#NEXUS [written Fri Dec 14 10:16:15 EST 2018 by Mesquite version 3.51 (build 898)] BEGIN TAXA; DIMENSIONS NTAX=104; TAXLABELS Nematoda Priapulida Hallucigenia Collinsium Luolishania Aysheaia Tardigrada Onychophora Jianshanopodia Megadictyon Opabinia Anomalocaris Hurdia Peytoia Schinderhannes Surusicaris Isoxys Yohoia Haikoucaris Leanchoiliidae Fortiforceps Kuamaia Marrella Burgessia Emeraldella Sidneyia Aglaspis Cindarella Kodymirus Retifacies Emucaris Tegopelte Cheloniellon Kwanyinaspis Naraoia Arthroaspis Olenoides Triarthrus Hongshiyanaspis Xandarella Sanctacaris Habelia Offacolus Dibasterium Chasmataspis Weinbergina Megalograptus Limulus Scorpiones Opiliones Araneae Archegozetes Shankouia Fuxianhuia Chengjiangocaris Euthycarcinoidea Odaraia Canadaspis Branchiocaris Tokummia Waptia Tanazios1 Tanazios2 Cascolus Nereocaris Perspicaris Arthropleura Pauropoda Symphyla Chilopoda Diplopoda Ostracoda Argulus Artemia Triops Copepoda Lepidocaris Cephalocarida Remipedia Nahecaris Cinerocaris Isopoda Euphausiacea Anaspidacea Stomatopoda Leptostraca Allacma Anurida Zygentoma Odonata Coleoptera Diptera Hymenoptera Oelandocaris cf.Leanchoilia_larva cf.Leanchoilia_metanauplius Yicaris Rehbachiella Bredocaris Martinssonia Skara Phospatocopina Agnostus_larva Peronopsis ; END; BEGIN CHARACTERS; DIMENSIONS NCHAR=225; FORMAT DATATYPE = STANDARD GAP = - MISSING = ? SYMBOLS = " 0 1 2 3 4 5 6"; MATRIX Nematoda 00-0-----0------0--0---------0-0-0------------------------------------0-----0--------------------------------------------------010000--0-------000-----------------------0-----------------------------------0------------------- Priapulida 00-0-----0------0--0---------0-0-0------------------------------------0-----0--------------------------------------------------010010--0-------000-----------------------0-----------------------------------0------------------- Hallucigenia 10-0-011001100111--0---------0-0-0-------------???????---------------00-----0--------------------------------------------------012100--0-------????----------------------?0---------------------------------40------------------- Collinsium 10-0-011011110111--0---------0-0-0-------------???????---------------00-----0--------------------------------------------------01001???????????????----------------------?0---------------------------------40-------------?----- Luolishania 10-0-01101111?111--0---------0-0-0-------------???????---------------00-----0--------------------------------------------------01??00--0-------????----------------------?0---------------------------------00------------------- Aysheaia 10-0-110010--1011--0---------0-0-0-------------???????---------------00-----0--1--0--------------------------------------------0110?0--0-------????----------------------?0---------------------------------00------------------- Tardigrada 11?0-10000?--1010--0---------0-0-0-------------00000?0---------------00-----0--------------------------------------------------011000--0-------010?----------------------00---------------------------------10------------------- Onychophora 10-0-110010--2001--0---------0-0-0-------------00000?0---------------00-----0--------------------------------------------------001000--0-------1010----------------------00---------------------------------10------------------- Jianshanopodia 10-0-1100110-?00?--0---------0-0-0-------------???????---------------00-----0--1--0--------------------------------------------01??0???1101000-????----------------------?0---------------------------------?0------------------- Megadictyon 10-0-1100??????0?--0---------0-0-0-------------???????---------------00-----0--1--0--------------------------------------------01??00--1101000-????----------------------?0---------------------------------?0------------------- Opabinia 1100-10-1--------1111????--100?0-0---------0---???????0?????0--------10?0-000??1?00--------------------------------------------00???0--1111000-????----------------------?0----------101??------------------?0--000-------------- Anomalocaris 1100-?0-1-0------0-11???0--10??101-------------???????0?0---0--------10-0-0000010111-------?-----------------------------------002100--1101000-????----------------------?0----------1010?------------------?0--000-------------- Hurdia 1100-?0-1-0------0-11????0-10????100101-------????????0?0---0--------10-0-0000011110-------?-----------------------------------00210???????????????----------------------?0----------1010?------------------?????00?-------?----- Peytoia 1100-10-1-0------0-11??????10??????????????---????????0?0---0--------10-0-0000011111-------?-----------------------------------002100--????????????----------------------?0----------10???------------------?0--000--------?----- Schinderhannes 110?-1???