Beyond nutmeg, mace, and cloves: Checklist of the liverworts and hornworts of Maluku Islands (Moluccas), Indonesia

Abstract

The first ever liverwort and hornwort checklist is provided for the Maluku Islands (Moluccas/Spice Islands) of Indonesia. We report 355 accepted and 16 doubtful species and reject 22 species previously reported for Maluku Islands. The list is based on the specimens housed in the Herbarium Bogoriense (BO) and reports from over 500 literature references, including monographs, regional studies, and molecular investigations. The Maluku Islands are part of the Wallacea Biodiversity Hotspot with many unique species found only in Wallacea. Publications focusing on liverworts and hornworts of Maluku Islands are few and scattered. Considering regionally widespread species that have been recorded elsewhere, we predict that further fieldwork exploring the diversity of habitats coupled with collections unveiled from regional herbaria, a number of new records remain to be reported.

Introduction

The Maluku Islands (Moluccas) is an island archipelago of over 1,000 islands situated in eastern Indonesia south of the Philippine island of Mindanao and east of Sulawesi, scattered in a series of irregular arcs spanning the Maluku, the Banda, and the Seram Seas (Zerner 1994; Leunufna and Evans 2014). The Maluku Islands were also formerly known as the Spice Islands because the archipelago held great geopolitical significance due to the abundance in spices as a commodity (Van Gils and Cox 1994). For instance, the spice nutmeg (Myristicafragrans Houtt.) a small-holder’s crop, with 3500 years of antiquity, is indigenous to the Banda Islands, and is now grown in about 12 countries including some secondary centers of domestication (Peter 2001; Sasikumar 2021).

The biogeographical region of Maluku Islands (Fig. 1) encompasses the Indonesian geopolitical provinces of North Moluccas (Maluku Utara Province) and part of South Moluccas (Maluku Province) (Rutgrink et al. 2018). The major island groups in the north Moluccas are Halmahera, Bacan, Morotai, Obi, Sula, Ternate, and Tidore whereas Seram (previously known as Ceram), Buru, Banda, and Ambon are situated in the south. Many of the islands of Maluku are comprised of fragments of continental crusts (Monk et al. 1997) and are one of the most complex biogeographical and oceanographic areas on Earth (Pitriana et al. 2020). The island groups have experienced considerable, but very different tectonic displacements. During the last 10 million years, the North Moluccas moved westwards along the north coast of New Guinea to their current position. The Southern Moluccas moved northwards from Australia, west of New Guinea (Rutgrink et al. 2018). Not surprisingly the flora shows a marked similarity to those of New Guinea and Australia but also includes many endemic species and at least two endemic genera (Parakibara Philipson and Siphokentia Burret) (Burret 1927; Philipson 1985).

Figure 1.

Moluccas in the Indonesian archipelago (inset) as defined in the present paper and the islands and archipelagos here used.

Famously, western exploration of the natural history of the Maluku Islands dates back to the 17^th century, starting with Georg Everhard Rumphius, and later, for example, Alfred Russel Wallace (Strack 1993; Lamoureux 1990). Notably, Wallace (1863), proposed a sharp boundary of the Indo-Australian Archipelago based upon land-mammal and land-bird distributions. It has since achieved iconic status and today its significance is recognized well beyond the confines of the biogeography community (Ali and Heaney 2021). Yet, it is noteworthy that the application in recent years of modern analytical techniques has not led to a consensus view on where the lines/areas should run/be placed (Ali and Heaney 2021). The Maluku Islands are part of what is termed as the Wallacea Biodiversity Hotspot, a region that covers an area of 338 thousand square kilometers with thousands of islands supporting highly diverse biological communities with many unique species found only in Wallacea (van Welzen et al. 2005; CEPF 2014). Wallacea was identified as one of the original 25 biodiversity hotspots in the world (Myers et al. 2000).

Maluku Islands are situated in the “Tropical and Subtropical Moist Broadleaved Forests” biome (Olson et al. 2001) and contribute approximately 24% of forest cover of the Wallacea land surface. The archipelago as here defined is separated into four ecoregions, eastern part of “Sulawesi Lowland Rainforests”, “Buru Rainforests”, “Halmahera Rainforests” and “Seram Rainforests” (Fig. 3, https://www.oneearth.org/bioregions/sulawesi-maluku-islands-au14/) and is considered home to much endemic flora (CEPF 2014). The Maluku Islands are one of the few places in the Indonesian archipelago where it is possible to find a complete altitudinal sequence of vegetation, with few places elsewhere in the tropics providing a comparable range (Edwards et al. 1993). Many of the forests in Maluku are lowland evergreen and semi-evergreen forests (Fig. 2), especially on Halmahera and adjacent islands, tropical montane forests, limestone (karst) geologies, and ultrabasic (serpentine) geologies (Ellen 1997; Abdo 2017).

Figure 3.

Ecoregions of the Moluccas Islands.

Figure 2.

Vegetation of the Aketajawe-Lolobata National Park in Halmahera. Photo: Ida Haerida.

Liverworts and hornworts are, together with mosses, commonly referred to as bryophytes. They represent three early diverging clades of land plants forming the second largest group of green land plants after flowering plants and are of critical biological, ecological, and phylogenetic significance. Bryophytes serve as the “macrophytes,” providing a matrix where many microscopic organisms live, including tardigrades, mites, rotifers, micro-molluscs, microalgae, microfungi, and prokaryotes (Gerson 1982; Huttunen et al. 2017; Muggia and Grube 2018). They also play an important role as possible indicators of climate change (e.g., Lindo et al. 2013; Ruklani et al. 2021), in nutrient cycling (Rieley et al. 1979) and through their water retention reducing soil nutrient loss and flooding risk (Anderson et al. 2010). Because of their physiological properties they have therefore been used extensively as ecological indicators (e.g., Pakeman et al. 2019; Déleg et al. 2021). Resolving relationships among bryophytes and their relationships to the remaining land plants is critical for understanding the evolution of fundamental innovations within land plants (Leebens-Mack et al. 2019).

