Skip to main content
NIHPA Author Manuscripts logoLink to NIHPA Author Manuscripts
. Author manuscript; available in PMC: 2023 Dec 8.
Published in final edited form as: Int J Sports Physiol Perform. 2022 Mar 17;17(7):1030–1036. doi: 10.1123/ijspp.2021-0021

Assessment of a Treadmill Speed Incline Conversion Chart: A Validation Study

Nicholas A Foreman 1, Emma J Lee 2, Christopher J Lundstrom 3
PMCID: PMC10707652  NIHMSID: NIHMS1946819  PMID: 35299154

Abstract

Purpose:

To investigate the validity of a treadmill speed incline conversion chart using physiological and subjective measures.

Methods:

Two groups of experienced runners (SLOW and FAST: divided based on their easy run pace) ran 6 speed incline combinations for 5 minutes each. Stages were equivalent according to the HillRunner.com chart, and stage order was randomized. Due to limitations of the chart, SLOW (n = 11) ran at inclines up to 4%, while FAST (n = 22) ran at inclines up 10%. Oxygen consumption (VO2), respiratory exchange ratio, heart rate, blood lactate, overall rating of perceived exertion (RPE), and leg RPE were measured for each stage. VO2 was compared against the VO2 predicted by the American College of Sports Medicine (ACSM) equation (ACSM VO2). Repeated-measures analysis of variance was used to detect differences between stages and inclines, and Hedges g was used as a measure of effect size.

Results:

Pooled results (0%–4%, N = 33) showed no incline effect on VO2, respiratory exchange ratio, heart rate, blood lactate, or RPE (P > .05; ηp2=.198), validating the chart at these inclines. At or above 6%, meaningful and significant increases occur in VO2 (g > 0.9, P < .05), with increases in heart rate, blood lactate, and leg RPE at higher inclines. ACSM VO2 underestimated oxygen consumption at all inclines up to 8% (P < .05) but not at 10% (45.9 [4.0] vs 46.7 [2.4] mL·kg−1·min−1; P = .186).

Conclusion:

The HillRunner.com chart is only valid at or below 4%. At higher inclines, supposedly equivalent stages result in increased exercise intensity. ACSM VO2 underestimates VO2 in trained runners at inclines up to 8%.

Keywords: running, training, ACSM prediction equation


Motorized treadmills are a common training tool used to approximate the effects of overground running, particularly during unfavorable outdoor conditions. They also allow for additional control of speed and incline without relying on local topography. This may be advantageous to runners training for ultramarathons, which are often completed on trails.1 As participation in ultramarathon running has been increasing rapidly over the last 20 years,2,3 more individuals may be interested in training on inclined surfaces to provide a competitive advantage, particularly in mountainous events. Running at incline without decreased speed leads to increased heart rate (HR), oxygen consumption (VO2), respiratory exchange ratio, and blood lactate concentrations (BL) when compared with level ground running.4,5 Thus, to create a comparable physiological response between flat and inclined running, runners may reduce their speed.

An important aspect of endurance training is accurate prescription of exercise intensities. Determining equivalent intensity levels at various speeds and inclines on motorized treadmills may be useful for exercise prescription and endurance training. The American College of Sports Medicine (ACSM) has created an equation to predict VO2 at various treadmill speeds and inclines, though the equation is limited to submaximal exercise.6 Free online resources also provide supposedly equivalent combinations of speed and incline and conversion to minutes-per-mile paces, a more familiar and commonly used measure of speed in the United States.7 One such chart, found at HillRunner.com, is widely used, with over 400,000 users visiting the page over 750,000 times in 2018 (see Supplementary Digital Material S5 [available online]). However, its purported basis is in unpublished research, so its validity and derivation remain unknown,8 limiting its credibility and potential use, both in research and to the public. Without validation, online conversion charts may under or overestimate physiological demands. Underestimation of exercise intensity may lead to unintended plateaus in performance, while overestimations may lead to overtraining9 or impaired recovery after exercise.10 Therefore, the primary aim of this study was to validate the Hillrunner.com Treadmill Pace Conversion chart (“the HillRunner.com chart,” see Supplementary Digital Material S6 [available online] for complete copy) by comparing subjective and physiological measures of exertion across theoretically equivalent intensity combinations of speeds and inclines in a sample of trained runners. A secondary aim was to compare measured VO2 with the predicted ACSM VO2 in this group.

