#NEXUS
[written Fri Dec 14 10:16:15 EST 2018 by Mesquite  version 3.51 (build 898)]

BEGIN TAXA;
	
	DIMENSIONS NTAX=104;
	TAXLABELS
		Nematoda Priapulida Hallucigenia Collinsium Luolishania Aysheaia Tardigrada Onychophora Jianshanopodia Megadictyon Opabinia Anomalocaris Hurdia Peytoia Schinderhannes Surusicaris Isoxys Yohoia Haikoucaris Leanchoiliidae Fortiforceps Kuamaia Marrella Burgessia Emeraldella Sidneyia Aglaspis Cindarella Kodymirus Retifacies Emucaris Tegopelte Cheloniellon Kwanyinaspis Naraoia Arthroaspis Olenoides Triarthrus Hongshiyanaspis Xandarella Sanctacaris Habelia Offacolus Dibasterium Chasmataspis Weinbergina Megalograptus Limulus Scorpiones Opiliones Araneae Archegozetes Shankouia Fuxianhuia Chengjiangocaris Euthycarcinoidea Odaraia Canadaspis Branchiocaris Tokummia Waptia Tanazios1 Tanazios2 Cascolus Nereocaris Perspicaris Arthropleura Pauropoda Symphyla Chilopoda Diplopoda Ostracoda Argulus Artemia Triops Copepoda Lepidocaris Cephalocarida Remipedia Nahecaris Cinerocaris Isopoda Euphausiacea Anaspidacea Stomatopoda Leptostraca Allacma Anurida Zygentoma Odonata Coleoptera Diptera Hymenoptera Oelandocaris cf.Leanchoilia_larva cf.Leanchoilia_metanauplius Yicaris Rehbachiella Bredocaris Martinssonia Skara Phospatocopina Agnostus_larva Peronopsis 
	;

END;


BEGIN CHARACTERS;
	
	DIMENSIONS  NCHAR=225;
	FORMAT DATATYPE = STANDARD GAP = - MISSING = ? SYMBOLS = "  0 1 2 3 4 5 6";
	MATRIX
	Nematoda                     00-0-----0------0--0---------0-0-0------------------------------------0-----0--------------------------------------------------010000--0-------000-----------------------0-----------------------------------0-------------------
	Priapulida                   00-0-----0------0--0---------0-0-0------------------------------------0-----0--------------------------------------------------010010--0-------000-----------------------0-----------------------------------0-------------------
	Hallucigenia                 10-0-011001100111--0---------0-0-0-------------???????---------------00-----0--------------------------------------------------012100--0-------????----------------------?0---------------------------------40-------------------
	Collinsium                   10-0-011011110111--0---------0-0-0-------------???????---------------00-----0--------------------------------------------------01001???????????????----------------------?0---------------------------------40-------------?-----
	Luolishania                  10-0-01101111?111--0---------0-0-0-------------???????---------------00-----0--------------------------------------------------01??00--0-------????----------------------?0---------------------------------00-------------------
	Aysheaia                     10-0-110010--1011--0---------0-0-0-------------???????---------------00-----0--1--0--------------------------------------------0110?0--0-------????----------------------?0---------------------------------00-------------------
	Tardigrada                   11?0-10000?--1010--0---------0-0-0-------------00000?0---------------00-----0--------------------------------------------------011000--0-------010?----------------------00---------------------------------10-------------------
	Onychophora                  10-0-110010--2001--0---------0-0-0-------------00000?0---------------00-----0--------------------------------------------------001000--0-------1010----------------------00---------------------------------10-------------------
	Jianshanopodia               10-0-1100110-?00?--0---------0-0-0-------------???????---------------00-----0--1--0--------------------------------------------01??0???1101000-????----------------------?0---------------------------------?0-------------------
	Megadictyon                  10-0-1100??????0?--0---------0-0-0-------------???????---------------00-----0--1--0--------------------------------------------01??00--1101000-????----------------------?0---------------------------------?0-------------------
	Opabinia                     1100-10-1--------1111????--100?0-0---------0---???????0?????0--------10?0-000??1?00--------------------------------------------00???0--1111000-????----------------------?0----------101??------------------?0--000--------------
	Anomalocaris                 1100-?0-1-0------0-11???0--10??101-------------???????0?0---0--------10-0-0000010111-------?-----------------------------------002100--1101000-????----------------------?0----------1010?------------------?0--000--------------
	Hurdia                       1100-?0-1-0------0-11????0-10????100101-------????????0?0---0--------10-0-0000011110-------?-----------------------------------00210???????????????----------------------?0----------1010?------------------?????00?-------?-----
	Peytoia                      1100-10-1-0------0-11??????10??????????????---????????0?0---0--------10-0-0000011111-------?-----------------------------------002100--????????????----------------------?0----------10???------------------?0--000--------?-----
	Schinderhannes               110?-1???--------0-11??????100?????????????0-?????????0????????------10?0-00000111??-------------------------------------------00210???????????????----------------------?0----------?????0-----------------?110??00-------?-----