Historically, relatively few publications have focused on plant collections from the Maluku Islands, despite the significance of the flora. Approximately 4,442 species of plants and fungi were recorded from Maluku Islands until 2017 (Retnowati and Rugayah 2019). Herbarium Amboinense, a classic work by Rumphius (1741–1755), contains descriptions of plants commonly found in Ambon and the surrounding islands. However, the floristic knowledge of liverworts and hornworts remains poor, especially compared to seed plants, and has scarcely been published. Seram and Ambon have been the most investigated areas for liverworts and hornworts and have been reported by several authors (e.g. Akiyama 2009; Doei 1987a, 1987b; Hasegawa 1986; Hattori 1986b; Inoue 1986; Mizutani 1986a). One of the most significant contributions to bryophytes of the Maluku Islands is the checklist of bryophytes from Seram and Ambon by Akiyama (2009) who recorded as many as 226 species of liverworts. One species, Lepidoziaintegrifolia Doei, is an endemic species of Seram (Doei 1987a).

We here present the first ever checklist of liverworts and hornworts for Maluku Islands in an effort to further promote bryological research, especially in the eastern part of Indonesia. The checklist complements recently published checklists for Indonesia including Söderström et al. (2010) for Java and Nadhifah et al. (2021) for the Lesser Sunda Islands. The significance of checklists is summarized by Söderström et al. (2008), including outlining the utility of checklists as powerful and important tools, and their applicability to taxonomy, systematics, and conservation.

Liverwort types from Maluku

Not less than 113 names (9 of them invalid) have original material from the Maluku Islands. Of the validly published names, 13 have holotypes and 47 of the rest are lectotypified. However, 43 names remain to be typified. Here we designate lectotypes, or validate earlier unsuccessful lectotypifications, for 13 names. For a discussion on lectotypification procedures, including the concept of inadvertent lectotypifications, see Söderström and Hentschel (2023). In the list below we cite the lectotypification source and indicate where further typifications are needed.

Chandonanthusgracilis Herzog, Hedwigia 66 (6): 341, 1926 (Herzog 1926b).

Original citation. Ceram: Hoale-Paß, ca. 1600 m, leg. E. Stresemann, 1911.

Lectotype (here designated, cf. Váňa et al. 2013: 30 as “holotype”). Indonesia, Moluccas, Seram, Hoale Pass, 1600 m., 1911, Stresemann s.n. (JE04005601, https://je.jacq.org/JE04005601).

Note. Only one specimen has been located but it cannot be ruled out that there exist duplicates elsewhere. Váňa et al. (2013: 30) call it “holotype” but this is too late for an inadvertent lectotypification. The specimen is also annotated as holotype by K. Amamoto in 2020. In case duplicates are found, we here formally designate it as a lectotype.

Currently accepted name.Plicanthushirtellus (F.Weber) R.M.Schust. (see Váňa et al. 2013).

Lepidozialongifolia Steph., Sp. Hepat. (Stephani) 3: 606, 1909 (Stephani 1909a).

Original citation. Hab. Amboina (Karsten).

Lectotype (here designated). Indonesia, Moluccas, Ambon, Karsten (G00069659 https://www.ville-ge.ch/musinfo/bd/cjb/chg/adetail.php?id=123550&lang=en).

Note. The lectotype is annotated “type” by Mizutani in Feb. 1975.

Lophocoleadeningeri Herzog, Beih. Bot. Centralbl. 38 (2): 321, 1921 (Herzog 1921).

Original citation. Buru: Im Urwald des Kapalamadang, leg. Dr. K. Deninger †, 1906.

Lectotype (here designated). Indonesia, Moluccas, Buru, Im Urwald des Kapalamadang, Deninger 1906 (JE04005314 https://je.jacq.org/JE04005314).

Note. JE04005314 is marked as “holotype” by R. Grolle. We are not aware of any other type material.

Lopholejeuneasagranavar.dentistipula Schiffn. ex P.Syd., Just’s Bot. Jahresber. 19 (1): 246, 1894 (Sydow 1894) (‘ sagraeana ’).

Original citation. Ins. Amboina, in iisdem locis ac typica, pauca tantum frustula sed c. per. et ♂ inventa sunt (7. 6. 75).

Lectotype (here designated, Zhu and Gradstein 2005: 85 as “holotype”). Indonesia, Moluccas, Ambon, 7 Jun 1875, D. Naumann (FH).

Note.Zhu and Gradstein (2005: 85) call it “holotype” but this is too late for an inadvertent lectotypification.

Currently accepted name.Caudalejeuneareniloba (Gottsche) Steph. (see Zhu and Gradstein 2005).

Madothecacrenilobula Herzog, Beih. Bot. Centralbl. 38 (2): 328, 1921 (Herzog 1921).

Original citation. Buru: Im Urwald des Kapalamadang, leg. Dr. K. Deninger †, 1906.

Lectotype (here designated). Indonesia, Moluccas, Buru, Im Urwald des Kapalamadang, 1906, K. Deninger (JE04001514 https://je.jacq.org/JE04001514, isotype JE04001515 https://je.jacq.org/JE04001515).

Note. The lectotype was annotated “Type!” and the isotype “isotype” by S. Hattori in 1968 but has apparently never been published.

Currently accepted name.Porellajavanica (Gottsche) Inoue (see Hattori 1975e).

Mastigobryumdeningeri Herzog, Beih. Bot. Centralbl. 38 (2): 322, 1921 (Herzog 1921).

Original citation. Buru: In den Bergen Mittelburus, ca. 1500 m, leg. Dr. K. Deninger †, 1911.