Methods

Participants

Participants were recruited from the University of Minnesota-Twin Cities and surrounding communities. Inclusion criteria were as follows: an average of 3 runs per week over the previous year, one run of at least 60 minutes in the last month, a typical easy run pace between 2.68 m·s−1 (10:00 min·mile−1) and 4.47 m·s−1 (6:00 min·mile−1), and at least 18 years of age. Participants were excluded if they had sustained a major injury in the previous 3 months. Potential participants were screened for eligibility via a Qualtrics (Qualtrics, Provo, UT) survey prior to enrollment. This study was reviewed and approved by the institutional review board at the University of Minnesota (STUDY00006703) prior to participant enrollment and all participants provided written informed consent. Thirty-four participants (9 females; 32.0 [10.3] y old) met inclusion criteria and were enrolled in the study.

Study Design

A cross-over design was used with each participant completing 6 stages in a randomized order. Because the HillRunner.com chart did not provide equivalent speeds for the higher inclines for slower runners, participants were assigned into FAST and SLOW groups based on their easy run pace. There was an unequal allocation ratio between the groups, with the ultimate ratio of FAST:SLOW being approximately 2:1. Randomization was balanced across orders, with each individual running a unique order of inclines. The overall study design is outlined in Figure 1.

Figure 1 —

Figure 1 —

Overview of study design.

Pretesting

Individuals interested in participating completed a screening and running history survey prior to enrollment in the study. Survey data were used to characterize participants. A complete list of survey questions is included (see Supplementary Digital Material S7 [available online]). To estimate each athlete's running ability, an estimation of maximal oxygen consumption over 1 minute (VDOT) was calculated using the Run Smart Project VDOT Running Calculator, based on work done by Jack Daniels.7 The VDOT score provides a measure of aerobic fitness without requiring participants to undergo maximal VO2 testing, though it may underestimate true maximal VO2 in a trained population.11 Additional details on VDOT calculation are included in the Supplementary Digital Material S1 [available online].

Participant Preparation

Athletes came to the Human and Sport Performance Lab at the University of Minnesota-Twin Cities for a single visit. Standard previsit exercise testing measures were followed, as detailed in Supplementary Digital Material S1 (available online).

Testing Protocol

Treadmill testing was completed on a Woodway Pro XL treadmill (Woodway, Waukesha, WI). Participants warmed-up with one minute of walking at 1.34 m·s−1 and 0% incline before beginning the incline testing. Participants ran for 5 minutes at each incline (FAST: 0%, 2%, 4%, 6%, 8%, and 10%; SLOW: 0%, 0%, 1%, 2%, 3%, and 4%). Inclines differed between groups because the HillRunner.com chart does not list pace and incline conversions for speeds lower than 3.35 m·s−1. For SLOW, the second 0% stage (0%_second) was included so the number of stages would be identical for each group. Speed was adjusted according to the HillRunner.com chart to create an equivalent exercise intensity in each stage. For each stage, the pace corresponded to the participant’s self-reported easy run pace from the survey, assumed to be performed in level overground running. Additional details on selection of paces are included in the Supplementary Digital Material S1 (available online). For each participant, the order of inclines was randomized using an online list randomizer to mitigate any potential confounding effects of fatigue or cardiovascular drift.12

Data Acquisition

Throughout testing, respiratory gases were measured, and standard calibration procedures were completed prior to measurement (Ultima CPX and BreezeSuite software; MGC Diagnostics, St. Paul, MN). Mid 5-of-7 averaging was used for all VO2 measurements. HR was also recorded continuously using a Polar chest strap monitor and Polar RS800cx watch (Polar Electro Oy). The company description applies to both the HR monitor and the watch, as both were made by the same company. In the final 20 seconds of each stage, participants were asked to provide their overall rating of perceived exertion (RPE; 6–20) using the Borg RPE scale,13 which was visually presented to them each time. Participants were also asked to provide an RPE for their legs (LRPE), to understand any differences between overall exertion and localized muscular fatigue associated with uphill running.14-16 The treadmill display was covered during testing, so participants were blinded to the speed and incline of each stage. At the end of each stage, participants stepped to the side of the treadmill for a one-minute standing rest. At the beginning of each rest period, BL was assessed using a portable analyzer (Lactate Plus analyzer; Nova Biomedical, Waltham, MA), with blood samples obtained using a lancet on the participant’s fingertip. A fan was placed perpendicular to the treadmill at the side of the participant in an attempt to limit cardiovascular drift throughout testing.