	Surusicaris                  1100-0100--------0-11????0-10100-100110----0-00???????0?0---0--------10-0-0011-11011-------1-----------------000---------------00??0???????????????----------------------?10---------10--00---------?------040????0?-------?-----
	Isoxys                       1100-????--------0-11????0-10??0-100110----0-00???????0?????0--------10?0-001?0?00?1?????????????-???--?????-??0------??0????--00??0???1111000-????----------------------????????????????00---------?---?????0--000--------------
	Yohoia                       1110-00-0--------0-11????0-0-100-11----00000-00???????1????????0??00-10?110011-0--0-????00110?01?01?0???130000000-----??00-0-00?0??00--0-------????0-1100000-101010-?00--?1?00?0???0-111001010-00????0001001?11200000-0-------0--
	Haikoucaris                  1110-00-0--------??11?????-0-10??11----00000-?0????????????????0???0-1??110011-0--0-????001?0???????0???1300???00-----??0?-0-0000??0???????????????0-0100000-1010??-?00--????????????111001010?00????00??????11??0000-0----?--0--
	Leanchoiliidae               1110-00-0--------1011??0?0-0-100-11----000?0-00?1?????1?1???0--0??00-1001110?1-0--0-010000110101101?0???130000000-------00-0-00?0??0??01111000-????0-0100000-10?111-100--?1?00?0???0-1110010101000-1?0001001611200000-0-------0--
	Fortiforceps                 1110-00-0--------0-11?????-1110??11----00000-?????????1????????0???0-1??110001-0--0-????001?0???????0???1300?0?00-----??0?-0-00?0??00--0-------????0-0-00010-1010??-020--????????????11??01000?00????0000000?111?0000-0-------0--
	Kuamaia                      1110-00-0--------0-11????1-0-1?1011----00?00-?0???????1?10--1010????-1000-000010--0-??0000110101001101?4140010?00-------00-0-0000??0???????????????0-0?00000-10010--100--?1????????101110?11101?0110?001????010-?0000-10000?00001
	Marrella                     1110-00-0--------0-0---------100-11----10?00-10???????1?1???1010??00-1000-000010--0-1---0-1101011000010-110000?00-------00-0-00?03101100-------????0-0-00010-01-----000--?1????????0-1110012101010--?0000000010-?0000-0-------0-1
	Burgessia                    1110-00-0--------0-??????0-??10??101-------0-00???????1????????0??00-10?0-000010--0-1---?-111??100100?0??????0?00-------00-0-00?0??011010---111????0-030-000-0?-----100--?1??????????11--01-1010----?001010??110?0000-0----0--0-0
	Emeraldella                  1110-00-0--------0-??????0-??110-11----00?00-10???????1?10--10101?00-1000-100010--0-??00?011?10100??0?0?140010?00-----??00-0-00?03?0???1120?00-????0-0100000-100110-100--?1?00?0???10111001010100110?0011101211010000-0--0----001
	Sidneyia                     1110-00-0--------0-11????0-101?0-11----00000-10???????1?10--1010??00-1000-000010--0-??00?011?10100??0?0?140010100-----??00-0-00?0310???11???00-????0-1100000-100111-100--?110010???10111001010-00110?1011101211110000-0--0----001
	Aglaspis                     1111-00-0--------0-11????100-1?0-11----00100-10???????1?1???1??0???0-1??0-000010--0-????001101010???01??????0??00-----??0?-0-00003?0???????????????0-0100000-100110-100--?1????????1011?-01?10100?????00?????11000100-0--00?--01?
	Cindarella                   1110-00-0--------0-11????0-10110-11----00000-?0???????1?10--11101000-1000-000010--0-1---??110?01001?01?21200?0?00-------00-0-000031011011010100????0-00000011100110-?00--?1????0---??111001010100100??0???0?210-00000-0--0----100
	Kodymirus                    111?-00-0--------0-???????-??100-11----00000-10???????1?10--10101000-1000-0000?0--0-?????-1??1011001010-1?0010100-----??00-0-0000310???????????????0-0100000-100110-100--?10-------1011?-010101001???101?101?110?0000-0--00?--00?
	Retifacies                   1110-00-0--------0-11????0-101?0-11----00000-10???????1?10--1010??00-1000-000010--0-1---?-110?01001?010?1?00?0100-------00-0-0000310???????????????0-0100000-100110-?00--?10-------10111001010100?????0??10??110?0000-1000-?01001
	Emucaris                     1110-00-0--------?-??????0-??100-11----?0?00-?0???????1?10--1110??00-1000-000010--0-1---?-110??10???0??????0???00-------0?-0-0000310???????????????0-0?0000?-1001??-??0--????????????11??01?1????????????????10-?0000-10100?0000?
	Tegopelte                    1110-00-0--------?-??????0-??1-0-11-----0000-?0???????1?10--1110??00-1000-0000?0--0-1---?-110??10?1?0??21200?0100-------00-0-0000310110112?0000????0-00-000?-1001---??0--?1????????10111001110100??0??0??????10-00000---00--0100?
	Cheloniellon                 1110-00-0--------?-1?????100-1?0-11----00001010???????1?1???1110??00-10?0-000010--0-1---?-110001?0?0??0?????00100-------00-0-0000310???????????????0-1100000-10011?-000--?10-------10110-0101010?????0000000610-?1000-0--10?--000
	Kwanyinaspis                 1110-00-0--------?-11????1-0-1???11----00000-?0????????????????0???0-100????0?????????????110?01?0110102120010100-------00-0-000031????????????????0-?10000?-100110-100--?10-------10111001010100????101010??110?0000-0-?10?--00?
	Naraoia                      1110-00-0--------0-11???-0-??110-11----00000-10???????1?10--11101?00-1000-000010--0-1---0-11010100110102120010100-------00-0-000031011011201111????0-000000?-1001---100--?10-------10111001110100100?0010100?10-?0000-1101-10(0 1)0(0 1)?
	Arthroaspis                  1110-00-0--------0-??????0-??10??11----00000-10???????1?10--11101000-1000-000010--0-1---??110?01001101?4140010?00-----??00-0-00?0310110112?0000????0-0000000-100110-100--?10-------1?111001010100110?101?101210-00000-1010--0100?