--------0-11??????100?????????????0-?????????0????????------10?0-00000111??-------------------------------------------00210???????????????----------------------?0----------?????0-----------------?110??00-------?----- Surusicaris 1100-0100--------0-11????0-10100-100110----0-00???????0?0---0--------10-0-0011-11011-------1-----------------000---------------00??0???????????????----------------------?10---------10--00---------?------040????0?-------?----- Isoxys 1100-????--------0-11????0-10??0-100110----0-00???????0?????0--------10?0-001?0?00?1?????????????-???--?????-??0------??0????--00??0???1111000-????----------------------????????????????00---------?---?????0--000-------------- Yohoia 1110-00-0--------0-11????0-0-100-11----00000-00???????1????????0??00-10?110011-0--0-????00110?01?01?0???130000000-----??00-0-00?0??00--0-------????0-1100000-101010-?00--?1?00?0???0-111001010-00????0001001?11200000-0-------0-- Haikoucaris 1110-00-0--------??11?????-0-10??11----00000-?0????????????????0???0-1??110011-0--0-????001?0???????0???1300???00-----??0?-0-0000??0???????????????0-0100000-1010??-?00--????????????111001010?00????00??????11??0000-0----?--0-- Leanchoiliidae 1110-00-0--------1011??0?0-0-100-11----000?0-00?1?????1?1???0--0??00-1001110?1-0--0-010000110101101?0???130000000-------00-0-00?0??0??01111000-????0-0100000-10?111-100--?1?00?0???0-1110010101000-1?0001001611200000-0-------0-- Fortiforceps 1110-00-0--------0-11?????-1110??11----00000-?????????1????????0???0-1??110001-0--0-????001?0???????0???1300?0?00-----??0?-0-00?0??00--0-------????0-0-00010-1010??-020--????????????11??01000?00????0000000?111?0000-0-------0-- Kuamaia 1110-00-0--------0-11????1-0-1?1011----00?00-?0???????1?10--1010????-1000-000010--0-??0000110101001101?4140010?00-------00-0-0000??0???????????????0-0?00000-10010--100--?1????????101110?11101?0110?001????010-?0000-10000?00001 Marrella 1110-00-0--------0-0---------100-11----10?00-10???????1?1???1010??00-1000-000010--0-1---0-1101011000010-110000?00-------00-0-00?03101100-------????0-0-00010-01-----000--?1????????0-1110012101010--?0000000010-?0000-0-------0-1 Burgessia 1110-00-0--------0-??????0-??10??101-------0-00???????1????????0??00-10?0-000010--0-1---?-111??100100?0??????0?00-------00-0-00?0??011010---111????0-030-000-0?-----100--?1??????????11--01-1010----?001010??110?0000-0----0--0-0 Emeraldella 1110-00-0--------0-??????0-??110-11----00?00-10???????1?10--10101?00-1000-100010--0-??00?011?10100??0?0?140010?00-----??00-0-00?03?0???1120?00-????0-0100000-100110-100--?1?00?0???10111001010100110?0011101211010000-0--0----001 Sidneyia 1110-00-0--------0-11????0-101?0-11----00000-10???????1?10--1010??00-1000-000010--0-??00?011?10100??0?0?140010100-----??00-0-00?0310???11???00-????0-1100000-100111-100--?110010???10111001010-00110?1011101211110000-0--0----001 Aglaspis 1111-00-0--------0-11????100-1?0-11----00100-10???????1?1???1??0???0-1??0-000010--0-????001101010???01??????0??00-----??0?-0-00003?0???????????????0-0100000-100110-100--?1????????1011?-01?10100?????00?????11000100-0--00?--01? Cindarella 1110-00-0--------0-11????0-10110-11----00000-?0???????1?10--11101000-1000-000010--0-1---??110?01001?01?21200?0?00-------00-0-000031011011010100????0-00000011100110-?00--?1????0---??111001010100100??0???0?210-00000-0--0----100 Kodymirus 111?-00-0--------0-???????-??100-11----00000-10???????1?10--10101000-1000-0000?0--0-?????-1??1011001010-1?0010100-----??00-0-0000310???????????????0-0100000-100110-100--?10-------1011?-010101001???101?101?110?0000-0--00?--00? Retifacies 1110-00-0--------0-11????0-101?0-11----00000-10???????1?10--1010??00-1000-000010--0-1---?-110?01001?010?1?00?0100-------00-0-0000310???????????????0-0100000-100110-?00--?10-------10111001010100?????0??10??110?0000-1000-?01001 Emucaris 1110-00-0--------?-??????0-??100-11----?0?00-?0???????1?10--1110??00-1000-000010--0-1---?-110??10???0??????0???00-------0?-0-0000310???????????????0-0?0000?-1001??-??0--????????????11??01?1????????????????10-?0000-10100?0000? Tegopelte 1110-00-0--------?-??????0-??1-0-11-----0000-?0???????1?10--1110??00-1000-0000?0--0-1---?-110??10?1?0??21200?0100-------00-0-0000310110112?0000????0-00-000?-1001---??0--?1????????10111001110100??0??0??????10-00000---00--0100? Cheloniellon 1110-00-0--------?-1?????100-1?0-11----00001010???????1?1???1110??00-10?0-000010--0-1---?-110001?0?0??0?????00100-------00-0-0000310???????????????0-1100000-10011?-000--?10-------10110-0101010?????0000000610-?1000-0--10?--000 Kwanyinaspis 1110-00-0--------?-11????1-0-1???11----00000-?0????????????????0???0-100????0?????????????110?01?0110102120010100-------00-0-000031????????????????0-?10000?-100110-100--?10-------10111001010100????101010??110?0000-0-?10?--00? Naraoia 1110-00-0--------0-11???