Data Analysis

The VO2 (in milliliter per kilogram per minute) and HR (in beats per minute) were measured continuously for each participant, and data from the last 2 minutes of each stage were averaged for both, based on work by Jones and Doust.17 Plots of VO2 were also examined visually to confirm attainment of a steady state. For these measures, data were organized, prepared, and averaged in Microsoft Excel (2013; Microsoft Corp). HR data were transferred to Polar ProTrainer software (Polar Electro Oy) and examined for artifact prior to selection of the final 2 minutes of each stage for averaging.

Statistical testing was completed using SPSS (version 25; IBM, Armonk, NY). An interim power analysis was completed after the first 10 participants completed testing. The analysis indicated that a sample size of 20 would be needed based on an alpha of .05 and a power of 0.80. After collection of all data, but prior to analysis, data were checked for normality and heteroskedasticity. Repeated-measures analysis of variance tests were used to compare VO2, HR, BL, RPE, and LRPE between the 6 incline stages in FAST and SLOW. For data collected at inclines of 0%, 2%, and 4%, data from FAST and SLOW were pooled (ALL) and analyzed together for greater statistical power. Repeated-measures analysis of variance tests by stage number in FAST were used to determine the effect of stage number. Post hoc comparisons were done with Fisher least-squares difference tests due to our equivalency hypothesis, and thus, the priority to reduce type II error. Partial eta squared (ηp2) was used as the measure of effect size for analysis of variance main effects. A corrected version of Hedges g, which adjusts for smaller sample sizes, was used for post hoc comparisons.18 Paired-sample t tests were used to compare RPE and LRPE and measured VO2 to VO2 predictions based on the ACSM equation.6 Bland–Altman plots were prepared to further compare measured VO2 and ACSM VO2.19 After a possible trend in the Bland–Altman plots was observed, linear models were fit to examine potential systematic bias in the plots. Assumptions were checked using R performance package (version 0.7.2).20 An alpha level of P ≤ .05 was set as the standard for significance.

Results

Results are presented as mean (SD), and 95% confidence intervals are provided in square brackets where relevant. Thirty-four participants (9 females; 32.0 [10.3] y old) met the inclusion criteria and were enrolled into 2 groups: 22 in FAST for runners training at or above 3.35 m·s−1 (≤8:00 min·mile−1) for their easy runs and 11 in SLOW for runners training between 2.68 and 3.34 m·s− (<10:00 min·mile−1 and >8:00 min·mile−1) for their easy runs. Participant characteristics can be found in Table 1. Of the 27 participants who responded to the screening survey regarding their routine hill training, 70% reported encountering hills as a natural part of their training. Of the 27, 54% reported running hill repeats, which they reported completing an average of 2.5 times per month. Mean values in FAST and SLOW are presented in Tables 2 and 3, respectively. Information on pairwise comparisons in FAST and ALL are included in the Supplementary Digital Materials S2 and S3 (available online). Higher inclines lead to increases in exercise intensity; this was observed for most variables in FAST (Table 2 and see Supplementary Digital Material S4 [available online]). At lower inclines seen in SLOW, significant differences were limited to comparisons with the second 0% stage (Table 3). Differences in VO2 by incline in FAST exceed the minimum detectable difference at 6% when compared with 0% (Figure 2). HR (P < .03; g > 0.461), BL (P < .006, g > 0.596), and LRPE (P = .265 for 8%, P < .01 for all others; g > 0.318) were significantly higher during the 10% stage compared with all other inclines in FAST. There were also differences in BL between 0% and 8% (g = 0.393; P = .06) and between 2% and 6%, 8%, and 10% incline (P < .02; g > 0.50).

Table 1.

Participant Characteristics

Variable FAST (n = 22) SLOW (n = 11)
Age, y 31.0 (10.6) 35.1 (9.7)
Mass, kg 66.7 (7.3)* 73.1 (10.2)
Height, cm 175.6 (5.5) 175.3 (10.3)
VDOTa 59.17 (6.6)** 39.26 (8.7)
Easy run pace, m·s−1 3.56 (0.15)** 2.93 (0.16)
Km.Wk.Avg, km·wk−1 75.3 (29.0)** 33.6 (13.8)
Km.Wk.Peak, km·wk−1 95.4 (31.3)** 55.7 (29.6)

Abbreviations: Km.Wk. Avg, average weekly training distance over the previous 6 months; Km.Wk.Peak, peak weekly training distance over the previous 6 months. Note: Values are presented as mean (SD).

a

These values are analogous to 5000-m times of 17:15 and 24:27 for FAST and SLOW, respectively.