	Olenoides                    1111-00-0--------0-11???0110-100-11----01100-10???????1?10--10101?00-1000-000010--0-1---0-11010100110102120010100-------00-0-00003101101120?00-????0-0100000-100111-100--?10-------10111001010100100?0011101210-01000-10101-10011
	Triarthrus                   1111-00-0--------0-11???0100-100-11----01100-10???????1?10--11101000-1000-000010--0-1---0-11010100110101110010100-------00-0-0000310???????????????0-0-00010-100111-100--?10-------1011100121010100-00011101210-00000-10101?100(0 1)0
	Hongshiyanaspis              1111-00-0--------0-11???0110-100-11----01100-10???????1?10--1110??00-1000-000010--0-1---0-11010100110104140010100-------00-0-00003101101120000-????0-0000000-100111-100--?10-------10111001010100110?001??0?210-00000-10101-10011
	Xandarella                   1110-00-0--------0-11????100-130-11----0?000-10???????1?10--11101?00-1010-000010--0-1---0-110121001???021200?0?00-------00-0-00003?0???1120100-????0-01000011100110-120--??????0?????11100101010010-?00??????10-00000-0--00---10?
	Sanctacaris                  1110-00-0--------0-11????0-0-130-11----00001010???????1?10--1110????-1011?0001-0--0-1---0-011101000?0??-15?110100-------00-0-000031010?0??????????????110000-1010??-???--?10-----??1?111011010-000-1010?????211210000-0----?--00-
	Habelia                      1110-00-0--------0-11????0-0-130-11----00001010???????1?10--1110????-10?1?0001-0--0-1---0-011101000?0??-150110100-------00-0-000031010?0-------????110110000-101011-100--?10-------10111011010-000-101000000211010000-0-------00-
	Offacolus                    1110-00-0--------??11????????13??11----00?01010???????1?????0--0???--1??0-0101-0--0-1---0-1111010110000-101101?00-------01-0-0000310???0-------????1-1310100-10100--100--?1??????????1110113-0--01--000-0000-11000000-0-------00-
	Dibasterium                  1110-00-0--------????????0???120-11----00001010???????1?????0--0???--1??0-0101-0--0-1---0-1111010110000-101101?00-------01-0-0000310???????????????1-1111100-10100----0--?1??????????1110113-0--01--000-0000-11010000-0----?--00-
	Chasmataspis                 1110-00-0--------1011????100-1??-11----10100-10???????1????????01110-10?0-0101-0--0-1---1-111101?100000-0---00?00-------01-0-00003?????????????????1-11?1100-10000--0-0--?1???00?????????11?-?--????001-0000-11000000-0--10?--01-
	Weinbergina                  1110-00-0--------1011?????0??13?-11----00?00-10???????1????????0???0-10?0-0101-0--0-1---1-1111010100000-0---00100-------00-0-00003?????????????????0-0?0?100-10000--0-0--?1????????10????11?-?--????000-0000211000000-0--0-?--00-
	Megalograptus                1110-00-0--------0-11????100-12?-11----10000-10???????1?1???0--01110-1000-0101-0--0-1---1-1?11011?00000-0---10100-------00-0-0000310???????????????1-111?100-10000--0-0--?10-------1?1111113-0--00--011-1001211?00000-0--10?--00-
	Limulus                      1110-00-0--------10110??0110-130-11----00110-1010000001110--0--0110--1000-0101-0--0-1---1-1111010100000-0---01100-------01-0-00003101100----11101110-0110100-10000--0-0--011001000-101110110-0--011-001-0000-11000000-1111000100-
	Scorpiones                   1110-00-0--------101?0000100-120-11----10000-1010000?01110--0--01110-1000-0101-0--0-1---1-1111011100000-0---00000-------00-0-00003100--1120000-01111-1111100-00-----0-0--110-------111111113-0--00--000-0000411000000-0---0---0--
	Opiliones                    1110-00-0--------?-10?000100-120-11----10000-10100????1110--0--0111--10?0-0101-0--0-1---1-1111011000000-0---00000-------00-0-00003100--11200?0?011?0-1310000-00-----0-0--110-------?10---11---------000-0000610-00000-0---0?--0--
	Araneae                      1110-00-0--------1?100010100-120-11----11000-1010000001110--0--0110--1000-0101-0--0-1---1-1111011000000-0---00000-------00-0-000031010010---11001110-13-0000-00-----0-0--110-------111111113-0--00--000-0000410-00000-0---0---0--
	Archegozetes                 1110-00-0--------0-0-0-------121111----10000-10100????1110--0--011?--1010-0101-0--0-1---1-1111011000000-0---00000-------00-0-00003100--11201110011?0-13-0000-00-----0-0--110-------0-0---11---------000-0000610-00000-0-------0--
	Shankouia                    1110-00-0--------??11????00111?10101-------1-10???????1?11101000??00-1?00-000000--0-0??????????11000000-?????0011102?0??00-0?0010310???0-------?1??1-0-0-010-10010--020--????????????110-010101000-?00000000?110?0000-0-------0-0
	Fuxianhuia                   1110-00-0--------??11??2000111?101000------1-10?????????11101000??00-1100-000000--0-0110?0110020--00000-???0?0011102?0??00-0?0010310???0-------?1??1-1-0-01??1101??-020--????????????110-01010-000-?00000000?110?000000-------0-0
	Chengjiangocaris             1110-00-0--------??11????00111?10101-------1-100????????11101000??00-1?00-000000--0-0110?0110020--00000-?---00011102?0??00-0?0010310???0-------?1??1-?-0-011010010--020--????????????110-010101000-?00000000?1????????0-------0-0
	Euthycarcinoidea             1110-00-0--------0-11????00111?1011----10001110???????1?11??10?0??00-10?0-000000--0-011????????0--------0---?001-?0?????00-0?0??0310???????????????1-1-0?0110110010-020--??????????0-0---01-1-------000000006110?0000-0--1-?--0-0
	Odaraia                      1110-00-0--------0-11????0-111???100110----?-10???????1????????0??00-1????0????0--0-?????????????????????????0???????0??0???????0??0???1???0?0-??????0-0-010-01-----000--?111??0???0-110-01?101??????00?000??1101100000-------0--
	Canadaspis                   1110-00-0--------0-11????0-101?1?100101----???0???????1????????0??00-11?0-000000--0-???????1???????00?0??????????????0??00???0??0??0???????????????10110-000-01-----00????1111?0???0-111001010-0?????0011001210-1110000----?--0-0
	Branchiocaris                1110-00-0--------0-??????0-?01110100100----1-10???????1?10--0--01?00-10?0-000000--0-011???00???0--------?????001-?02?0??0????00?0310???0-------????1?0-0-010-01-----101-1?1111?0???0-110-0101010?????000?00??10-010000???-----0-0
	Tokummia                     1110-00-0--------0-??????0-?01111100100----1-10???????1?10--0--01?00-10?0-000000--0-011???001??0--------?????001-?0200??00???00?0310???0-------????1?0-0-010-01-----101-1?111110???0-110-0101010?????0001001410-010001???-----0-0
	Waptia                       1110-00-0--------0-11????0-111?11100111----1-10???????1?10--0--0???0-11?0-000010--0-011111010110--------0---0001-102001100-0100?0310???0-------????1-111-001101-----0-0--?11?1?????0-0-1011--0-010--?00-0010410-0100020-------0-0
	Tanazios1                    1110-00-0--------0-0---------120-11----00001110?????????1100???0???0-1000-0001-0--0-?01000110??110000?0-1??0?0?1010?00--00-000110310???????????????0-0-00010-1011??-0?0--?110??10??0-110-0150100???-100??????10-?1100?0----?--0-?