-0-??110-11----00000-10???????1?10--11101?00-1000-000010--0-1---0-11010100110102120010100-------00-0-000031011011201111????0-000000?-1001---100--?10-------10111001110100100?0010100?10-?0000-1101-10(0 1)0(0 1)? Arthroaspis 1110-00-0--------0-??????0-??10??11----00000-10???????1?10--11101000-1000-000010--0-1---??110?01001101?4140010?00-----??00-0-00?0310110112?0000????0-0000000-100110-100--?10-------1?111001010100110?101?101210-00000-1010--0100? Olenoides 1111-00-0--------0-11???0110-100-11----01100-10???????1?10--10101?00-1000-000010--0-1---0-11010100110102120010100-------00-0-00003101101120?00-????0-0100000-100111-100--?10-------10111001010100100?0011101210-01000-10101-10011 Triarthrus 1111-00-0--------0-11???0100-100-11----01100-10???????1?10--11101000-1000-000010--0-1---0-11010100110101110010100-------00-0-0000310???????????????0-0-00010-100111-100--?10-------1011100121010100-00011101210-00000-10101?100(0 1)0 Hongshiyanaspis 1111-00-0--------0-11???0110-100-11----01100-10???????1?10--1110??00-1000-000010--0-1---0-11010100110104140010100-------00-0-00003101101120000-????0-0000000-100111-100--?10-------10111001010100110?001??0?210-00000-10101-10011 Xandarella 1110-00-0--------0-11????100-130-11----0?000-10???????1?10--11101?00-1010-000010--0-1---0-110121001???021200?0?00-------00-0-00003?0???1120100-????0-01000011100110-120--??????0?????11100101010010-?00??????10-00000-0--00---10? Sanctacaris 1110-00-0--------0-11????0-0-130-11----00001010???????1?10--1110????-1011?0001-0--0-1---0-011101000?0??-15?110100-------00-0-000031010?0??????????????110000-1010??-???--?10-----??1?111011010-000-1010?????211210000-0----?--00- Habelia 1110-00-0--------0-11????0-0-130-11----00001010???????1?10--1110????-10?1?0001-0--0-1---0-011101000?0??-150110100-------00-0-000031010?0-------????110110000-101011-100--?10-------10111011010-000-101000000211010000-0-------00- Offacolus 1110-00-0--------??11????????13??11----00?01010???????1?????0--0???--1??0-0101-0--0-1---0-1111010110000-101101?00-------01-0-0000310???0-------????1-1310100-10100--100--?1??????????1110113-0--01--000-0000-11000000-0-------00- Dibasterium 1110-00-0--------????????0???120-11----00001010???????1?????0--0???--1??0-0101-0--0-1---0-1111010110000-101101?00-------01-0-0000310???????????????1-1111100-10100----0--?1??????????1110113-0--01--000-0000-11010000-0----?--00- Chasmataspis 1110-00-0--------1011????100-1??-11----10100-10???????1????????01110-10?0-0101-0--0-1---1-111101?100000-0---00?00-------01-0-00003?????????????????1-11?1100-10000--0-0--?1???00?????????11?-?--????001-0000-11000000-0--10?--01- Weinbergina 1110-00-0--------1011?????0??13?-11----00?00-10???????1????????0???0-10?0-0101-0--0-1---1-1111010100000-0---00100-------00-0-00003?????????????????0-0?0?100-10000--0-0--?1????????10????11?-?--????000-0000211000000-0--0-?--00- Megalograptus 1110-00-0--------0-11????100-12?-11----10000-10???????1?1???0--01110-1000-0101-0--0-1---1-1?11011?00000-0---10100-------00-0-0000310???????????????1-111?100-10000--0-0--?10-------1?1111113-0--00--011-1001211?00000-0--10?--00- Limulus 1110-00-0--------10110??0110-130-11----00110-1010000001110--0--0110--1000-0101-0--0-1---1-1111010100000-0---01100-------01-0-00003101100----11101110-0110100-10000--0-0--011001000-101110110-0--011-001-0000-11000000-1111000100- Scorpiones 1110-00-0--------101?0000100-120-11----10000-1010000?01110--0--01110-1000-0101-0--0-1---1-1111011100000-0---00000-------00-0-00003100--1120000-01111-1111100-00-----0-0--110-------111111113-0--00--000-0000411000000-0---0---0-- Opiliones 1110-00-0--------?-10?000100-120-11----10000-10100????1110--0--0111--10?0-0101-0--0-1---1-1111011000000-0---00000-------00-0-00003100--11200?0?011?0-1310000-00-----0-0--110-------?10---11---------000-0000610-00000-0---0?--0-- Araneae 1110-00-0--------1?100010100-120-11----11000-1010000001110--0--0110--1000-0101-0--0-1---1-1111011000000-0---00000-------00-0-000031010010---11001110-13-0000-00-----0-0--110-------111111113-0--00--000-0000410-00000-0---0---0-- Archegozetes 1110-00-0--------0-0-0-------121111----10000-10100????1110--0--011?--1010-0101-0--0-1---1-1111011000000-0---00000-------00-0-00003100--11201110011?0-13-0000-00-----0-0--110-------0-0---11---------000-0000610-00000-0-------0-- Shankouia 1110-00-0--------??11????00111?10101-------1-10???????1?11101000??00-1?00-000000--0-0??????????11000000-?????0011102?0??00-0?0010310???0-------?1??1-0-0-010-10010--020--????????????110-010101000-?00000000?110?0000-0-------0-0 Fuxianhuia 1110-00-0--------??11??2000111?101000------1-10?????????11101000??00-1100-000000--0-0110?0110020--00000-???0?0011102?0??00-0?0010310???0-------?1??1-1-0-01??1101??-020--????????????110-01010-000-?00000000?110?000000-------0-0 Chengjiangocaris 1110-00-0--------??11????00111?10101-------1-100????????11101000??00-1?00-000000--0-0110?0110020--00000-?