*

Different from SLOW, P < .05.

**

Different from SLOW, P < .01.

Table 2.

Measures of Exercise Intensity in FAST by Percentage Incline

Incline
0% 2% 4% 6% 8% 10%
VO2, mL·kg−1·min−1 42.18 (3.6)c,d,e,f 42.56 (4.1)c,d,e,f 43.22 (3.9)a,b,e,f 43.91 (4.1)a,b,e,f 44.66 (4.3)a,b,c,f 45.86 (4)a,b,c,d,e
RER 0.92 (0.07) 0.92 (0.07) 0.93 (0.09) 0.92 (0.06) 0.93 (0.08) 0.93 (0.07)
BL, mM 1.36 (0.87)f 1.24 (0.85)d,e,f 1.47 (1.17)f 1.54 (1.15)b,f 1.60 (1.15)b,f 1.94 (1.2)a,b,c,d,e
Heart rate, beats·min−1 146.3 (11.8)f 148.3 (10.7)f 147.7 (11.7)f 146.9 (12.3)f 148.1 (13.4)f 151.3 (12.5)a,b,c,d,e
RPE 10.7 (1.8) 10.6 (1.9) 11.0 (2.0) 10.5 (2.0) 10.9 (2.1) 11.4 (1.9)
LRPE 10.1 (1.7)f 10.2 (1.8)f 10.7 (2.2)f 10.4 (2.2)f 10.9 (2.3)f 11.9 (2.2)

Abbreviations: BL, blood lactate concentration; LRPE, RPE for the legs; RER, respiratory exchange ratio; RPE, rating of perceived exertion; VO2, oxygen consumption. Significantly different (P < .05) from a0%, b2%, c4%, d6%, e8%, f10%.

Table 3.

Measures of Exercise Intensity in SLOW by Percentage Incline

Incline
0% 0%_Second 1% 2% 3% 4%
VO2, mL·kg−1·min−1 37.29 (3.2) 36.91 (4.2) 37.1 (3.5) 37.18 (3.7) 36.97 (3.3) 36.62 (2.9)
RER 0.90 (0.04) 0.89 (0.05) 0.90 (0.04) 0.91 (0.04) 0.90 (0.04) 0.9 (0.04)
BL, mM 2.15 (1) 1.82 (0.84) 2.06 (1.26) 2.31 (1.27) 2.21 (1.37) 1.9 (0.70)
Heart rate, beats·min−1 153.3 (16.6)a 160.8 (14.9) 156.2 (14.9)a 152.6 (16.3)a 156.2 (14.8)a 154.4 (14.9)a
RPE 12.0 (1.9)a 13.5 (1.0) 12.0 (1.0)a 11.8 (2.0)a 12.1 (1.5)a 12 (1.3)a
LRPE 11.5 (2.5) 12.6 (2.2) 11.9 (1.6) 11.8 (2.6) 12.6 (2.1) 12.8 (1.8)

Abbreviations: BL, blood lactate concentration; LRPE, RPE for their legs; RER, respiratory exchange ratio; RPE, rating of perceived exertion; VO2, oxygen consumption.

a

Significantly different (P < .05) from 0%_second.

Figure 2 —

Figure 2 —

Difference in VO2 by incline in FAST labeled with effect size. Corrected Hedges g is used as the measure of effect size. Vertical bars indicate the 95% CIs for the difference in VO2. The horizontal line across the figure approximates the minimum detectable difference by the metabolic cart used in this study. Dark bars indicate a significant difference (ANOVA, P < .05). ANOVA indicates analysis of variance; VO2, oxygen consumption.

Data collected from SLOW and ALL further clarify changes seen at lower inclines. In ALL, there were significant differences in LRPE between 0% and 4% ([−1.35 to −0.18]; P = .012; g = 0.433) and between 2% and 4% ([−1.29 to −0.06]; P = .031; g = 0.365). Similar trends in LRPE were seen in the SLOW subgroup when comparing 0% with 3% ([−2.52 to −0.03]; g = 0.563) and 4% ([−2.19 to 0.01]; g = 0.586]). P values and effect sizes for the main effect of each variable by study group are included in the supplemental materials (see Supplementary Digital Material S4 [available online]).