Lectotype (here designated). Indonesia, Moluccas, Buru, Mittelburu, ca. 1500 m, 1911, Deninger 10 (JE04006158 https://je.jacq.org/JE04006158, isolectotype JE04006157 https://je.jacq.org/JE04006157).

Note. The selected lectotype is richer than the isolectotype and has a label in Herzog’s handwriting marked “n.sp.”. The isolectotype is marked “Typus” and annotated as type and syn. to Bazzanialongicaulis by Kitagawa 18 Nov 1978 but no typification seems to have been published.

Currently accepted name.Bazzanialongicaulis (Sande Lac.) Schiffn., syn. nov.>

Mastigobryumnigricans Herzog, Beih. Bot. Centralbl. 38 (2): 322, 1921 (Herzog 1921).

Original citation. Buru: Im Urwald des Kapalamadang, leg. Dr. K. Deninger †, 1906.

Lectotype (here designated). Indonesia, Moluccas, Buru, Kapalamadang, 1906, K. Deninger 112 (JE04006129 https://je.jacq.org/JE04006129).

Note. The lectotype is marked “n.sp.” on the label and annotated by N. Kitagawa 1991 as type.

Mastigobryumstresemannii Herzog, Beih. Bot. Centralbl. 38 (2): 324, 1921 (Herzog 1921).

Original citation. Ceram: Paßhöhe zwischen Mansela und Wolu (Useahánpaß), ca. 1750 m, leg. E. Stresemann, 1911.

Lectotype (here designated). Indonesia, Moluccas, Seram, between Mansela and Wolu ca. 1750 m, 1911, E. Stresemann 110, (JE04005944 https://je.jacq.org/JE04005944).

Note. The lectotype is marked “n.sp.” on the label by Herzog and annotated as type by N. Kitagawa in 1978.

Currently accepted name.Bazzaniastresemannii (Herzog) N.Kitag.

Mastigobryumvermiculare Herzog, Hedwigia 66 (6): 339, 1926 (Herzog 1926b).

Original citation. Ceram: Hoale, ca.1100m, leg. E. Stresemann, 1911.

Lectotype (here designated). Indonesia, Moluccas, Seram, Hoale, ca. 1100 m, 1911, Stresemann (JE04006130 https://je.jacq.org/JE04006130).

Note. Only one specimen is online in JE and without any annotation.

Pycnolejeuneaventricosa Schiffn. ex P.Syd., Just’s Bot. Jahresber. 19 (1): 246, 1894 (Sydow 1894).

Original citation. In insula Amboina cum aliis Lejeuniis parcissime, cum fl. ♀ et periantbiis vetustis (7. 6. 75) [cf. Schiffner 1890: 32].

Lectotype (here designated, cf. Zhu and Lai 2005: 200 as “holotype”). Indonesia, Moluccas, Ambon, Gazellen Expedition, 7.6.1875, Naumann (FH00284228 https://kiki.huh.harvard.edu/databases/specimen_search.php?mode=details&id=206370).

Note. “Holotype” by Zhu and Lai (2005: 200) is too late to be an inadvertent lectotype (cf. ICN2018 Art. 7.11) and is here designated as lectotype.

Currently accepted name.Cheilolejeuneaventricosa (Schiffn. ex P.Syd.) Xiao L.He

Schistochilaamboinensis Steph., Sp. Hepat. (Stephani) 4: 77, 1909 (Stephani 1909c).

Original citation. Hab. Amboina.

Lectotype (here designated, cf. So 2003a: 87 as “holotype”). Indonesia, Moluccas, Ambon, 1883, Luerssen 1270 (G00067852 https://www.ville-ge.ch/musinfo/bd/cjb/chg/adetail.php?id=131418&lang=en).

Note.So (2003a: 87 as “holotype”) is too late for an inadvertent lectotypification.

Currently accepted name.Schistochilabeccariana (De Not.) Trevis. (see So 2003a).

Schistochilainversa Herzog, Hedwigia 66 (6): 342, 1926 (Herzog 1926b).

Original citation. Ceram: Hoale-Paß, ca. 1600 m, leg. E. Stresemann, 1911; in Bruchstücken aus Lepidoziaceramensis und L. Ferdinandi Mülleri herausgelesen.

Lectotype (here designated, cf. So 2003a: 98 as “holotype”). Indonesia, Moluccas, Seram, Hoale-Pass, ca. 1600 m, 1911, E. Stresemann s.n. (JE04002842 https://je.jacq.org/JE04002842).

Note.So (2003a: 98 as “holotype”) is too late to be an inadvertent lectotypification.

Currently accepted name.Schistochilasciurea (Nees) Schiffn. (see So 2003a).

Schistochilapurpurascens Herzog, Hedwigia 66 (6): 341, 1926 (Herzog 1926b).

Original citation. Ceram: Hoale-Paß, ca. 1600 m, leg. E. Stresemann, 1911.

Lectotype (here designated, cf. So 2003a: 80 as “holotype”). Indonesia, Moluccas, Seram, Hoale-Pass, ca. 1000 m, Mar-Aug 1911, E. Stresemann 201 (JE04002840 https://je.jacq.org/JE04002840, isotype L-937.235.12 https://data.biodiversitydata.nl/naturalis/specimen/L%20%200444630).

Note. “holotype” by So (2003a: 80) is too late for an inadvertent lectotypification.

Currently accepted name.Schistochilaacuminata Steph. (see So 2003a).

