Harmaline-induced tremor does not require electrical coupling in IO. A, Force transducer output from representative spontaneous tremor in a WT mouse before (thick line) and after (thin line) harmaline administration. Transducer output units are arbitrary but fixed across all measurements. B, Power spectra from force measurements in control (thick) and harmaline (thin) states; harmaline tremor is evident as a sharp, strong peak at ~14 Hz. C, Similar force measurements in a representative KO mouse before and after harmaline (same scale as in A). D, The power spectrum from the KO mouse tremor is similar to that of the WT mouse.