---00011102?0??00-0?0010310???0-------?1??1-?-0-011010010--020--????????????110-010101000-?00000000?1????????0-------0-0 Euthycarcinoidea 1110-00-0--------0-11????00111?1011----10001110???????1?11??10?0??00-10?0-000000--0-011????????0--------0---?001-?0?????00-0?0??0310???????????????1-1-0?0110110010-020--??????????0-0---01-1-------000000006110?0000-0--1-?--0-0 Odaraia 1110-00-0--------0-11????0-111???100110----?-10???????1????????0??00-1????0????0--0-?????????????????????????0???????0??0???????0??0???1???0?0-??????0-0-010-01-----000--?111??0???0-110-01?101??????00?000??1101100000-------0-- Canadaspis 1110-00-0--------0-11????0-101?1?100101----???0???????1????????0??00-11?0-000000--0-???????1???????00?0??????????????0??00???0??0??0???????????????10110-000-01-----00????1111?0???0-111001010-0?????0011001210-1110000----?--0-0 Branchiocaris 1110-00-0--------0-??????0-?01110100100----1-10???????1?10--0--01?00-10?0-000000--0-011???00???0--------?????001-?02?0??0????00?0310???0-------????1?0-0-010-01-----101-1?1111?0???0-110-0101010?????000?00??10-010000???-----0-0 Tokummia 1110-00-0--------0-??????0-?01111100100----1-10???????1?10--0--01?00-10?0-000000--0-011???001??0--------?????001-?0200??00???00?0310???0-------????1?0-0-010-01-----101-1?111110???0-110-0101010?????0001001410-010001???-----0-0 Waptia 1110-00-0--------0-11????0-111?11100111----1-10???????1?10--0--0???0-11?0-000010--0-011111010110--------0---0001-102001100-0100?0310???0-------????1-111-001101-----0-0--?11?1?????0-0-1011--0-010--?00-0010410-0100020-------0-0 Tanazios1 1110-00-0--------0-0---------120-11----00001110?????????1100???0???0-1000-0001-0--0-?01000110??110000?0-1??0?0?1010?00--00-000110310???????????????0-0-00010-1011??-0?0--?110??10??0-110-0150100???-100??????10-?1100?0----?--0-? Tanazios2 1110-00-0--------0-0---------120-11----00001110?????????1100???0???0-1100-000010--0-?11000110??0--------1??0?0?1-10?00--00-000110310???????????????0-0-00010-1011??-0?0--?110??10??0-110-0150100???-100??????10-?1100?0----?--0-? Cascolus 1110-00-0--------0-11????0-11110-11----11000010???????1?1100???01000-100101001-0--0-011000010??11?100?0013000001010300--00-0000?0310???????????????0-?100000-00-----0?0--?1100?10??0-110-01000?0???11000?????1????????0----?--0-0 Nereocaris 1110-00-0--------0-11????0-111???100110----?-?0????????????????0???0-1????0????0--0-????????????????????????????????????????????0310?????????????????1-?-01??01-----0????????????????????01?0?-???????00?????10-0100000----?--0-? Perspicaris 1110-00-0--------0-11????0-111?0-100101----?-10????????????????0???0-10?0-000000--0-????????????????????????????????????????????0310?????????????????10?-00??01-----?????????????????11???10??-???????0??????10-0100000----?--0-? Arthropleura 1110-00-0--------0-0---------11??11----1000111????????1?????????????-1??0-000010--0-0111?10?1???????????????????0???????????????0310???????????????0-0-000111100010-020--?11?00001?0-0---01-1-1-----?0001000610-?0000-0--0-?--0-? Pauropoda 1110-00-0--------0-0---------110-11----1000111001?????10111110001001-100111001-0--0-011111001--0--------0---0001-000010-10-1001103100--0-------111?0-0200001100-----100--01100000100-0---01-1-1-----00000000610-00000-0-------0-0 Symphyla 1111-00-0--------0-0---------110-11----1000111001?????10111110011001-1000-000010--0-011111001--0--------0---0001-01001100000001103100--0-------111?0-0100001200-----100--01100000100-0---01-1-1-----00000000610-0100030-------0-0 Chilopoda 1110-00-0--------0-1?0011100-110-11----10001110010100010111010001001-1000-000010--0-0111110011-0--------0---0001-01?111000-0101103100--0-------11110-0-00010-00-----1010101100000100-0---01-1-1-----00000000611100000-0---0---0-0 Diplopoda 1111-00-0--------0-120011100-110-11----10001110010??0010111110001001-1000-000000--0-011111001--0--------0---0001-010110-10-1001103100--0-------11110-0-00011100-----100--01100000100-0---01-1-1-----00000000610-00000-0---0---0-0 Ostracoda 1111-00-0--------1-0---------110-100100----1-100??????101100???01100-1000-000000--0-0111110011111010001013--0001010?00?-0010001103101000-------?11?0-110-000-01-----110--011001000-0-110-01000-000-000000010?10-0100010-------0-1 Argulus 1110-00-0--------0-11?1?0100-11??11-----1000-1001?????1011000--?1?0--100??0001-0--0-0111110011111000001-0---1001000?000-00-01?01031010010---110?11?1-1400000-00-----000--011000100-0-110-01-00-100--00000010-10-0?0?0?0---0---0-0 Artemia 1110-00-0--------111111100-11110-11----1000110000110001011000--01000-0?0--000--0--0-011111001--110?0001-0---10010001000-0011001103100--10---111111?101-00010-01-----0-0--011011000-0-110-01501-000--10000010-10-0100030----0--0-0 Triops 1111-00-0--------12111110100-110-1001110---1-10001100010111110001000-000--000--0--0-011111001--11000001-0---10010002000-0011001103100--10---1111110101-0-011101-----0-11?011011000-0-110-01501-000--10000010-10-0100010---00--0-0 Copepoda 1110-00-0--------0-11--------11??11----10000-1001?????101101???0??00-1000-000000--0-011111001011101000111?1010010103001-0010101103101000-------?11?111300000-01-----01111011000110-01110-01401-101--00000010-10-0100030-------0-1 Lepidocaris 1110-00-0--------??0---??0---110-11----10001110???????1?1100???0??00-10?0-0001-0--0-0111110?101110100010????00010001000-00??00110310???????????????0-1-00010-1110??