	Tanazios2                    1110-00-0--------0-0---------120-11----00001110?????????1100???0???0-1100-000010--0-?11000110??0--------1??0?0?1-10?00--00-000110310???????????????0-0-00010-1011??-0?0--?110??10??0-110-0150100???-100??????10-?1100?0----?--0-?
	Cascolus                     1110-00-0--------0-11????0-11110-11----11000010???????1?1100???01000-100101001-0--0-011000010??11?100?0013000001010300--00-0000?0310???????????????0-?100000-00-----0?0--?1100?10??0-110-01000?0???11000?????1????????0----?--0-0
	Nereocaris                   1110-00-0--------0-11????0-111???100110----?-?0????????????????0???0-1????0????0--0-????????????????????????????????????????????0310?????????????????1-?-01??01-----0????????????????????01?0?-???????00?????10-0100000----?--0-?
	Perspicaris                  1110-00-0--------0-11????0-111?0-100101----?-10????????????????0???0-10?0-000000--0-????????????????????????????????????????????0310?????????????????10?-00??01-----?????????????????11???10??-???????0??????10-0100000----?--0-?
	Arthropleura                 1110-00-0--------0-0---------11??11----1000111????????1?????????????-1??0-000010--0-0111?10?1???????????????????0???????????????0310???????????????0-0-000111100010-020--?11?00001?0-0---01-1-1-----?0001000610-?0000-0--0-?--0-?
	Pauropoda                    1110-00-0--------0-0---------110-11----1000111001?????10111110001001-100111001-0--0-011111001--0--------0---0001-000010-10-1001103100--0-------111?0-0200001100-----100--01100000100-0---01-1-1-----00000000610-00000-0-------0-0
	Symphyla                     1111-00-0--------0-0---------110-11----1000111001?????10111110011001-1000-000010--0-011111001--0--------0---0001-01001100000001103100--0-------111?0-0100001200-----100--01100000100-0---01-1-1-----00000000610-0100030-------0-0
	Chilopoda                    1110-00-0--------0-1?0011100-110-11----10001110010100010111010001001-1000-000010--0-0111110011-0--------0---0001-01?111000-0101103100--0-------11110-0-00010-00-----1010101100000100-0---01-1-1-----00000000611100000-0---0---0-0
	Diplopoda                    1111-00-0--------0-120011100-110-11----10001110010??0010111110001001-1000-000000--0-011111001--0--------0---0001-010110-10-1001103100--0-------11110-0-00011100-----100--01100000100-0---01-1-1-----00000000610-00000-0---0---0-0
	Ostracoda                    1111-00-0--------1-0---------110-100100----1-100??????101100???01100-1000-000000--0-0111110011111010001013--0001010?00?-0010001103101000-------?11?0-110-000-01-----110--011001000-0-110-01000-000-000000010?10-0100010-------0-1
	Argulus                      1110-00-0--------0-11?1?0100-11??11-----1000-1001?????1011000--?1?0--100??0001-0--0-0111110011111000001-0---1001000?000-00-01?01031010010---110?11?1-1400000-00-----000--011000100-0-110-01-00-100--00000010-10-0?0?0?0---0---0-0
	Artemia                      1110-00-0--------111111100-11110-11----1000110000110001011000--01000-0?0--000--0--0-011111001--110?0001-0---10010001000-0011001103100--10---111111?101-00010-01-----0-0--011011000-0-110-01501-000--10000010-10-0100030----0--0-0
	Triops                       1111-00-0--------12111110100-110-1001110---1-10001100010111110001000-000--000--0--0-011111001--11000001-0---10010002000-0011001103100--10---1111110101-0-011101-----0-11?011011000-0-110-01501-000--10000010-10-0100010---00--0-0
	Copepoda                     1110-00-0--------0-11--------11??11----10000-1001?????101101???0??00-1000-000000--0-011111001011101000111?1010010103001-0010101103101000-------?11?111300000-01-----01111011000110-01110-01401-101--00000010-10-0100030-------0-1
	Lepidocaris                  1110-00-0--------??0---??0---110-11----10001110???????1?1100???0??00-10?0-0001-0--0-0111110?101110100010????00010001000-00??00110310???????????????0-1-00010-1110??-0-101?11011100-0-110-01001-1???0?0010010-10-0110030----?--0-0
	Cephalocarida                1110-00-0--------0-0---------110-11----00000-1000??1?010110011001100-1000-000000--0-01110000001110110011130010010000001-00-0101103100--0-------?11?1-1-00010-11110--000--01?01?000-0-110-01000-000-010010010210-0100030-------0-1
	Remipedia                    1110-00-0--------0-0---------11??11----10001110001?11?10111????01000-11011000010--0-001010001101101000130---10010000001-00-0101103100--11?000--?11?0-0-00010-10000--101?1011010000-0-110-014101100--00000010?10-0100020--0----0-1
	Nahecaris                    1110-00-0--------0-11????0-111???100111----1-10???????1?110????0???0-100101001-0--0-011???0??1?1101010?1?????0010?0000??0????0110310???????????????12000-000-01-----1?0--01????????0-110-0100?010????00000?0?1100100000----?--0-?