In ALL, FAST, and SLOW, measured VO2 was lower than ACSM VO2 at all inclines (P < .05) except 10% (P = .186). Linear models to assess for a relationship in the Bland–Altman data showed no significant relationship at 0% (P = . 173), 2% (P = .647), or 4% (P = .750). There was a significant relationship at 6% (P = .048), but the homoscedasticity assumption was violated (P = .004). Significant relationships exist at 8% (P < .001, R2 = .49) and 10% (P = .004, R2 = .32). Therefore, regression lines were only included on the 8% (Figure 3E) and 10% plots (Figure 3F).

Figure 3 —

Figure 3 —

Bland–Altman plots of ACSM and measured VO2. Plots include average difference (long dashes) and limits of agreement (dotted line), each with a 95% CI (shaded regions). All plots are scaled from 30 to 55 mL·kg−1·min−1 on the x-axis. Points greater than 0 on the y-axis indicate an overestimation by the ACSM equation. When significant, linear regression lines were added. (A) 0%, (B) 2%, (C) 4%, (D) 6%, (E) 8%, and (F) 10%. ACSM indicates American College of Sports Medicine; VO2, oxygen consumption.

Discussion

The present study aimed to determine whether the Hillrunner.com chart could be used to accurately adjust speeds to produce equivalent efforts at different inclines from 0% to 10%, and to evaluate the accuracy of the ACSM equation for predicting VO2 across these inclines. While we found the Hillrunner.com chart to be accurate in producing equivalent efforts at inclines at or below 4%, it led to higher physiological and subjective efforts at higher inclines. The ACSM equation, on the other hand, overestimated VO2 at 0% to 4% in ALL, 0% to 4% in SLOW, and 0% to 8% in FAST. A novel relationship was seen at higher inclines (8%–10%) such that VO2 was no longer overestimated when VO2 was high.

These findings mirror previous research by Minetti et al,21 who showed increases in VO2 and the energy cost of running at 10% incline compared with 0%, even with some decrease in treadmill speed at 10%. Overestimation of VO2 by the ACSM equation is also in agreement with previous research. These findings may be explained by previous literature showing a reduction in submaximal VO2 (ie, improved running economy) at a given workload through chronic endurance training.22,23 In each of the previous studies and in the present study, the submaximal VO2 would likely be lower than that of the general population on which the ACSM equation is based.23 In line with previous research,24 the limits of agreement in the Bland–Altman plots are wide (~12 mL·kg−1·min−1), suggesting a high degree of individual variability when applying the ACSM equation (Figure 3). Plots of VO2 at inclines below 8% show a consistent overestimation in ACSM VO2. At inclines of 8% and 10%, a linear relationship exists between the differences in measured VO2 versus ACSM VO2; at a lower mean VO2, the ACSM equation overestimates VO2 and becomes more accurate as mean VO2 increases. We are unable to determine the cause of the transition from overestimation to accurate prediction at high inclines as VO2 increases. We caution against use of the ACSM equation in a trained population until the relationships between speed, incline, and magnitude of VO2 are better understood.

Physiological measures provide specific insight into the differences at higher inclines and the implications for use of the HillRunner.com chart. Increases in VO2 at 2% and 4% had 95% confidence intervals within the typical error of measurement for the metabolic system used25 (Figure 2). At higher inclines, VO2 continues to increase, with large effect sizes between low and high inclines (Figure 2), suggesting the chart is not valid beyond 4%. As incline increased from 0% to 8% in FAST, we observed no changes in HR, and corresponding effect sizes were low (see Supplementary Digital Material S3 [available online]). A difference of 3 to 5 beats per minute was observed when comparing HR at 10% to HR at lower inclines. Across inclines, the 95% confidence intervals for HR were quite wide (4–6 beats per minute, see Supplementary Digital Material S2 [available online]), indicating that individual HR responses may vary. For individuals whose training is guided by HR zones, these increases in HR may be enough to transition from a low-intensity zone to a moderate-intensity zone, which has been linked to reduced training adaptations and impaired race performance over a 5-month season.26 We found BL in FAST began to increase by up to 0.5 mM at 6% when compared with both 0% and 2% (see Supplementary Digital Material S2 [available online]), and there were significant differences between 10% and all lower inclines. Depending on the intensity of exercise, these increases may also be enough to transition beyond a lactate steady state, leading to marked increases in BL, HR, and respiratory rate.27 Increases in LRPE followed a comparable trend, with significant differences between inclines below 6% when compared with 10%. Based on data from FAST, the decreases in speed provided by the chart do not sufficiently correct for increased intensity above 4%.