-0-101?11011100-0-110-01001-1???0?0010010-10-0110030----?--0-0 Cephalocarida 1110-00-0--------0-0---------110-11----00000-1000??1?010110011001100-1000-000000--0-01110000001110110011130010010000001-00-0101103100--0-------?11?1-1-00010-11110--000--01?01?000-0-110-01000-000-010010010210-0100030-------0-1 Remipedia 1110-00-0--------0-0---------11??11----10001110001?11?10111????01000-11011000010--0-001010001101101000130---10010000001-00-0101103100--11?000--?11?0-0-00010-10000--101?1011010000-0-110-014101100--00000010?10-0100020--0----0-1 Nahecaris 1110-00-0--------0-11????0-111???100111----1-10???????1?110????0???0-100101001-0--0-011???0??1?1101010?1?????0010?0000??0????0110310???????????????12000-000-01-----1?0--01????????0-110-0100?010????00000?0?1100100000----?--0-? Cinerocaris 1110-00-0--------0-11????0-1111??100110----1-?0???????1?110????0???0-100101001-0--0-0?11110011111010???10---100101??001100-000110310???????????????12000-000-01-----??0--01??1?????0-110-0101?010??0100100?0?110?100000----?--0-? Isopoda 1111-00-0--------0-111130100-111011----0000111001111111011101000??00-1000-000000--0-011111001--11000001-0---10010?0000100010001103100--10---1?01110120100000-10000--101110110000?110-110-01-10------00000010?10-01001-0--1----0-0 Euphausiacea 1111-00-0--------111111200-111111101---00011-1001?????10111010001000-100101001-0--0-011111001--1101000130---10010?00001000100011031011110---111?11?12010-000-10100--110--01101111110-110-010110100-110000010511000011-0----0--0-0 Anaspidacea 1111-00-0--------0-1111200-1111?-11----10001110011???01011101000??00-100101001-0--0-011111001--110101-130---10010100001000100011031011110---111?11?120100000-01-----111?10110?010110-110-01211-010--10000010?11100001-0----0--0-0 Stomatopoda 1111-00-0--------111111200-11111011----0011111101111?010111010001?00-100101001-0--0-011111001--110101-130---10010100001000100011031011111200111111?110100000-10100--1011001100011110-110-010110000-010000010?11100001-0----0--0-0 Leptostraca 1110-00-0--------0-1111100-1111??100111----1-0101111?010111010001100-100100001-0--0-011111001--11000001-0---10010100001000101011031011010---100?11?12000-000-01-----0-0--0110110?110-110-015010100--10000010510-0100030-------0-1 Allacma 1110000-0--------0-111110100-110-11----1000111000?????1011101001100001000-000010--0-011111001--0--------0---0001-00301100000111103100--0-------111?131400000-00----0100--01100000100-0---01-10-0----00000000410-01000-0---0---0-0 Anurida 1110000-0--------0-111110100-110-11----1000111000?????1011101001100001000-000000--0-011111001--0--------0---0001-00301100000111103100--0-------111?131200000-10000-0100--01100000100-0---01-10-0----00000000410-00000-0--00---0-0 Zygentoma 1110000-0--------11111110100-110-11----1000111000111101011101001100001000-100010--0-0111110011-0--------0---0001-000011100001011031010010---1001110131100000-01----0100--01100000100-0---01-10-0----0000000041100100010---0---0-0 Odonata 1110000-0--------11111130100-110-11----10001110011111?1011101001100001000-000010--0-011111001--0--------0---0001-00001110000101103101010-------111?131100000-01----0100--01100000100-0---01-10-0----00000000410-00000-0---0---0-0 Coleoptera 1110100-0--------0-111130100-110-11----1000111001111111011101001100001000-000010--0-011111001--0--------0---0001-00001110000101103101010-------1110131100000-00----1100--01100000100-0---01-10-0----00000000410-00000-0---0---0-0 Diptera 1110100-0--------11111130100-110-11----1000111001111111011101001100011000-0001-0--0-011111001--0--------0---0001-00001110000101103101010-------1110131100000-00----1100--01100000100-0---01-10-0----00000000410-00000-0---0---0-0 Hymenoptera 1110100-0--------11111130100-110-11----1000111001111111011101001100011000-000010--0-0111110011-0--------0---0001-000011100001011031010110---110?11?1-1100000-00----1100--01100000100-0---01-10-0----00000000410-00000-0---0---0-0 Oelandocaris 1110-00-0--------100-????0-0-110-11----10000-00???????1?????11101100-10111?001-0--0-01??0?110??1101?00001?10?0000-----??00-000000310???????????????0-0100000-10100--??0--?1?00?000-0-110-01000100101000?010??1??0?????0----?--0-- cf.Leanchoilia_larva 1110-00-0--------????????????11??11----00000-00?????????110????0????