	Cinerocaris                  1110-00-0--------0-11????0-1111??100110----1-?0???????1?110????0???0-100101001-0--0-0?11110011111010???10---100101??001100-000110310???????????????12000-000-01-----??0--01??1?????0-110-0101?010??0100100?0?110?100000----?--0-?
	Isopoda                      1111-00-0--------0-111130100-111011----0000111001111111011101000??00-1000-000000--0-011111001--11000001-0---10010?0000100010001103100--10---1?01110120100000-10000--101110110000?110-110-01-10------00000010?10-01001-0--1----0-0
	Euphausiacea                 1111-00-0--------111111200-111111101---00011-1001?????10111010001000-100101001-0--0-011111001--1101000130---10010?00001000100011031011110---111?11?12010-000-10100--110--01101111110-110-010110100-110000010511000011-0----0--0-0
	Anaspidacea                  1111-00-0--------0-1111200-1111?-11----10001110011???01011101000??00-100101001-0--0-011111001--110101-130---10010100001000100011031011110---111?11?120100000-01-----111?10110?010110-110-01211-010--10000010?11100001-0----0--0-0
	Stomatopoda                  1111-00-0--------111111200-11111011----0011111101111?010111010001?00-100101001-0--0-011111001--110101-130---10010100001000100011031011111200111111?110100000-10100--1011001100011110-110-010110000-010000010?11100001-0----0--0-0
	Leptostraca                  1110-00-0--------0-1111100-1111??100111----1-0101111?010111010001100-100100001-0--0-011111001--11000001-0---10010100001000101011031011010---100?11?12000-000-01-----0-0--0110110?110-110-015010100--10000010510-0100030-------0-1
	Allacma                      1110000-0--------0-111110100-110-11----1000111000?????1011101001100001000-000010--0-011111001--0--------0---0001-00301100000111103100--0-------111?131400000-00----0100--01100000100-0---01-10-0----00000000410-01000-0---0---0-0
	Anurida                      1110000-0--------0-111110100-110-11----1000111000?????1011101001100001000-000000--0-011111001--0--------0---0001-00301100000111103100--0-------111?131200000-10000-0100--01100000100-0---01-10-0----00000000410-00000-0--00---0-0
	Zygentoma                    1110000-0--------11111110100-110-11----1000111000111101011101001100001000-100010--0-0111110011-0--------0---0001-000011100001011031010010---1001110131100000-01----0100--01100000100-0---01-10-0----0000000041100100010---0---0-0
	Odonata                      1110000-0--------11111130100-110-11----10001110011111?1011101001100001000-000010--0-011111001--0--------0---0001-00001110000101103101010-------111?131100000-01----0100--01100000100-0---01-10-0----00000000410-00000-0---0---0-0
	Coleoptera                   1110100-0--------0-111130100-110-11----1000111001111111011101001100001000-000010--0-011111001--0--------0---0001-00001110000101103101010-------1110131100000-00----1100--01100000100-0---01-10-0----00000000410-00000-0---0---0-0
	Diptera                      1110100-0--------11111130100-110-11----1000111001111111011101001100011000-0001-0--0-011111001--0--------0---0001-00001110000101103101010-------1110131100000-00----1100--01100000100-0---01-10-0----00000000410-00000-0---0---0-0
	Hymenoptera                  1110100-0--------11111130100-110-11----1000111001111111011101001100011000-000010--0-0111110011-0--------0---0001-000011100001011031010110---110?11?1-1100000-00----1100--01100000100-0---01-10-0----00000000410-00000-0---0---0-0
	Oelandocaris                 1110-00-0--------100-????0-0-110-11----10000-00???????1?????11101100-10111?001-0--0-01??0?110??1101?00001?10?0000-----??00-000000310???????????????0-0100000-10100--??0--?1?00?000-0-110-01000100101000?010??1??0?????0----?--0--
	cf.Leanchoilia_larva         1110-00-0--------????????????11??11----00000-00?????????110????0????-10?1110?1-0--0-010000110??1101?0????????0000-------0?-000000??0???????????????0-0100000-1010??-?00--?1?0??0???0-11??01010100??1??0??????112?0000-0----?--0--
	cf.Leanchoilia_metanauplius  1110-00-0--------????????000-110-11----00000-?0???????1?110????0???0-10?1110?1-0--0-??0000110??1?01?0??01000?0??0???----0?-000000??0???????????????0-0400000-1010??-??0--?1?0??0???0-1????1?00?0010???0??????112?0000-0----?--0--
	Yicaris                      1110-00-0--------100---------110-11----00000-00???????1????????0???0-101000001-0--0-???000110101101?0??11?001001010?00--00-0000?0?10???????????????0-0300000-01-----100--?1??110???0-110-010110010-11001001?510-?1100?0----?--0-?