Materials and methods

Nomenclature and taxonomy follow the world checklist of hornworts and liverworts (Söderström et al. 2016) with some updates from recent literature. Sources include more than 500 publications found through the work with Early Land Plants Today (ELPT) database of liverwort taxonomy and distribution, and with some consultation with taxonomic experts. In addition, we also examined the specimens collected from the Maluku Islands in the Herbarium Bogoriense (BO). The checklist follows a similar format of a series of previous liverwort and hornwort regional checklists by the authors, e.g., Java, Lesser Sunda Islands (Söderström et al. 2010; Nadhifah et al. 2021). References citing primary reports (citing specimens or material seen by the author) are given in bold. Secondary references not citing any material seen are given in normal font. In cases where we are not aware of any primary source, or the earliest report is a secondary source without any clear connection to a primary source published later, we comment on the situation. In the current checklist we note under which name a taxon is reported except when it is reported under the currently accepted name.

We here accept reports of 355 species from the area in addition to 16 reported species that we consider as doubtful and 21 species reported whose occurrence we reject from the islands. Species number known varies a lot between the islands/archipelagos (Table 1, Fig. 4). We are not aware of a single report from Sula while 258 species are known from Seram. Thus, we expect that with further investigations the number of species will rise considerably for many of the islands in the region.

Table 1.

Number of accepted, doubtful and rejected species of liverworts and hornworts from Moluccas and the recognized areas (islands/archipelagos) within the Moluccas.

Area	Accepted	Doubtful	Rejected
Ambon	143	11	10
Bacan	26	0	0
Banda	7	2	0
Buru	13	0	2
Halmahera	31	3	0
Morotai	2	0	0
Obi	2	0	0
Seram	258	7	5
Sulu	0	0	0
Ternate/Tidore	22	0	2
Moluccas	355	16	22

Figure 4.

Species richness on the different islands/archipelagos of Moluccas. The darker the more species known (cf. Table 1). White (Sulu I) means that no liverwort or hornwort species are known from the area.

Taxa accepted for Moluccas

Anthocerotophyta

Dendroceros Nees

Dendrocerosacutilobus Steph. MOLUCCAS: Ambon: LECTOTYPE [Piippo 1993a: 44], LECTOTYPE of Dendroceroskarstenii [to be designated], Schiffner 1898 as Dendroceroskarstenii, Stephani 1909b, Stephani 1917 as Dendroceroskarstenii, Stephani 1917, Bonner 1965 also as Dendroceroskarstenii, Hasegawa 1980, 1982, 1986, 2002, Piippo 1993a, Peñalosa et al. 2019. Seram: Akiyama 1986, Hasegawa 1986, 2002, Akiyama 2009.

Dendroceroscavernosus J.Haseg. MOLUCCAS: Seram: Akiyama 1986, Hasegawa 1986, 2002, Akiyama 2009.

Dendroceroscrispus (Sw.) Nees. MOLUCCAS: Amélio et al. 2021. Ambon: Schiffner 1893b.

Dendrocerosfoliicola J.Haseg. MOLUCCAS: Seram: Akiyama 1986, Hasegawa 1986 as Dendroceros ‘foliicala’, Akiyama 2009.

Dendrocerosjavanicus (Nees) Nees. MOLUCCAS: Seram: Akiyama 1986, 2009, Hasegawa 1986.

Dendrocerospedunculatus Steph. MOLUCCAS: Ambon: LECTOTYPE [Hasegawa 1980: 306], Stephani 1909b, 1917, Bonner 1965, Hasegawa 1980, Piippo 1993a, Peñalosa et al. 2019.

Dendrocerosseramensis J.Haseg. MOLUCCAS: Seram: HOLOTYPE, Akiyama 1986, 2009, Hasegawa 1986, Engel 1993, Crosby and Engel 2006, Peñalosa et al. 2019.

Folioceros D.C.Bharadwaj

Foliocerosamboinensis (Schiffn.) Piippo. MOLUCCAS: Ambon: Schiffner 1890, 1898 both as Anthocerosamboinensis, Sydow 1894 as Anthocerosamboinensis, Stephani 1916 as Aspiromitusamboinensis, Meijer 1954 as Anthocerosamboinensis, Bonner 1962b as Aspiromitusamboinensis and Anthocerosamboinensis, Hasegawa 1993 as Anthocerosamboinensis.

Foliocerosappendiculatus (Steph.) J.Haseg. MOLUCCAS: Seram: Akiyama 1986, Hasegawa 1986, 2002 as Anthocerosappendiculatus, Akiyama 2009.

Foliocerosglandulosus (Lehm. et Lindenb.) D.C.Bharadwaj. MOLUCCAS: Inoue and Miller 1965 as Anthocerosglandulosus, Inoue 1965 as Anthocerosglandulosus, Miller et al. 1983 as Anthocerosglandulosus, Bapna and Kachroo 2000a. Banda: Meijer 1957 as Anthocerosglandulosus. ?Seram: Piippo 1993a. Note: we are not aware of any first hand report from Seram.

Megaceros Campb.

Megacerosflagellaris (Mitt.) Steph. MOLUCCAS: Ambon: Schiffner 1893b, 1898 both as Anthocerosgrandis, Merrill 1906 as Anthocerosgrandis, Miller et al. 1983 as Megacerosgrandis. Seram: Akiyama 1986, 2009, Hasegawa 1986.

Phaeoceros Prosk.

Phaeoceroscarolinianus (Michx.) Prosk. MOLUCCAS: Seram: Akiyama 1986, 2009 both as Phaeoceroslaevissubsp.carolinianus, Hasegawa 1986 as Phaeoceroslaevissubsp.carolinianus.

Phaeomegaceros R.J.Duff, J.C.Villarreal, Cargill et Renzaglia

Phaeomegaceroshirticalyx (Steph.) R.J.Duff, J.C.Villarreal, Cargill et Renzaglia. MOLUCCAS: Ambon: Hasegawa 1986 as Phaeocerospolyandrus, Campbell and Hasegawa 1993 as Phaeoceroshirticalyx, Akiyama 2009 as Phaeocerospolyandrus.

Marchantiophyta

Acrolejeunea (Spruce) Steph.