Data from SLOW and ALL support the accuracy of the chart at lower inclines. Physiological measures of exercise intensity were not significantly different, and effect sizes were relatively small for both the main effect (all ηp2 in ALL < .2) and for pairwise comparisons between inclines (all g < 0.31). While there was a significant increase in LRPE at these inclines, effect sizes were not notable in ALL (g < 0.4), and 95% confidence intervals suggest this difference is of little practical relevance (see Supplementary Digital Material S2 [available online]). Effect sizes were larger in SLOW than ALL (g ≈ 0.6), but 95% confidence intervals were also wider (2 units in SLOW vs <1.2 in ALL). Given the greater power and tighter confidence intervals in ALL, the chart is likely accurate at all inclines up to 4%.

As this was a validation study, we felt it was most important to limit type II error; if inclines were reported as being equivalent, we wanted to ensure they were truly equivalent. As a result, we chose to use Fischer least significant difference instead of a Bonferroni correction. Had a Bonferroni correction been applied, the majority of notable findings would meet the threshold for significance (P ≤ .0083).

Evaluation of Testing Duration

We observed increases in HR, RPE, and LRPE over time. However, study design thoughtfully included randomization of inclines to mitigate any stage order effect. Therefore, these increases do not affect the validity of our overall results. Time-dependent elevations in HR, characteristic of cardiovascular drift, were observed. This is likely the cause of increased HR throughout testing in the present study.28 A change in VO2 over time was not observed, which is in agreement with previous literature showing the absence of a VO2 slow component during low-intensity exercise or at incline.29,30

Practical Applications

The HillRunner.com chart can be used by trained runners to create treadmill speed–incline combinations equivalent to overground speeds ranging from 2.68 to 3.62 m·s−1 (10:00–7:25 min·mile−1), provided the desired incline is <6%. Inclines greater than 4% are likely to induce increases in physiological and psychological measures of exercise intensity. We assessed only efforts that were submaximal in nature, and further studies are required to validate the HillRunner.com chart at higher speeds and maximal efforts. Our findings do not support use of the ACSM prediction equation to predict submaximal VO2 at inclines under 10% in a trained population.

As noted in the study design, the HillRunner.com chart does not have any paces slower than 2.68 m·s−1 at a 10% incline, making the higher inclines impractical for some recreational athletes, walkers, or clinical populations. The present study used moderately trained runners to evaluate the validity of the HillRunner.com chart. As such, these findings cannot be applied to untrained or highly trained, elite runners. Furthermore, a limited number of female runners was included in the study, so it is unknown whether prescription using the HillRunner.com chart differs by sex. There is no previous indication or known physiological mechanism for a sex difference, but one still may exist.

Conclusion

The HillRunner.com chart may be used for accurate prescription of submaximal exercise intensity in trained runners at speeds ranging from 2.68 to 4.47 m·s−1 (10:00–6:45 min·mile−1), and at inclines up to 4%. Across the same range of speeds, adjustments in speed, as recommended by the HillRunner.com chart, are not enough to maintain exercise intensity. Beyond a 4% incline, increases in VO2, HR, and BL should be expected compared with speed adjustments recommended at lower inclines. Further decreases in speed beyond what the HillRunner.com chart currently recommends may be required for trained runners wishing to create equivalent efforts at these higher inclines. We show a novel relationship between magnitude of VO2 and the degree of VO2 overestimation by the ACSM equation. Based on findings of the present study, the ACSM equation should not be relied upon to accurately predict submaximal VO2 at inclines under 10% in a trained population.

Supplementary Material

S5
S6
S7
S4
S3
S2
S1

Acknowledgments

The current work was supported by funding from the University of Minnesota’s Undergraduate Research Opportunities Program. The project described was also supported by grant number T32AG029796. The authors are grateful to the participants for their time and efforts. Conceptualization was done by N. A.F and C.J.L.; methodology by N. A.F., C.J.L., and E.J.L.; formal analysis by C.J.L; investigation by N.A.F., C.J.L., and E.J.L.; data curation by N.A.F.; writing—original draft by N.A.F.; and writing—review and editing by C.J.L. and E.J.L.