	Rehbachiella                 1110-00-0--------100---------110-100110----0-00???????1?11000--01100-10100000000--0-01110110001110110011130010010100001100-010010310???????????????10110-000-01--0--0-0--?11011100-0-110-01001-110-000010010510-0110030----?--0-1
	Bredocaris                   1110-00-0--------100---------100-10???1----0-00???????1?11000--01000-1010-000000--0-011100110011101100111100100101020-1100-000000310???????????????0-030-000-01-----000--?11011100-0-110-01-001110--00011100?10-0100010----?--0-1
	Martinssonia                 1110-00-0--------0-0---------11?-11----10000-00???????1?11000--01100-1000-000000--0-0111011100110011001111001001010??-1100-010000310???????????????0-1300000-01-----100--?11011000-0-110-01?00-?10--0000?????10-1100010----?--0-1
	Skara                        1110-00-0--------0-0---------110-11----10000-00???????1?11000--0??00-1000-000000--0-011101100011001100111?00100101000-1100-010010310???????????????0-1100000-01-----01????11011100-0-110-01000-010-10001010051??????0?0----?--0-1
	Phospatocopina               1111-00-0--------100---------110-100100----1-00???????1?11000--011?0-10100000000--0-011101010011101100111?--1001010?001100-010110310???????????????0-??0-000-0?-----010--011001000-0-110-01?00?010-000010010-10-0100010----?--0-1
	Agnostus_larva               1111-00-0--------100---------1?0-11----11100-10???????1?11??11101000-1010-0000?0--0-01000?1?0001100-0-00101010100-------00-??0000310???????????????0-0400000-100111-000--?10-0-----1-110-01-10100??-00011000510-?0000-1011-?00011
	Peronopsis                   1111-00-0--------?00---------120-11----11100-10?????????????1110??0?-1010-000010--0-0100?0110??1100????01010???00-------?0-00000031011010--0111????0-0400000-100111-0?0--????????????11?-01?10?00????00??????10-00000-1011-100001

;

END;
BEGIN MRBAYES;
SET autoclose = yes nowarn = yes;
LOG START filename = Peronopsis_wap18.output;
OUTGROUP Priapulida;
LSET rates = gamma coding = variable;
CONSTRAINT Root = 3 - .;
CONSTRAINT Node1 PARTIAL =   Coleoptera Diptera : Priapulida Nematoda Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae Chilopoda Diplopoda Pauropoda Symphyla Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma Zygentoma Odonata Hymenoptera;
CONSTRAINT Node2 PARTIAL =   Coleoptera Diptera Hymenoptera : Priapulida Nematoda Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae Chilopoda Diplopoda Pauropoda Symphyla Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma Zygentoma Odonata;
CONSTRAINT Node3 PARTIAL =   Odonata Coleoptera Diptera Hymenoptera : Priapulida Nematoda Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae Chilopoda Diplopoda Pauropoda Symphyla Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma Zygentoma;
CONSTRAINT Node4 PARTIAL =   Zygentoma Odonata Coleoptera Diptera Hymenoptera : Priapulida Nematoda Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae Chilopoda Diplopoda Pauropoda Symphyla Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma;
CONSTRAINT Node5 PARTIAL =   Anurida Allacma : Priapulida Nematoda Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae Chilopoda Diplopoda Pauropoda Symphyla Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Zygentoma Odonata Coleoptera Diptera Hymenoptera;
CONSTRAINT Node6 PARTIAL =   Anurida Allacma Zygentoma Odonata Coleoptera Diptera Hymenoptera : Priapulida Nematoda Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae Chilopoda Diplopoda Pauropoda Symphyla Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia;
CONSTRAINT Node7 PARTIAL =   Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea : Priapulida Nematoda Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae Chilopoda Diplopoda Pauropoda Symphyla Argulus Ostracoda Triops Artemia Cephalocarida Remipedia Anurida Allacma Zygentoma Odonata Coleoptera Diptera Hymenoptera;
CONSTRAINT Node8 PARTIAL =   Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma Zygentoma Odonata Coleoptera Diptera Hymenoptera : Priapulida Nematoda Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae Chilopoda Diplopoda Pauropoda Symphyla Argulus Ostracoda;
CONSTRAINT Node9 PARTIAL =   Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma Zygentoma Odonata Coleoptera Diptera Hymenoptera : Priapulida Nematoda Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae Chilopoda Diplopoda Pauropoda Symphyla;
CONSTRAINT Node10 PARTIAL =  Argulus Ostracoda : Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma Zygentoma Odonata Coleoptera Diptera Hymenoptera Priapulida Nematoda Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae Chilopoda Diplopoda Pauropoda Symphyla;
CONSTRAINT Node11 PARTIAL =  Diplopoda Pauropoda : Priapulida Nematoda Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae Chilopoda Symphyla Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma Zygentoma Odonata Coleoptera Diptera Hymenoptera;