Acrolejeuneafertilis (Reinw., Blume et Nees) Schiffn. MOLUCCAS: LECTOTYPE of Ptychocoleustener [Gradstein 1975: 86], Bapna and Kachroo 2000b, Singh et al. 2010. Ambon: Sande Lacoste 1864 as Phragmicomafertilis, Schiffner 1898, Gradstein 1975.

Acrolejeuneapycnoclada (Taylor) Schiffn. MOLUCCAS: Mizutani 1977, Bapna and Kachroo 2000b, Haerida et al. 2010, Siregar et al. 2014, Siregar 2015. Ambon: Verdoorn 1934a, 1934b both as Ptychocoleuspycnocladus, Swanson and Miller 1969 as Ptychocoleuspycnocladus, Kamimura 1974 as Ptychocoleuspycnocladus, Miller et al. 1983 also as Ptychocoleuspycnocladus. Seram: Mizutani 1986a, Akiyama 2009.

— subsp. pycnoclada. MOLUCCAS: Ambon: LECTOTYPE of Acrolejeunearostrata [Gradstein 1975: 109], LECTOTYPE of Ptychocoleusbrunneus [Gradstein 1975: 109 as “holotype”], ORIGINAL MATERIAL of Acrolejeunearostratavar.major, ORIGINAL MATERIAL of Acrolejeunearostratavar.minor, Schiffner 1890 as Acrolejeunearostrata and its var. major and var. minor, Bonner 1962b as Acrolejeunearostrata and its var. Major and var. minor, Gradstein 1975, Miller et al. 1983 as Ptychocoleusbrunneus, Geissler and Bischler 1987 as Lejeunearostrata var. Minor, Geissler and Bischler 1989 as Ptychocoleusbrunneus.

Acrolejeuneasecurifolia (Nees) Steph. MOLUCCAS: Seram: Mizutani 1986a as Acrolejeuneasecurifolia, Akiyama 2009.

— subsp. hartmannii (Steph.) Gradst. MOLUCCAS: Ambon: Gradstein 1975, Bonner 1977, Gradstein et al. 2002, Piippo et al. 2002.

Acromastigum A.Evans

Acromastigumaurescens A.Evans. MOLUCCAS: Ambon: Grolle 1978.

Acromastigumbancanum (Sande Lac.) A.Evans. MOLUCCAS: Ambon: Grolle 1978 as Acromastigumbidenticulatum.

Acromastigumdivaricatum (Nees) A.Evans ex Reimers. MOLUCCAS: Ambon: Schiffner 1893b, 1898 both as Bazzaniadivaricata, Evans 1934, Brown and Renner 2014. Seram: Akiyama 1986, 2009.

Acromastigumechinatiforme (De Not.) A.Evans. MOLUCCAS: Ambon: Schiffner 1893b, 1898 as Bazzaniaechinatiformis, Stephani 1909a as Mastigobryumechinatiforme, Evans 1934, Bonner 1962b, 1963a as Bazzaniaechinatiformis, Grolle 1965, 1978, Pócs 1971, Miller et al. 1983, Piippo 1991, Schuster 2000a. Seram: Grolle 1978, Miller et al. 1983, Piippo 1991.

Acromastiguminaequilaterum (Lehm. et Lindenb.) A.Evans. MOLUCCAS: Ambon: Evans 1934, Jovet-Ast 1958, Bonner 1962b, Sharma and Srivastava 1993, Schuster 2000a.

Acromastigumlobuliferum A.Evans. MOLUCCAS: Ambon: Grolle 1978.

Acroscyphella N.Kitag. et Grolle

Acroscyphellatjiwideiensis (Sande Lac.) N.Kitag. et Grolle. MOLUCCAS: Piippo 1985 as Clasmatocoleatjiwideiensis. Ambon: LECTOTYPE of Chiloscyphusfalcifolius [Bonner 1963b: 733], Stephani 1907c as Chiloscyphusfalcifolius, Bonner 1963b as Chiloscyphusfalcifolius, Grolle 1984 as Clasmatocoleatjiwideiensis, 1986, Enroth 1991 as Clasmatocoleatjiwideiensis.

Allorgella Tixier

Allorgellasemperiana (Steph.) Bechteler, G.E.Lee, Schäf.-Verw. et Heinrichs. MOLUCCAS: Seram: Mizutani 1986a as Otolejeuneasemperiana, Akiyama 2009 as Otolejeuneasemperiana.

Allorgellazantenii (Grolle) Bechteler, G.E.Lee, Schäf.-Verw. et Heinrichs. MOLUCCAS: Seram: Bi et al. 2019.

Anastrophyllopsis (R.M.Schust.) Váňa et L.Söderstr.

Anastrophyllopsisrevoluta (Steph.) Váňa et L.Söderstr. MOLUCCAS: Ah-Peng et al. 2010 as Anastrophyllumrevolutum. Seram: Akiyama 1986 as Anastrophyllumrevolutum, Váňa and Piippo 1989b as Anastrophyllumrevolutum, Váňa 1991b as Anastrophyllumrevolutum, Akiyama 2009 as Anastrophyllumrevolutum.

Anastrophyllum (Spruce) Steph.

Anastrophyllumpiligerum (Nees) Steph. MOLUCCAS: Váňa and Engel 2013. Halmahera: Váňa and Piippo 1989b, Váňa 1991b.

Aneura Dumort.

Aneuraamboinensis Steph. MOLUCCAS: Ambon: LECTOTYPE [Bonner 1962b: 86], Stephani 1899b, Bonner 1962b.

Aneurapinguis (L.) Dumort. MOLUCCAS: Ambon: Schiffner 1893b as Aneurapinguisf.normalis, 1898 as Riccardiapinguis.

Balantiopsis Mitt.