-10?1110?1-0--0-010000110??1101?0????????0000-------0?-000000??0???????????????0-0100000-1010??-?00--?1?0??0???0-11??01010100??1??0??????112?0000-0----?--0-- cf.Leanchoilia_metanauplius 1110-00-0--------????????000-110-11----00000-?0???????1?110????0???0-10?1110?1-0--0-??0000110??1?01?0??01000?0??0???----0?-000000??0???????????????0-0400000-1010??-??0--?1?0??0???0-1????1?00?0010???0??????112?0000-0----?--0-- Yicaris 1110-00-0--------100---------110-11----00000-00???????1????????0???0-101000001-0--0-???000110101101?0??11?001001010?00--00-0000?0?10???????????????0-0300000-01-----100--?1??110???0-110-010110010-11001001?510-?1100?0----?--0-? Rehbachiella 1110-00-0--------100---------110-100110----0-00???????1?11000--01100-10100000000--0-01110110001110110011130010010100001100-010010310???????????????10110-000-01--0--0-0--?11011100-0-110-01001-110-000010010510-0110030----?--0-1 Bredocaris 1110-00-0--------100---------100-10???1----0-00???????1?11000--01000-1010-000000--0-011100110011101100111100100101020-1100-000000310???????????????0-030-000-01-----000--?11011100-0-110-01-001110--00011100?10-0100010----?--0-1 Martinssonia 1110-00-0--------0-0---------11?-11----10000-00???????1?11000--01100-1000-000000--0-0111011100110011001111001001010??-1100-010000310???????????????0-1300000-01-----100--?11011000-0-110-01?00-?10--0000?????10-1100010----?--0-1 Skara 1110-00-0--------0-0---------110-11----10000-00???????1?11000--0??00-1000-000000--0-011101100011001100111?00100101000-1100-010010310???????????????0-1100000-01-----01????11011100-0-110-01000-010-10001010051??????0?0----?--0-1 Phospatocopina 1111-00-0--------100---------110-100100----1-00???????1?11000--011?0-10100000000--0-011101010011101100111?--1001010?001100-010110310???????????????0-??0-000-0?-----010--011001000-0-110-01?00?010-000010010-10-0100010----?--0-1 Agnostus_larva 1111-00-0--------100---------1?0-11----11100-10???????1?11??11101000-1010-0000?0--0-01000?1?0001100-0-00101010100-------00-??0000310???????????????0-0400000-100111-000--?10-0-----1-110-01-10100??-00011000510-?0000-1011-?00011 Peronopsis 1111-00-0--------?00---------120-11----11100-10?????????????1110??0?-1010-000010--0-0100?0110??1100????01010???00-------?0-00000031011010--0111????0-0400000-100111-0?0--????????????11?-01?10?00????00??????10-00000-1011-100001 ; END; BEGIN MRBAYES; SET autoclose = yes nowarn = yes; LOG START filename = Peronopsis_wap18.output; OUTGROUP Priapulida; LSET rates = gamma coding = variable; CONSTRAINT Root = 3 - .; CONSTRAINT Node1 PARTIAL = Coleoptera Diptera : Priapulida Nematoda Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae Chilopoda Diplopoda Pauropoda Symphyla Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma Zygentoma Odonata Hymenoptera; CONSTRAINT Node2 PARTIAL = Coleoptera Diptera Hymenoptera : Priapulida Nematoda Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae Chilopoda Diplopoda Pauropoda Symphyla Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma Zygentoma Odonata; CONSTRAINT Node3 PARTIAL = Odonata Coleoptera Diptera Hymenoptera : Priapulida Nematoda Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae Chilopoda Diplopoda Pauropoda Symphyla Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma Zygentoma; CONSTRAINT Node4 PARTIAL = Zygentoma Odonata Coleoptera Diptera Hymenoptera : Priapulida Nematoda Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae Chilopoda Diplopoda Pauropoda Symphyla Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma; CONSTRAINT Node5 PARTIAL = Anurida Allacma : Priapulida Nematoda Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae Chilopoda Diplopoda Pauropoda Symphyla Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Zygentoma Odonata Coleoptera Diptera Hymenoptera; CONSTRAINT Node6 PARTIAL = Anurida Allacma Zygentoma Odonata Coleoptera Diptera Hymenoptera : Priapulida Nematoda Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae Chilopoda Diplopoda Pauropoda Symphyla Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia; CONSTRAINT Node7 PARTIAL = Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea : Priapulida Nematoda Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae Chilopoda Diplopoda Pauropoda Symphyla Argulus Ostracoda Triops Artemia Cephalocarida Remipedia Anurida Allacma Zygentoma Odonata Coleoptera Diptera Hymenoptera; CONSTRAINT Node8 PARTIAL = Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma Zygentoma Odonata Coleoptera Diptera Hymenoptera : Priapulida Nematoda Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae Chilopoda Diplopoda Pauropoda Symphyla Argulus Ostracoda; CONSTRAINT Node9 PARTIAL = Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma Zygentoma Odonata Coleoptera Diptera Hymenoptera : Priapulida Nematoda Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae Chilopoda Diplopoda Pauropoda Symphyla; CONSTRAINT Node10 PARTIAL = Argulus Ostracoda : Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma Zygentoma Odonata Coleoptera Diptera Hymenoptera Priapulida Nematoda Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae Chilopoda Diplopoda Pauropoda Symphyla; CONSTRAINT Node11 PARTIAL = Diplopoda Pauropoda : Priapulida Nematoda Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae Chilopoda Symphyla Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma Zygentoma Odonata Coleoptera Diptera Hymenoptera; CONSTRAINT Node12 PARTIAL = Diplopoda Pauropoda Symphyla : Priapulida Nematoda Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae Chilopoda Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma Zygentoma Odonata Coleoptera Diptera Hymenoptera; CONSTRAINT Node13 PARTIAL = Chilopoda Diplopoda Pauropoda Symphyla : Priapulida Nematoda Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma Zygentoma Odonata Coleoptera Diptera Hymenoptera; CONSTRAINT Node14 PARTIAL = Chilopoda Diplopoda Pauropoda Symphyla Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma Zygentoma Odonata Coleoptera Diptera Hymenoptera : Priapulida Nematoda Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae; CONSTRAINT Node15 PARTIAL = Archegozetes Opiliones Scorpiones Araneae : Priapulida Nematoda Tardigrada Onychophora Limulus Chilopoda Diplopoda Pauropoda Symphyla Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma Zygentoma Odonata Coleoptera Diptera Hymenoptera; CONSTRAINT Node16 PARTIAL = Limulus Archegozetes Opiliones Scorpiones Araneae : Priapulida Nematoda Tardigrada Onychophora Chilopoda Diplopoda Pauropoda Symphyla Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma Zygentoma Odonata Coleoptera Diptera Hymenoptera; CONSTRAINT Node17 PARTIAL = Limulus Archegozetes Opiliones Scorpiones Araneae Chilopoda Diplopoda Pauropoda Symphyla Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma Zygentoma Odonata Coleoptera Diptera Hymenoptera : Priapulida Nematoda Tardigrada Onychophora; CONSTRAINT Node18 PARTIAL = Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae Chilopoda Diplopoda Pauropoda Symphyla Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma Zygentoma Odonata Coleoptera Diptera Hymenoptera : Priapulida Nematoda; calibrate root = offsetexp ( 540 , 550 ) Hallucigenia = uniform ( 506 , 529 ) Collinsium = uniform ( 515 , 520 ) Luolishania = uniform ( 515 , 520 ) Aysheaia = uniform ( 506 , 508 ) Jianshanopodia = uniform ( 515 , 520 ) Megadictyon = uniform ( 515 , 520 ) Opabinia = uniform ( 506 , 508 ) Anomalocaris = uniform ( 506 , 520 ) Hurdia = uniform ( 506 , 508 ) Peytoia = uniform ( 506 , 508 ) Schinderhannes = uniform ( 390 , 396 ) Surusicaris = uniform ( 506 , 508 ) Isoxys = uniform ( 506 , 520 ) Yohoia = uniform ( 506 , 508 ) Haikoucaris = uniform ( 515 , 520 ) Leanchoiliidae = uniform ( 495 , 520 ) Fortiforceps = uniform ( 515 , 520 ) Kuamaia = uniform ( 515 , 520 ) Marrella = uniform ( 506 , 508 ) Burgessia = uniform ( 506 , 508 ) Emeraldella = uniform ( 506 , 508 ) Sidneyia = uniform ( 506 , 508 ) Aglaspis = uniform ( 485 , 497 ) Cindarella = uniform ( 515 , 520 ) Kodymirus = uniform ( 520 , 530 ) Retifacies = uniform ( 515 , 520 ) Emucaris = uniform ( 510 , 515 ) Tegopelte = uniform ( 506 , 508 ) Cheloniellon = uniform ( 390 , 396 ) Kwanyinaspis = uniform ( 515 , 520 ) Naraoia = uniform ( 420 , 520 ) Arthroaspis = uniform ( 515 , 520 ) Olenoides = uniform ( 506 , 508 ) Triarthrus = uniform ( 445 , 450 ) Hongshiyanaspis = uniform ( 515 , 520 ) Xandarella = uniform ( 515 , 520 ) Sanctacaris = uniform ( 506 , 508 ) Habelia = uniform ( 506 , 508 ) Offacolus = uniform ( 420 , 425 ) Dibasterium = uniform ( 420 , 425 ) Chasmataspis = uniform ( 458 , 470 ) Weinbergina = uniform ( 390 , 396 ) Megalograptus = uniform ( 445 , 460 ) Scorpiones = uniform ( 430 , 438 ) Shankouia = uniform ( 515 , 520 ) Fuxianhuia = uniform ( 515 , 520 ) Chengjiangocaris = uniform ( 515 , 520 ) Euthycarcinoidea = uniform ( 237 , 497 ) Odaraia = uniform ( 506 , 508 ) Canadaspis = uniform ( 506 , 520 ) Branchiocaris = uniform ( 506 , 508 ) Tokummia = uniform ( 506 , 508 ) Waptia = uniform ( 506 , 520 ) Tanazios1 = uniform ( 420 , 425 ) Tanazios2 = uniform ( 420 , 425 ) Cascolus = uniform ( 420 , 425 ) Nereocaris = uniform ( 506 , 508 ) Perspicaris = uniform ( 506 , 508 ) Arthropleura = uniform ( 299 , 315 ) Ostracoda = uniform ( 480 , 485 ) Lepidocaris = uniform ( 400 , 412 ) Nahecaris = uniform ( 390 , 396 ) Cinerocaris = uniform ( 420 , 425 ) Odonata = uniform ( 320 , 325 ) Oelandocaris = uniform ( 490 , 500 ) cf.Leanchoilia_larva = uniform ( 515 , 520 ) cf.Leanchoilia_metanauplius = uniform ( 515 , 520 ) Yicaris = uniform ( 515 , 520 ) Rehbachiella = uniform ( 490 , 500 ) Bredocaris = uniform ( 490 , 500 ) Martinssonia = uniform ( 490 , 500 ) Skara = uniform ( 490 , 500 ) Phospatocopina = uniform ( 490 , 500 ) Agnostus_larva = uniform ( 490 , 500 ) Peronopsis = uniform ( 506 , 508 ); PRSET topologypr = CONSTRAINT ( Root , Node1 , Node2 , Node3 , Node4 , Node5 , Node6 , Node7 , Node8 , Node9 , Node10 , Node11 , Node12 , Node13 , Node14 , Node15 , Node16 , Node17 , Node18 ); prset brlenspr = clock : uniform; prset ratepr = variable; prset clockvarpr = igr; prset igrvarpr = exp ( 10 ); prset nodeagepr = calibrated; prset clockratepr = normal ( 0.001 , 0.01 ); MCMCP temp = 0.1 samplefreq = 1000 printfr = 10000 nruns = 4 nchain = 4 relburnin = yes burninfrac = 0.20; MCMCP savebrlens = yes filename = Peronopsis_wap18; SHOWMOVES; MCMC ngen = 20000000; SUMT filename = Peronopsis_wap18 contype = allcompat relburnin = yes burninfrac = 0.20; SUMP filename = Peronopsis_wap18 relburnin = yes burninfrac = 0.20; END; BEGIN ASSUMPTIONS; TYPESET * UNTITLED = unord: 1- 225; END; BEGIN MESQUITECHARMODELS; ProbModelSet * UNTITLED = 'Mk1 (est.)': 1- 225; END; Begin MESQUITE; MESQUITESCRIPTVERSION 2; TITLE AUTO; tell ProjectCoordinator; timeSaved 1544800575777; getEmployee #mesquite.minimal.ManageTaxa.ManageTaxa; tell It; setID 0 5477453712120517936; tell It; setDefaultOrder 0 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 40 34 44 37 38 42 44 45 46 47 48 49 50 51 52 53 54 55 57 58 59 60 61 62 63 64 65 66 67 68 69 70 71 72 73 74 75 76 77 78 79 80 81 82 83 84 85 86 87 88 89 90 91 92 93 94 95 96 97 98 99 100 103 104 105 106 107 109 108 110; attachments ; endTell; endTell; getEmployee #mesquite.charMatrices.ManageCharacters.ManageCharacters; tell It; setID 0 3270826805688825120; mqVersion 351; checksumv 0 3 1274248034 null getNumChars 225 numChars 225 getNumTaxa 104 numTaxa 104 short true bits 127 states 127 sumSquaresStatesOnly 96293.0 sumSquares 96293.0 longCompressibleToShort false usingShortMatrix true NumFiles 1 NumMatrices 1; mqVersion; endTell; getWindow; tell It; suppress; setResourcesState false false 100; setPopoutState 300; setExplanationSize 0; setAnnotationSize 0; setFontIncAnnot 0; setFontIncExp 0; setSize 3840 1960; setLocation -13 0; setFont SanSerif; setFontSize 10; getToolPalette; tell It; endTell; desuppress; endTell; getEmployee #mesquite.charMatrices.BasicDataWindowCoord.BasicDataWindowCoord; tell It; showDataWindow #3270826805688825120 #mesquite.charMatrices.BasicDataWindowMaker.BasicDataWindowMaker; tell It; getWindow; tell It; setExplanationSize 30; setAnnotationSize 20; setFontIncAnnot 0; setFontIncExp 0; setSize 3740 1888; setLocation -13 0; setFont SanSerif; setFontSize 10; getToolPalette; tell It; setTool mesquite.charMatrices.BasicDataWindowMaker.BasicDataWindow.arrow; endTell; setActive; setTool mesquite.charMatrices.BasicDataWindowMaker.BasicDataWindow.arrow; colorCells #mesquite.charMatrices.NoColor.NoColor; colorRowNames #mesquite.charMatrices.TaxonGroupColor.TaxonGroupColor; colorColumnNames #mesquite.charMatrices.CharGroupColor.CharGroupColor; colorText #mesquite.charMatrices.NoColor.NoColor; setBackground White; toggleShowNames on; toggleShowTaxonNames on; toggleTight off; toggleThinRows off; toggleShowChanges on; toggleSeparateLines off; toggleShowStates on; toggleAutoWCharNames on; toggleAutoTaxonNames off; toggleShowDefaultCharNames off; toggleConstrainCW on; toggleBirdsEye off; toggleShowPaleGrid off; toggleShowPaleCellColors off; toggleShowPaleExcluded off; togglePaleInapplicable on; toggleShowBoldCellText off; toggleAllowAutosize on; toggleColorsPanel off; toggleDiagonal on; setDiagonalHeight 80; toggleLinkedScrolling on; toggleScrollLinkedTables off; endTell; showWindow; getWindow; tell It; forceAutosize; endTell; getEmployee #mesquite.charMatrices.AlterData.AlterData; tell It; toggleBySubmenus off; endTell; getEmployee #mesquite.charMatrices.ColorByState.ColorByState; tell It; setStateLimit 9; toggleUniformMaximum on; endTell; getEmployee #mesquite.charMatrices.ColorCells.ColorCells; tell It; setColor Red; removeColor off; endTell; getEmployee #mesquite.categ.StateNamesStrip.StateNamesStrip; tell It; showStrip off; endTell; getEmployee #mesquite.charMatrices.AnnotPanel.AnnotPanel; tell It; togglePanel off; endTell; getEmployee #mesquite.charMatrices.CharReferenceStrip.CharReferenceStrip; tell It; showStrip off; endTell; getEmployee #mesquite.charMatrices.QuickKeySelector.QuickKeySelector; tell It; autotabOff; endTell; getEmployee #mesquite.charMatrices.SelSummaryStrip.SelSummaryStrip; tell It; showStrip off; endTell; getEmployee #mesquite.categ.SmallStateNamesEditor.SmallStateNamesEditor; tell It; panelOpen true; endTell; endTell; endTell; endTell; end;