CONSTRAINT Node12 PARTIAL =  Diplopoda Pauropoda Symphyla : Priapulida Nematoda Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae Chilopoda Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma Zygentoma Odonata Coleoptera Diptera Hymenoptera;
CONSTRAINT Node13 PARTIAL =  Chilopoda Diplopoda Pauropoda Symphyla : Priapulida Nematoda Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma Zygentoma Odonata Coleoptera Diptera Hymenoptera;
CONSTRAINT Node14 PARTIAL =  Chilopoda Diplopoda Pauropoda Symphyla Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma Zygentoma Odonata Coleoptera Diptera Hymenoptera : Priapulida Nematoda Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae;
CONSTRAINT Node15 PARTIAL =  Archegozetes Opiliones Scorpiones Araneae : Priapulida Nematoda Tardigrada Onychophora Limulus Chilopoda Diplopoda Pauropoda Symphyla Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma Zygentoma Odonata Coleoptera Diptera Hymenoptera;
CONSTRAINT Node16 PARTIAL =  Limulus Archegozetes Opiliones Scorpiones Araneae : Priapulida Nematoda Tardigrada Onychophora Chilopoda Diplopoda Pauropoda Symphyla Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma Zygentoma Odonata Coleoptera Diptera Hymenoptera;
CONSTRAINT Node17 PARTIAL =  Limulus Archegozetes Opiliones Scorpiones Araneae Chilopoda Diplopoda Pauropoda Symphyla Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma Zygentoma Odonata Coleoptera Diptera Hymenoptera : Priapulida Nematoda Tardigrada Onychophora;
CONSTRAINT Node18 PARTIAL =  Tardigrada Onychophora Limulus Archegozetes Opiliones Scorpiones Araneae Chilopoda Diplopoda Pauropoda Symphyla Argulus Ostracoda Triops Artemia Copepoda Leptostraca Stomatopoda Isopoda Euphausiacea Cephalocarida Remipedia Anurida Allacma Zygentoma Odonata Coleoptera Diptera Hymenoptera : Priapulida Nematoda;
calibrate
 root = offsetexp ( 540 , 550 )
 Hallucigenia = uniform ( 506 , 529 )
 Collinsium = uniform ( 515 , 520 )
 Luolishania = uniform ( 515 , 520 )
 Aysheaia = uniform ( 506 , 508 )
 Jianshanopodia = uniform ( 515 , 520 )
 Megadictyon = uniform ( 515 , 520 )
 Opabinia = uniform ( 506 , 508 )
 Anomalocaris = uniform ( 506 , 520 )
 Hurdia = uniform ( 506 , 508 )
 Peytoia = uniform ( 506 , 508 )
 Schinderhannes = uniform ( 390 , 396 )
 Surusicaris = uniform ( 506 , 508 )
 Isoxys = uniform ( 506 , 520 )
 Yohoia = uniform ( 506 , 508 )
 Haikoucaris = uniform ( 515 , 520 )
 Leanchoiliidae = uniform ( 495 , 520 )
 Fortiforceps = uniform ( 515 , 520 )
 Kuamaia = uniform ( 515 , 520 )
 Marrella = uniform ( 506 , 508 )
 Burgessia = uniform ( 506 , 508 )
 Emeraldella = uniform ( 506 , 508 )
 Sidneyia = uniform ( 506 , 508 )
 Aglaspis = uniform ( 485 , 497 )
 Cindarella = uniform ( 515 , 520 )
 Kodymirus = uniform ( 520 , 530 )
 Retifacies = uniform ( 515 , 520 )
 Emucaris = uniform ( 510 , 515 )
 Tegopelte = uniform ( 506 , 508 )
 Cheloniellon = uniform ( 390 , 396 )
 Kwanyinaspis = uniform ( 515 , 520 )
 Naraoia = uniform ( 420 , 520 )
 Arthroaspis = uniform ( 515 , 520 )
 Olenoides = uniform ( 506 , 508 )
 Triarthrus = uniform ( 445 , 450 )
 Hongshiyanaspis = uniform ( 515 , 520 )
 Xandarella = uniform ( 515 , 520 )
 Sanctacaris = uniform ( 506 , 508 )
 Habelia = uniform ( 506 , 508 )
 Offacolus = uniform ( 420 , 425 )
 Dibasterium = uniform ( 420 , 425 )
 Chasmataspis = uniform ( 458 , 470 )
 Weinbergina = uniform ( 390 , 396 )
 Megalograptus = uniform ( 445 , 460 )
 Scorpiones = uniform ( 430 , 438 )
 Shankouia = uniform ( 515 , 520 )
 Fuxianhuia = uniform ( 515 , 520 )
 Chengjiangocaris = uniform ( 515 , 520 )
 Euthycarcinoidea = uniform ( 237 , 497 )
 Odaraia = uniform ( 506 , 508 )
 Canadaspis = uniform ( 506 , 520 )
 Branchiocaris = uniform ( 506 , 508 )
 Tokummia = uniform ( 506 , 508 )
 Waptia = uniform ( 506 , 520 )
 Tanazios1 = uniform ( 420 , 425 )
 Tanazios2 = uniform ( 420 , 425 )
 Cascolus = uniform ( 420 , 425 )
 Nereocaris = uniform ( 506 , 508 )
 Perspicaris = uniform ( 506 , 508 )
 Arthropleura = uniform ( 299 , 315 )
 Ostracoda = uniform ( 480 , 485 )
 Lepidocaris = uniform ( 400 , 412 )
 Nahecaris = uniform ( 390 , 396 )
 Cinerocaris = uniform ( 420 , 425 )
 Odonata = uniform ( 320 , 325 )
 Oelandocaris = uniform ( 490 , 500 )
 cf.Leanchoilia_larva = uniform ( 515 , 520 )
 cf.Leanchoilia_metanauplius = uniform ( 515 , 520 )
 Yicaris = uniform ( 515 , 520 )
 Rehbachiella = uniform ( 490 , 500 )
 Bredocaris = uniform ( 490 , 500 )
 Martinssonia = uniform ( 490 , 500 )
 Skara = uniform ( 490 , 500 )
 Phospatocopina = uniform ( 490 , 500 )
 Agnostus_larva = uniform ( 490 , 500 )
 Peronopsis = uniform ( 506 , 508 );
PRSET topologypr = CONSTRAINT (
 Root ,
 Node1 ,
 Node2 ,
 Node3 ,
 Node4 ,
 Node5 ,
 Node6 ,
 Node7 ,
 Node8 ,
 Node9 ,
 Node10 ,
 Node11 ,
 Node12 ,
 Node13 ,
 Node14 ,
 Node15 ,
 Node16 ,
 Node17 ,
 Node18