BalantiopsisciliarisS.Hatt.subsp.novoguineensis S.Hatt. MOLUCCAS: Seram: Akiyama 1986, 2009.

Bazzania Gray

Bazzaniaasymmetrica (Steph.) N.Kitag. MOLUCCAS: Seram: Herzog 1926b as Mastigobryumasymmetricum.

Bazzaniacadens N.Kitag. MOLUCCAS: Seram: Akiyama 1986, 2009.

Bazzaniacincinnata (De Not.) Trevis. MOLUCCAS: Seram: Akiyama 1986, 2009.

Bazzaniacommutata (Lindenb. et Gottsche) Schiffn. MOLUCCAS: Seram: Akiyama 1986, 2009.

Bazzaniaconophylla (Sande Lac.) Schiffn. MOLUCCAS: Ambon: LECTOTYPE of Mastigobryumlongidens [Kitagawa 1973: 263 as “type”], Stephani 1886a, 1886b, 1908c all as Mastigobryumlongidens, Schiffner 1893b, 1898 as Bazzanialongidens, Bonner 1963a as Bazzanialongidens, Kitagawa 1973. Seram: Akiyama 1986, 2009.

Bazzaniacrassitexta Steph. MOLUCCAS: Ambon: LECTOTYPE [Bonner 1963a: 340], Stephani 1893c, 1908c as Mastigobryumcrassitextum, Schiffner 1898, Bonner 1963a, Kitagawa 1979b.

Bazzaniadistans (Nees) Trevis. MOLUCCAS: Seram: Akiyama 1986, 2009.

Bazzaniaerosa (Reinw., Blume et Nees) Trevis. MOLUCCAS: Del Rosario 1975b. Halmahera: Meijer 1960, Kitagawa 1978, Miller et al. 1983, Meagher 2015, Lestari and Ariyanti 2017, Siregar et al. 2018, Haerida et al. 2023. Seram: Akiyama 1986, 2009.

Bazzaniafallax (Sande Lac.) Schiffn. MOLUCCAS: Ambon: Stephani 1908c as Mastigobryumfallax.

Bazzaniahorridula Schiffn. MOLUCCAS: Geissler and Bischler 1985 as Mastigobryumhorridulum. Ambon: LECTOTYPE [Kitagawa 1967: 254], Schiffner 1893b, 1898, Stephani 1908d as Mastigobryumhorridulum, Bonner 1963a, Kitagawa 1967, Grolle 1968b, Del Rosario 1975b, He et al. 2013, Cheah and Yong 2016, Lestari and Ariyanti 2017.

Bazzaniainsignis (De Not.) Trevis. MOLUCCAS: Ambon: Schiffner 1893b, 1898, Stephani 1908c as Mastigobryuminsigne, Evans 1932, Del Rosario 1975b. Seram: Akiyama 1986, 2009.

Bazzaniaintermedia (Gottsche et Lindenb.) Trevis. MOLUCCAS: Ambon: Schiffner 1893b, 1898, Stephani 1908c as Mastigobryumintermedium, 1908d as Mastigobryumconcinnum, Bonner 1963a also as Bazzaniaconcinna, Inoue and Miller 1965, Pócs et al. 1967, Pócs 1968, Tixier 1971, Miller et al. 1983 as Bazzaniaconcinna, Bapna and Kachroo 2000a. ?Banda: Pócs 1968. Note: we are not aware of any first hand report from Banda,

— var. sarawakiana (De Not.) Schiffn. MOLUCCAS: ?Ambon: Bonner 1963a. Note: we are not aware of any first hand report of this variety from Moluccas and we are do not know if it is worth recognizing.

Bazzaniairregularis (Steph.) Schiffn. MOLUCCAS: Ambon: LECTOTYPE of Mastigobryumirregulare [to be designated], Stephani 1886b, 1908c both as Mastigobryumirregulare, Schiffner 1898, Bonner 1963a.

Bazzaniajavanica (Sande Lac.) Schiffn. MOLUCCAS: Khotimperwati et al. 2018. Seram: Sande Lacoste 1864 as Mastigobryumjavanicum, Schiffner 1898, Stephani 1908e as Mastigobryumjavanicum, Meijer 1960, Kitagawa 1967, Del Rosario 1975b, Miller et al. 1983, Meagher 2015.

Bazzaniakernii Steph. MOLUCCAS: Seram: Akiyama 1986, 2009.

Bazzaniakokawana N.Kitag. et T.Kodama. MOLUCCAS: Seram: Akiyama 1986, 2009.

Bazzanialongicaulis (Sande Lac.) Schiffn. MOLUCCAS: Buru: LECTOTYPE of Mastigobryumdeningeri [here designated], Herzog 1921 as Mastigobryumdeningeri, Herzog 1926a as Mastigobryumdeningeri. Seram: Akiyama 1986, 2009.

Bazzanialoricata (Reinw., Blume et Nees) Trevis. MOLUCCAS: Kitagawa 1967, Siregar and Pasaribu 2019. Seram: Akiyama 1986, 2009.

Bazzaniamerrillana (Steph.) Inoue ex Bonner. MOLUCCAS: ?Buru: Herzog 1926a as Mastigobryummerrillanum. Seram: Herzog 1921 as Mastigobryum ‘Merillanum’. Note: we are not aware of any first hand report from Buru.

Bazzaniaparvitexta Steph. MOLUCCAS: Seram: Akiyama 1986, 2009.

Bazzaniapectinata (Lindenb. et Gottsche) Schiffn. MOLUCCAS: Siregar and Pasaribu 2019. Ambon: Schiffner 1893b, 1898, Stephani 1908c as Mastigobryumpectinatum, Evans 1932, Bonner 1963a, 1977, Sari 2014, Khotimperwati et al. 2018. Seram: Akiyama 1986, 2009.