Contributor Information

Nicholas A. Foreman, School of Kinesiology, University of Minnesota, Minneapolis, MN, USA.

Emma J. Lee, Div of Rehabilitation Science, Dept of Rehabilitation Medicine, University of Minnesota, Minneapolis, MN, USA.

Christopher J. Lundstrom, School of Kinesiology, University of Minnesota, Minneapolis, MN, USA.

References

  • 1.Hoffman MD, Ong JC, Wang G. Historical analysis of participation in 161 km ultramarathons in North America. Int J Hist Sport. 2010;27(11):1877–1891. doi: 10.1080/09523367.2010.494385 [DOI] [PubMed] [Google Scholar]
  • 2.Cejka N, Rust CA, Lepers R, Onywera V, Rosemann T, Knechtle B. Participation and performance trends in 100-km ultra-marathons worldwide. J Sports Sci. 2014:32(4):354–366. doi: 10.1080/02640414.2013.825729 [DOI] [PubMed] [Google Scholar]
  • 3.Ronto P. The state of ultra running 2020. Published 2020. Accessed August 1, 2020. https://runrepeat.com/state-of-ultra-running [Google Scholar]
  • 4.Padulo J, Powell D, Milia R, Ardigò LP. A paradigm of uphill running. PLoS One. 2013;8(7):e69006. doi: 10.1371/journal.pone.0069006 [DOI] [PMC free article] [PubMed] [Google Scholar]
  • 5.Klein RM, Potteiger JA, Zebas CJ. Metabolic and biomechanical variables of two incline conditions during distance running. Med Sci Sports Exerc. 1997:29(12):1625–1630. doi: 10.1097/00005768-199712000-00012 [DOI] [PubMed] [Google Scholar]
  • 6.Arena R, Riebe D, Thompson PD, eds. ACSM's Guidelines for Exercise Testing and Prescription. 9th ed. Philadelphia, PA: Wolters Kluwer, Lippincott Williams & Wilkins; 2014. [Google Scholar]
  • 7.Daniels J. Daniels’ Running Formula. 3rd ed. Champaign, IL: Human Kinetics Publishers Inc.; 2014. [Google Scholar]
  • 8.Hill R. Treadmill Pace Conversions. Accessed August 1, 2020. https://www.hillrunner.com/calculators/treadmill-pace-conversions/ [Google Scholar]
  • 9.Lehmann M, Foster C, Keul J. Overtraining in endurance athletes: a brief review. Med Sci Sports Exerc. 1993:25(7):854–862. doi: 10.1249/00005768-199307000-00015 [DOI] [PubMed] [Google Scholar]
  • 10.Seiler S, Haugen O, Kuffel E. Autonomic recovery after exercise in trained athletes: Intensity and duration effects. Med Sci Sports Exerc. 2007:39(8):1366–1373. doi: 10.1249/mss.0b013e318060fl7d [DOI] [PubMed] [Google Scholar]
  • 11.Scudamore EM, Barry VW, Coons JM. An evaluation of time-trial-based predictions of v[combining dot above]o2max and recommended training paces for collegiate and recreational runners. J strength Cond Res. 2018:32(4):1137–1143. doi: 10.1519/JSC.0000000000001942 [DOI] [PubMed] [Google Scholar]
  • 12.Haahr M, Haahr S. List Randomizer. Accessed July 1, 2019. https://www.random.org/lists/ [Google Scholar]
  • 13.Borg GA. Psychophysical bases of perceived exertion. Med Sci Sports Exerc. 1982;14:377–381. [PubMed] [Google Scholar]
  • 14.Bolgar MR, Baker CE, Goss FL, Nagle E, Robertson RJ. Effect of exercise intensity on differentiated and undifferentiated ratings of perceived exertion during cycle and treadmill exercise in recreationally active and trained women. J Sports Sci Med. 2010;9(4):557. [PMC free article] [PubMed] [Google Scholar]
  • 15.Pandolf KB, Billings DS, Drolet LL, Pimental NA, Sawka MN. Differentiated ratings of perceived exertion and various physiological responses during prolonged upper and lower body exercise. Eur J Appl Physiol Occup Physiol. 1984:53(1):5–11. doi: 10.1007/BF00964681 [DOI] [PubMed] [Google Scholar]
