 CHROMOSOME_X 16901985 16901986 MACS_peak_2956 CHROMOSOME_X 16928240 16928241 MACS_peak_2962.1 CHROMOSOME_X 16929665 16929666 MACS_peak_2962.2 CHROMOSOME_X 16930937 16930938 MACS_peak_2962.3 CHROMOSOME_X 16931558 16931559 MACS_peak_2962.4 CHROMOSOME_X 16932372 16932373 MACS_peak_2962.5 CHROMOSOME_X 16932496 16932497 MACS_peak_2962.6 CHROMOSOME_X 16934144 16934145 MACS_peak_2963 CHROMOSOME_X 16940294 16940295 MACS_peak_2964.1 CHROMOSOME_X 16941147 16941148 MACS_peak_2964.2 CHROMOSOME_X 16941289 16941290 MACS_peak_2964.3 CHROMOSOME_X 16941436 16941437 MACS_peak_2964.4 CHROMOSOME_X 16942566 16942567 MACS_peak_2964.5 CHROMOSOME_X 16943338 16943339 MACS_peak_2964.6 CHROMOSOME_X 16944104 16944105 MACS_peak_2964.7 CHROMOSOME_X 16961511 16961512 MACS_peak_2966 CHROMOSOME_X 16976381 16976382 MACS_peak_2967 CHROMOSOME_X 16988435 16988436 MACS_peak_2971 CHROMOSOME_X 16991553 16991554 MACS_peak_2972 CHROMOSOME_X 17003779 17003780 MACS_peak_2973 CHROMOSOME_X 17025561 17025562 MACS_peak_2976 CHROMOSOME_X 17030612 17030613 MACS_peak_2977 CHROMOSOME_X 17033631 17033632 MACS_peak_2978 CHROMOSOME_X 17040386 17040387 MACS_peak_2980 CHROMOSOME_X 17043018 17043019 MACS_peak_2981 CHROMOSOME_X 17058466 17058467 MACS_peak_2984 CHROMOSOME_X 17063843 17063844 MACS_peak_2985 CHROMOSOME_X 17070411 17070412 MACS_peak_2986 CHROMOSOME_X 17092569 17092570 MACS_peak_2987 CHROMOSOME_X 17097850 17097851 MACS_peak_2988 CHROMOSOME_X 17104993 17104994 MACS_peak_2989 CHROMOSOME_X 17108791 17108792 MACS_peak_2990 CHROMOSOME_X 17115710 17115711 MACS_peak_2992 CHROMOSOME_X 17134740 17134741 MACS_peak_2993 CHROMOSOME_X 17139043 17139044 MACS_peak_2994 CHROMOSOME_X 17150133 17150134 MACS_peak_2995.1 CHROMOSOME_X 17151258 17151259 MACS_peak_2995.2 CHROMOSOME_X 17161009 17161010 MACS_peak_2996 CHROMOSOME_X 17172905 17172906 MACS_peak_2998 CHROMOSOME_X 17177247 17177248 MACS_peak_2999 CHROMOSOME_X 17181339 17181340 MACS_peak_3000 CHROMOSOME_X 17184503 17184504 MACS_peak_3001 CHROMOSOME_X 17192975 17192976 MACS_peak_3004 CHROMOSOME_X 17200389 17200390 MACS_peak_3006 CHROMOSOME_X 17201682 17201683 MACS_peak_3007 CHROMOSOME_X 17203602 17203603 MACS_peak_3008 CHROMOSOME_X 17216282 17216283 MACS_peak_3011.1 CHROMOSOME_X 17217583 17217584 MACS_peak_3011.2 CHROMOSOME_X 17222012 17222013 MACS_peak_3012 CHROMOSOME_X 17235874 17235875 MACS_peak_3013.1 CHROMOSOME_X 17236308 17236309 MACS_peak_3013.2 CHROMOSOME_X 17239287 17239288 MACS_peak_3013.3 CHROMOSOME_X 17284047 17284048 MACS_peak_3018 CHROMOSOME_X 17297584 17297585 MACS_peak_3019 CHROMOSOME_X 17302004 17302005 MACS_peak_3020 CHROMOSOME_X 