Bazzaniapraerupta (Reinw., Blume et Nees) Trevis. MOLUCCAS: Khotimperwati et al. 2018. Seram: Meijer 1960, Miller et al. 1983.

Bazzaniaspiralis (Reinw., Blume et Nees) Meijer. MOLUCCAS: Halmahera: Meijer 1960, Miller et al. 1983. Seram: Akiyama 1986, 2009.

Bazzaniastresemannii (Herzog) N.Kitag. MOLUCCAS: Seram: LECTOTYPE of Mastigobryumstresemannii [here designated], Herzog 1921, 1926a both as Mastigobryumstresemannii, Kitagawa 1979b.

Bazzaniasubtilis (Sande Lac.) Trevis. MOLUCCAS: Ambon: Stephani 1908d as Mastigobryumsubtile, Meijer 1960 [with doubt], Bonner 1963a, Del Rosario 1975b, Meagher 2010, Sari 2014, Thouvenot et al. 2018.

Bazzaniatridens(Reinw., Blume et Nees)Trevis.var.tridens. MOLUCCAS: Khotimperwati et al. 2018, Siregar and Pasaribu 2019. Ambon: LECTOTYPE of Mastigobryumamboinense [Kitagawa 1972: 449 as “type”], Stephani 1924 as Mastigobryumamboinense, Kitagawa 1972 as Bazzaniaoshimensis, Miller et al. 1983 as Bazzaniaaustralis and Bazzaniaintermedia, Sharma and Srivastava 1993. ?Banda: Pócs 1969b. Seram: Sande Lacoste 1864 as Mastigobryumtridens, Schiffner 1898, Stephani 1908c as Mastigobryumtridens, Evans 1932, Meijer 1960, Bonner 1963a, Inoue and Miller 1965, Inoue 1965, Mizutani 1967, Hattori 1968, Del Rosario 1975b, Pócs 1971, Kamimura 1975, Miller et al. 1983, Akiyama 1986, Akiyama 2009, 2009 both also as Bazzaniaaustralis, Sharma and Srivastava 1993, Bapna and Kachroo 2000a, Khotimperwati et al. 2018.

Bazzaniauncigera (Reinw., Blume et Nees) Trevis. MOLUCCAS: Ambon: Schiffner 1893b, 1898, Stephani 1908e as Mastigobryumuncigerum, Bonner 1963a, Hattori 1975b, Miller et al. 1983.

Bazzaniavittata (Gottsche) Trevis. MOLUCCAS: ?Ambon: Stephani 1909a as Mastigobryumvittatum, Hattori 1951, Miller et al. 1983, Sari 2014, Siregar et al. 2018. Seram: Akiyama 1986, 2009. Note: we are not aware of any first hand report from Ambon.

Bazzaniawallichiana (Lindenb.) Trevis. MOLUCCAS: Ambon: Stephani 1908d as Mastigobryumwallichianum, Hattori 1951.

Calatholejeunea K.I.Goebel

Calatholejeuneaparadoxa (Schiffn.) K.I.Goebel. MOLUCCAS: Ambon: LECTOTYPE of Lejeuneaparadoxa [Mizutani 1984: 331 as “holotype”], Schiffner 1893b as Lejeuneaparadoxa, 1898 as Colurolejeuneaparadoxa, Stephani 1916 as Coluraparadoxa, Jovet-Ast 1954, Bischler 1961, Bonner 1963a, 1963b as Coluraparadoxa, Grolle 1966a, Schuster 1969, Mizutani 1984, Gradstein 1985, Piippo 1992.

Calypogeia Raddi

Calypogeiagoebelii (Schiffn.) Steph. MOLUCCAS: Seram: Akiyama 1986, 2009.

Caudalejeunea Schiffn.

Caudalejeuneacristiloba (Steph.) Gradst. MOLUCCAS: Mizutani 1977 as Caudalejeuneacircinata. ?Ambon: Thiers and Gradstein 1989, Gradstein et al. 2002, Sass-Gyarmati 2003. Seram: Verdoorn 1934a as Caudalejeuneacircinata, 1934b as Caudalejeuneacircinata, Jovet-Ast 1958 as Caudalejeuneacircinata, Miller et al. 1983 as Caudalejeuneacircinata, Mizutani 1986a as Caudalejeuneacircinata, Akiyama 2009 as Caudalejeuneacircinata. Note: we are not aware of any first hand report from Ambon.

Caudalejeuneareniloba (Gottsche) Steph. MOLUCCAS: Mizutani 1977, Siregar 2015. Ambon: LECTOTYPE of Lopholejeuneasagranavar.dentistipula [here designated], Sande Lacoste 1864 as Phragmicomareniloba, Schiffner 1890 as Lopholejeunea ‘Sagraeana’var.dentistipula, 1898 as Lopholejeuneadentistipula and Thysananthusrenilobus, Sydow 1894 as Lopholejeunea ‘Sagraeana’var.dentistipula, Miller et al. 1983 also as Thysananthusrenilobus, Geissler and Bischler 1987 as Lejeuneasagranavar.dentistipula, Mizutani 1988, Bapna and Kachroo 2000b, Zhu and Gradstein 2005 as Lopholejeuneadentistipula, Siregar et al. 2014, Rosyanti et al. 2018. Seram: Sande Lacoste 1864 as Lejeunearecurvistipula and Phragmicomareniloba, Schiffner 1898 as Caudalejeunearecurvistipula and Thysananthusrenilobus, Verdoorn 1934a, 1934b, Jovet-Ast 1958, Miller 1968, Swanson and Miller 1969, Tixier 1970a, 1970b as Caudalejeunea ‘renilyba’, Miller et al. 1983 also as Thysananthusrenilobus, Mizutani 1986a, Mizutani 1988 also as Caudalejeunearecurvistipula, Bapna and Kachroo 2000b, Akiyama 2009, Siregar et al. 2014, Siregar 2015, Rosyanti et al. 2018, Ginting and Batubara 2019.

