AnimaliaNymphalidaeNymphalidaeNakaharaShinichiLamasGerardoTylerStephanieMarínMario AlejandroHuertasBlancaWillmottKeith R.MielkeOlaf H. H.EspelandMarianneA revision of the new genus Amiga Nakahara, Willmott & Espeland, gen. n., described for Papilioarnaca Fabricius, 1776 (Lepidoptera, Nymphalidae, Satyrinae)Zookeys311201920198218515210.3897/zookeys.821.31782 Amiga arnaca adela http://zoobank.org/737B03DC-05CC-4D17-B734-13E3E3B38918 Nakahara & Espelandsubsp. n.Figs 2i–l, 3d, 6Description and diagnosis.

MALE: forewing length 19–21 mm (n = 5): Differs from the nominotypical subspecies in the following respects: broad, faint, iridescent bluish-lilac reflection between the ventral forewing postdiscal band and the umbra is absent (but see also below).

FEMALE: forewing length 21–22 mm (n = 4): Differs from the nominotypical subspecies by the same ventral forewing character provided for the male.

Variation.

As explained under the variation section under the genus, the absence of the bluish-lilac reflection on the ventral forewing is consistent in specimens from western Colombia and western Ecuador. However, this character appears in a few specimens from Panama, Costa Rica, and Nicaragua (e.g., FLMNH-MGCL 208036, 257145, 263067), although the degree of reflection is variable.

Types.

HOLOTYPE ♂: // ECUADOR: Esmeraldas, W Maldonado–Selva Alegre rd., El Cerro 235 m, 0°58'22"N,78°55'19"W 21, 25.vii.2011, K. Willmott, & J. Hall FLMNH-MGCL-151127 // DNA voucher LEP-09931 // (FLMNH, to be deposited in INABIO). (Fig. 2k).

Other examined specimens

(244 ♂, 104 ♀). See Appendix for the data of these specimens. These specimens are not included in the type series as labelling will likely not be completed for over 300 specimens.

Etymology.

This species-group name is based on the Greek adjective “adelos”, meaning “indistinct” or “inconspicuous”, in reference to its lack of bluish reflection on the ventral forewing. This species-group name is treated as a Latinized feminine adjective in accordance with the feminine generic name.

Distribution

(Fig. 6). This subspecies occurs from Nicaragua to western Ecuador.

Amigaarnaca spp. specimens spanning its range (dorsal on left, ventral on right): a nominotypical subspecies from Suriname, neotype male (USNM) b nominotypical subspecies from Guyana, female (FLMNH-MGCL 263373) c nominotypical subspecies from E Ecuador, male (FLMNH-MGCL 257121) d nominotypical subspecies from Peru, female (FLMNH-MGCL 262953); e nominotypical subspecies from N Brazil, male (FLMNH-MGCL1036223) f Nominotypical subspecies from N Brazil, female (FLMNH-MGCL 207984) g Nominotypical subspecies from SE Brazil, male (FLMNH-MGCL 1036213); h nominotypical subspecies from SE Brazil, female (FLMNH-MGCL 1036218) iA.arnacaadela from Costa Rica, male (FLMNH-MGCL 207991) jA.arnacaadela from Costa Rica, female (FLMNH-MGCL 207992) kA.arnacaadela from W Ecuador, holotype male (FLMNH-MGCL 151127) lA.arnacaadela from W Ecuador, female (FLMNH-MGCL 257087) mA.arnacasericeella, male from Mexico (FLMNH-MGCL 207900) nA.arnacasericeella from Mexico, female (FLMNH-MGCL 207896) oA.arnacaindianacristoi from NW Venezuela, paratype male (FLMNH-MGCL 263107) pA.arnacaindianacristoi from N Venezuela, paratype female (FLMNH-MGCL 1036235).

Amigaarnacaarnaca comb. n. wing venation: a male (FLMNH-MGCL specimen 257164) b female (FLMNH-MGCL specimen 257166).

Distribution map of Amigaarnaca ssp.