17305157 17305158 MACS_peak_3021 CHROMOSOME_X 17315008 17315009 MACS_peak_3023 CHROMOSOME_X 17317447 17317448 MACS_peak_3024 CHROMOSOME_X 17327383 17327384 MACS_peak_3026 CHROMOSOME_X 17330619 17330620 MACS_peak_3028 CHROMOSOME_X 17339822 17339823 MACS_peak_3029 CHROMOSOME_X 17353560 17353561 MACS_peak_3030 CHROMOSOME_X 17369675 17369676 MACS_peak_3032 CHROMOSOME_X 17386463 17386464 MACS_peak_3035 CHROMOSOME_X 17392286 17392287 MACS_peak_3036 CHROMOSOME_X 17403926 17403927 MACS_peak_3038 CHROMOSOME_X 17408936 17408937 MACS_peak_3039 CHROMOSOME_X 17413821 17413822 MACS_peak_3040 CHROMOSOME_X 17416638 17416639 MACS_peak_3041 CHROMOSOME_X 17427849 17427850 MACS_peak_3043 CHROMOSOME_X 17434426 17434427 MACS_peak_3044 CHROMOSOME_X 17451511 17451512 MACS_peak_3045 CHROMOSOME_X 17456828 17456829 MACS_peak_3046 CHROMOSOME_X 17458997 17458998 MACS_peak_3047 CHROMOSOME_X 17462047 17462048 MACS_peak_3048.1 CHROMOSOME_X 17464053 17464054 MACS_peak_3048.2 CHROMOSOME_X 17466718 17466719 MACS_peak_3049 CHROMOSOME_X 17471945 17471946 MACS_peak_3050 CHROMOSOME_X 17475026 17475027 MACS_peak_3051 CHROMOSOME_X 17479704 17479705 MACS_peak_3052.1 CHROMOSOME_X 17480257 17480258 MACS_peak_3052.2 CHROMOSOME_X 17496859 17496860 MACS_peak_3054 CHROMOSOME_X 17505859 17505860 MACS_peak_3055 CHROMOSOME_X 17508445 17508446 MACS_peak_3056 CHROMOSOME_X 17513627 17513628 MACS_peak_3058 CHROMOSOME_X 17516942 17516943 MACS_peak_3059.1 CHROMOSOME_X 17520984 17520985 MACS_peak_3059.2 CHROMOSOME_X 17528253 17528254 MACS_peak_3059.3 CHROMOSOME_X 17534247 17534248 MACS_peak_3060 CHROMOSOME_X 17537757 17537758 MACS_peak_3061 CHROMOSOME_X 17540068 17540069 MACS_peak_3062.1 CHROMOSOME_X 17544462 17544463 MACS_peak_3062.2 CHROMOSOME_X 17545648 17545649 MACS_peak_3062.3 CHROMOSOME_X 17562320 17562321 MACS_peak_3065 CHROMOSOME_X 17587060 17587061 MACS_peak_3067 CHROMOSOME_X 17598819 17598820 MACS_peak_3068 CHROMOSOME_X 17610491 17610492 MACS_peak_3070 CHROMOSOME_X 17628500 17628501 MACS_peak_3072 CHROMOSOME_X 17631057 17631058 MACS_peak_3073.1 CHROMOSOME_X 17631801 17631802 MACS_peak_3073.2 CHROMOSOME_X 17633283 17633284 MACS_peak_3074 CHROMOSOME_X 17640928 17640929 MACS_peak_3075 CHROMOSOME_X 17650048 17650049 MACS_peak_3076 CHROMOSOME_X 17656861 17656862 MACS_peak_3077 CHROMOSOME_X 17667342 17667343 MACS_peak_3078 CHROMOSOME_X 17670589 17670590 MACS_peak_3080 CHROMOSOME_X 17677523 17677524 MACS_peak_3081 CHROMOSOME_X 17688389 17688390 MACS_peak_3083 CHROMOSOME_X 17714492 17714493 MACS_peak_3084.1 CHROMOSOME_X 17714970 17714971 MACS_peak_3084.2 CHROMOSOME_X 17717203 17717204 MACS_peak_3085