Table 2.

The frequencies of the maternal-offspring genotype combinations before selection. Genotypes have alleles ordered by their parent-of-origin, with the maternally inherited allele appearing first. The combinations are a function of the frequency of the four ordered genotypes (f_ij, where i is the maternally inherited allele) and the frequencies of the offspring genotypes within the maternal genotypes (u_ij, where again i is the maternally inherited allele). The frequencies of the ordered genotypes (f_ij) appear at the bottom of the columns for the offspring genotypes (the genotype frequencies are the same as those of the four possible maternal genotypes and so are only listed under the offspring genotypes for simplicity) and the frequencies of the offspring genotypes within the maternal genotypes (u_ij) are: u₁₁ = F +p₁(1 − F), u₁₂ = p₂(1 − F), u₂₁ = p₁(1 − F), u₂₂ = F +p₂(1 − F), and. Cells containing a ‘—’ indicate combinations that do not exist.

	Offspring genotype
	A1A1	A1A2	A2A1	A2A2
Maternal genotype
A1A1	f11u11	f11u12	—	—
A1A2	½f12u11	½f12u12	½f12u21	½f12u22
A2A1	½f12u11	½f12u12	½f12u21	½f12u22
A2A2	—	—	f22u21	f22u22
Genotype Frequencies	$f_{11}=p_1^2+p_1{p}_{2}F$	f12 = p1p2 − p1p2F	f21 = p1p2 − p1p2F	$f_{22}=p_2^2+p_1{p}_{2}F$