AnimaliaColeopteraHydrophilidaeGirónJennifer C.ShortAndrew Edward Z.Review of the Neotropical water scavenger beetle genus Tobochares Short & García, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new recordsZookeys222202110199314010.3897/zookeys.1019.59881C005666E-376B-50FC-8C4C-154284A560E4 Tobochares sulcatus species groupDiagnosis.

This species group can be recognized by the oblique and posteriorly directed anterior margin of the eye in lateral view, which emarginates the eye in lateral view (Fig. 2A–C), the broad, ovoid to diamond-shaped glabrous patch on the metaventrite (Kohlenberg and Short 2017: fig. 10), the dense and uniform distribution of the hydrofuge pubescence on the abdominal ventrites (Kohlenberg and Short 2017: fig. 13A), and the moderate (in number and size) metatibial spines.

Composition.

This species group includes the following species: Tobochares benettii sp. nov., T. canaliculatus Kohlenberg & Short, 2017, T. goias sp. nov., T. kasikasima Short, 2013, T. kusad Kohlenberg & Short, 2017, T. pallidus Kohlenberg & Short, 2017, T. sipaliwini Short & Kadosoe, 2011, T. striatus Short, 2013, and T. sulcatus Short & García, 2007; see Table 2.

10.3897/zookeys.1019.59881.figure280986B23-97A3-5C94-BA74-4263A3B5D6E7

Heads of Tobochares spp. A, BT. goias, black mark pointing to canthus emarginating the eye A dorsal view B anterolateral view CT. fususD, ET. luteomargo, black mark pointing to straight anterior margin of the eye D dorsal view E anterolateral view FT. romanoaeGT. pemonHT. communisIT. microps.

https://binary.pensoft.net/fig/512600

Checklist of Tobochares species, their assigned species group, and known distribution. Asterisks (*) denote new country records for previously described species.

Species groupSpeciesKnown distribution
communis Tobochares akoerio sp. nov.Suriname
Tobochares anthonyae sp. nov.Venezuela (Bolívar)
Tobochares arawak sp. nov.Guyana
Tobochares atures sp. nov.Venezuela (Bolívar)
Tobochares canaima sp. nov.Venezuela (Amazonas)
Tobochares communis sp. nov.Brazil (Amapá, Roraima), Guyana, Suriname, Venezuela (Bolívar)
Tobochares kappel sp. nov.Suriname
Tobochares kolokoe sp. nov.Suriname
Tobochares microps sp. nov.Suriname
Tobochares pemon sp. nov.Suriname
Tobochares romanoae sp. nov.Brazil (Roraima)
emarginatus Tobochares canthus Kohlenberg & Short, 2017Venezuela (Amazonas)
Tobochares emarginatus Kohlenberg & Short, 2017Suriname
Tobochares fusus sp. nov.Brazil (Amapá), French Guiana
luteomargo Tobochares luteomargo sp. nov.Venezuela (Bolívar)
sulcatus Tobochares benettii sp. nov.Brazil (Amazonas)
Tobochares canaliculatus Kohlenberg & Short, 2017Venezuela (Amazonas)
Tobochares goias sp. nov.Brazil (Goiás)
Tobochares kasikasima Short, 2013Suriname
Tobochares kusad Kohlenberg & Short, 2017Guyana, Brazil* (Roraima)
Tobochares pallidus Kohlenberg & Short, 2017Venezuela (Amazonas, Bolívar)
Tobochares sipaliwini Short & Kadosoe, 2011Suriname, Brazil* (Roraima)
Tobochares sp. A in Kohlenberg & Short, 2017Venezuela (Amazonas)
Tobochares striatus Short, 2013Suriname
Tobochares sulcatus Short & García, 2007Venezuela (Amazonas, Bolívar)
KohlenbergATShortAEZ (2017) Revision of the Neotropical water scavenger beetle genus Tobochares Short and García, 2007 (Coleoptera, Hydrophilidae, Acidocerinae).ZooKeys669: 113146. https://doi.org/10.3897/zookeys.669.11773