  • 16.Aliverti A, Kayser B, Lo Mauro A, et al. Respiratory and leg muscles perceived exertion during exercise at altitude. Respir Physiol Neurobiol. 2011;177(2):162–168. doi: 10.1016/j.resp.2011.03.014 [DOI] [PubMed] [Google Scholar]
  • 17.Jones AM, Doust JH. A 1% treadmill grade most accurately reflects the energetic cost of outdoor running, J Sports Sci. 1996:14(4):321–327. doi: 10.1080/02640419608727717 [DOI] [PubMed] [Google Scholar]
  • 18.Hedges LV, Olkin I. Statistical Methods in Meta-Analysis. 1985. Orlando, FL: Academic Press, Inc. doi: 10.2307/1164953 [DOI] [Google Scholar]
  • 19.Bland JM, Altman DG. Statistical methods for assessing agreement between two methods of clinical measurement. Lancet. 1986;327(8476):307–310. doi: 10.1016/S0140-6736(86)90837-8 [DOI] [PubMed] [Google Scholar]
  • 20.Lüdecke D, Ben-Shachar MS, Patil I, Waggoner P, Makowski D. Performance: An R package for assessment, comparison and testing of statistical models. Journal of Open Source Software. 2021:6(60):3139. doi: 10.21105/joss.03139 [DOI] [Google Scholar]
  • 21.Minetti AE, Moia C, Roi GS, Susta D, Ferretti G. Energy cost of walking and running at extreme uphill and downhill slopes, J Appl Physiol. 2002:93(3):1039–1046. doi: 10.1152/japplphysiol.01177.2001 [DOI] [PubMed] [Google Scholar]
  • 22.Morgan DW, Bransford DR, Costill DL, Daniels JT, Howley ET, Krahenbuhl GS. Variation in the aerobic demand of running among trained and untrained subjects. Med Sci Sports Exerc. 1995:27(3):404–409. doi: 10.1249/00005768-199503000-00017 [DOI] [PubMed] [Google Scholar]
  • 23.Barnes KR, Kilding AE. Running economy: measurement, norms, and determining factors. Sports Med—Open. 2015;1(1):8. doi: 10.1186/s40798-015-0007-y [DOI] [PMC free article] [PubMed] [Google Scholar]
  • 24.Hall C, Figueroa A, Fernhall B, Kanaley JA. Energy expenditure of walking and running: comparison with prediction equations. Med Sci Sports Exerc. 2004;36(12):2128–2134. doi: 10.1249/01.MSS.0000147584.87788.0E [DOI] [PubMed] [Google Scholar]
  • 25.Miodownik S, Carlon VA, Ferri E, Burda B, Melendez JA. System of automated gas-exchange analysis for the investigation of metabolic processes. J Appl Physiol. 2000;89(1):373–378. doi: 10.1152/jappl.2000.89.1.373 [DOI] [PubMed] [Google Scholar]
  • 26.Esteve-Lanao J, Foster C, Seiler S, Lucia A. Impact of Raining intensity distribution on performance in endurance athletes, J Strength Cond Res. 2007:21(3):943–949. doi: 10.1519/r-19725.1 [DOI] [PubMed] [Google Scholar]
  • 27.Faude O, Kindermann W, Meyer T. Lactate threshold concepts: how valid are they? Sports Med. 2009:39(6):469–490. doi: 10.2165/00007256-200939060-00003 [DOI] [PubMed] [Google Scholar]
  • 28.Wingo JE. Exercise intensity prescription during heat stress: a brief review. Scand J Med Sci Sports. 2015;25(suppl 1):90–95. doi: 10.1111/sms.12381 [DOI] [PubMed] [Google Scholar]
  • 29.Reis VM, Guidetti L, Duarte JA, et al. Slow component of VO2 during level and uphill treadmill running: relationship to aerobic fitness in endurance runners, J Sports Med Phys Fitness. 2007;47(2):135–140. [PubMed] [Google Scholar]
  • 30.Pringle JSM, Carter H, Doust JH, Jones AM. Oxygen uptake kinetics during horizontal and uphill treadmill running in humans. Eur J Appl Physiol. 2002:88(1–2):163–169. doi: 10.1007/s00421-002-0687-0 [DOI] [PubMed] [Google Scholar]

Associated Data

This section collects any data citations, data availability statements, or supplementary materials included in this article.

Supplementary Materials

S5
S6
S7
S4
S3
S2
S1

RESOURCES