 );
prset brlenspr = clock : uniform;
prset ratepr = variable;
prset clockvarpr = igr;
prset igrvarpr = exp ( 10 );
prset nodeagepr = calibrated;
prset clockratepr = normal ( 0.001 , 0.01 );
MCMCP temp = 0.1 samplefreq = 1000 printfr = 10000 nruns = 4 nchain = 4 relburnin = yes burninfrac = 0.20;
MCMCP savebrlens = yes filename = Peronopsis_wap18;
SHOWMOVES;
MCMC ngen = 20000000;
SUMT filename = Peronopsis_wap18 contype = allcompat relburnin = yes burninfrac = 0.20;
SUMP filename = Peronopsis_wap18 relburnin = yes burninfrac = 0.20;

END;
BEGIN ASSUMPTIONS;
	TYPESET * UNTITLED   =  unord:  1- 225;

END;

BEGIN MESQUITECHARMODELS;
	ProbModelSet * UNTITLED   =  'Mk1 (est.)':  1- 225;
END;

Begin MESQUITE;
		MESQUITESCRIPTVERSION 2;
		TITLE AUTO;
		tell ProjectCoordinator;
		timeSaved 1544800575777;
		getEmployee #mesquite.minimal.ManageTaxa.ManageTaxa;
		tell It;
			setID 0 5477453712120517936;
			tell It;
				setDefaultOrder  0 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 40 34 44 37 38 42 44 45 46 47 48 49 50 51 52 53 54 55 57 58 59 60 61 62 63 64 65 66 67 68 69 70 71 72 73 74 75 76 77 78 79 80 81 82 83 84 85 86 87 88 89 90 91 92 93 94 95 96 97 98 99 100 103 104 105 106 107 109 108 110;
				attachments ;
			endTell;
		endTell;
		getEmployee #mesquite.charMatrices.ManageCharacters.ManageCharacters;
		tell It;
			setID 0 3270826805688825120;
			mqVersion 351;
			checksumv 0 3 1274248034 null  getNumChars 225 numChars 225 getNumTaxa 104 numTaxa 104   short true   bits 127   states 127   sumSquaresStatesOnly 96293.0 sumSquares 96293.0 longCompressibleToShort false usingShortMatrix true   NumFiles 1 NumMatrices 1;
			mqVersion;
		endTell;
		getWindow;
		tell It;
			suppress;
			setResourcesState false false 100;
			setPopoutState 300;
			setExplanationSize 0;
			setAnnotationSize 0;
			setFontIncAnnot 0;
			setFontIncExp 0;
			setSize 3840 1960;
			setLocation -13 0;
			setFont SanSerif;
			setFontSize 10;
			getToolPalette;
			tell It;
			endTell;
			desuppress;
		endTell;
		getEmployee  #mesquite.charMatrices.BasicDataWindowCoord.BasicDataWindowCoord;
		tell It;
			showDataWindow #3270826805688825120 #mesquite.charMatrices.BasicDataWindowMaker.BasicDataWindowMaker;
			tell It;
				getWindow;
				tell It;
					setExplanationSize 30;
					setAnnotationSize 20;
					setFontIncAnnot 0;
					setFontIncExp 0;
					setSize 3740 1888;
					setLocation -13 0;
					setFont SanSerif;
					setFontSize 10;
					getToolPalette;
					tell It;
						setTool mesquite.charMatrices.BasicDataWindowMaker.BasicDataWindow.arrow;
					endTell;
					setActive;
					setTool mesquite.charMatrices.BasicDataWindowMaker.BasicDataWindow.arrow;
					colorCells  #mesquite.charMatrices.NoColor.NoColor;
					colorRowNames  #mesquite.charMatrices.TaxonGroupColor.TaxonGroupColor;
					colorColumnNames  #mesquite.charMatrices.CharGroupColor.CharGroupColor;
					colorText  #mesquite.charMatrices.NoColor.NoColor;
					setBackground White;
					toggleShowNames on;
					toggleShowTaxonNames on;
					toggleTight off;
					toggleThinRows off;
					toggleShowChanges on;
					toggleSeparateLines off;
					toggleShowStates on;
					toggleAutoWCharNames on;
					toggleAutoTaxonNames off;
					toggleShowDefaultCharNames off;
					toggleConstrainCW on;
					toggleBirdsEye off;
					toggleShowPaleGrid off;
					toggleShowPaleCellColors off;
					toggleShowPaleExcluded off;
					togglePaleInapplicable on;
					toggleShowBoldCellText off;
					toggleAllowAutosize on;
					toggleColorsPanel off;
					toggleDiagonal on;
					setDiagonalHeight 80;
					toggleLinkedScrolling on;
					toggleScrollLinkedTables off;
				endTell;
				showWindow;
				getWindow;
				tell It;
					forceAutosize;
				endTell;
				getEmployee #mesquite.charMatrices.AlterData.AlterData;
				tell It;
					toggleBySubmenus off;
				endTell;
				getEmployee #mesquite.charMatrices.ColorByState.ColorByState;
				tell It;
					setStateLimit 9;
					toggleUniformMaximum on;
				endTell;
				getEmployee #mesquite.charMatrices.ColorCells.ColorCells;
				tell It;
					setColor Red;
					removeColor off;
				endTell;
				getEmployee #mesquite.categ.StateNamesStrip.StateNamesStrip;
				tell It;
					showStrip off;
				endTell;
				getEmployee #mesquite.charMatrices.AnnotPanel.AnnotPanel;
				tell It;
					togglePanel off;
				endTell;
				getEmployee #mesquite.charMatrices.CharReferenceStrip.CharReferenceStrip;
				tell It;
					showStrip off;
				endTell;
				getEmployee #mesquite.charMatrices.QuickKeySelector.QuickKeySelector;
				tell It;
					autotabOff;
				endTell;
				getEmployee #mesquite.charMatrices.SelSummaryStrip.SelSummaryStrip;
				tell It;
					showStrip off;
				endTell;
				getEmployee #mesquite.categ.SmallStateNamesEditor.SmallStateNamesEditor;
				tell It;
					panelOpen true;
				endTell;
			endTell;
		endTell;
		endTell;
end;