Cephalozia (Dumort.) Dumort.

Cephaloziastolonacea (Herzog) Váňa. MOLUCCAS: Ambon: Váňa 1993 as Metahygrobiellastolonacea, Schuster 2002 as Metahygrobiellastolonacea.

Ceratolejeunea (Spruce) J.B.Jack et Steph.

Ceratolejeuneabelangeriana (Gottsche) Steph. MOLUCCAS: Seram: Mizutani 1986a, Zhu et al. 2005, Akiyama 2009.

Ceratolejeuneamoniliata Herzog. MOLUCCAS: Seram: Mizutani 1986a, Akiyama 2009.

Cheilolejeunea (Spruce) Steph.

Cheilolejeuneaceylanica (Gottsche) R.M.Schust. et Kachroo. MOLUCCAS: Mizutani 1978. Bacan: Hoffmann 1935 as Pycnolejeuneaceylanica. Ambon: Schiffner 1893b as Lejeuneaconnivens, 1898 as Pycnolejeuneaceylanica, Tixier 1967b, 1972, 1973b, Kamimura 1974, Miller et al. 1983 also as Xenolejeuneaceylanica, Braggins et al. 2014. Seram: Mizutani 1986a, 2009.

Cheilolejeuneafalsinervis (Sande Lac.) R.M.Schust. et Kachroo. MOLUCCAS: Bacan: Zwickel 1932 as Pycnolejeuneafalsinervis, Hoffmann 1935 as Pycnolejeuneafalsinervis.

Cheilolejeuneagigantea (Steph.) R.M.Schust. et Kachroo. MOLUCCAS: Ambon: LECTOTYPE of Pycnolejeuneagigantea [Thiers 1992: 16 as “holotype”], Stephani 1896b as Pycnolejeuneagigantea, Schiffner 1898 as Pycnolejeuneagigantea, Stephani 1914 as Pycnolejeuneagigantea, Hoffmann 1935 as Pycnolejeuneagigantea, Thiers 1992.

Cheilolejeuneaincisa (Gottsche) R.M.Schust. et Kachroo. MOLUCCAS: Bacan: Hoffmann 1935 as Pycnolejeuneaincisa.

Cheilolejeuneaintertexta (Lindenb.) Steph. MOLUCCAS: Ambon: Grolle 1979b, Mizutani 1982, Miller et al. 1983, Asthana et al. 1995, Zhu and So 1999, Piippo et al. 2002. Seram: Mizutani 1986a, Akiyama 2009.

Cheilolejeunealindenbergii (Gottsche) Mizut. MOLUCCAS: Ambon: LECTOTYPE of Euosmolejeuneaintegristipula [Bonner 1965: 153], Stephani 1896b as Euosmolejeuneaintegristipula, Schiffner 1898 as Euosmolejeuneaintegristipula, Bonner 1965 as Euosmolejeuneaintegristipula, Mizutani 1972, Pócs et al. 1994, Szabó 1997.

Cheilolejeunealongidens (Steph.) R.M.Schust. et Kachroo. MOLUCCAS: Ambon: Thiers 1986 as Pycnolejeuneademissa.

Cheilolejeuneaocclusa (Herzog) T.Kodama et N.Kitag. MOLUCCAS: Seram: Mizutani 1986a, Akiyama 2009.

Cheilolejeuneatrapezia (Nees) Kachroo et R.M.Schust. MOLUCCAS: Ambon: Schiffner 1893b as Lejeuneatrapezia, 1898 as Pycnolejeuneatrapezia, Hoffmann 1935 as Pycnolejeuneameyeniana, Zhu and Grolle 2004. Seram: Mizutani 1986a as Cheilolejeuneameyeniana and Cheilolejeuneaimbricata, Akiyama 2009 as Cheilolejeuneaimbricata and Cheilolejeuneameyeniana.

Cheilolejeuneatrifaria (Reinw., Blume et Nees) Mizut. MOLUCCAS: Mizutani 1977. Ternate/Tidore: LECTOTYPE of Lejeuneaheterophylla [to be designated], Sande Lacoste 1856a, 1856b both as Lejeuneaheterophylla, Schiffner 1898 as Euosmolejeuneaheterophylla, Bonner 1965 as Euosmolejeuneaheterophylla. Ambon: Schiffner 1890 as Euosmolejeuneatrifaria, 1898 as Lejeuneatrifaria, Sydow 1894 as Euosmolejeuneatrifaria, Miller et al. 1983. Seram: Sande Lacoste 1864 as Lejeuneaheterophylla, Schiffner 1898 as Euosmolejeuneaheterophylla, Bonner 1965 as Euosmolejeuneaheterophylla.

Cheilolejeuneaventricosa (Schiffn. ex P.Syd.) Xiao L.He. MOLUCCAS: Furuki and Yamaguchi 2005, Pócs et al. 2013. Ambon: LECTOTYPE of Pycnolejeuneaventricosa [here designated], Schiffner 1890, 1898 both as Pycnolejeuneaventricosa, Sydow 1894 as Pycnolejeuneaventricosa, Stephani 1914 as Pycnolejeuneaventricosa, Zhu et al. 2002, Zhu and Lai 2005, Pócs and Streimann 2006, Renner 2011.

Cheilolejeuneavittata (Steph. ex G.Hoffm.) R.M.Schust. et Kachroo. MOLUCCAS: Seram: Mizutani 1986a, Zhu and So 1999, Zhu et al. 2002, Akiyama 2009.

Chiastocaulon Carl

Chiastocaulonbraunianum (Nees) S.D.F.Patzak, M.A.M.Renner, Schäf.-Verw. et Heinrichs. MOLUCCAS: Halmahera: Sande Lacoste 1864 as Plagiochilabrauniana, Schiffner 1898, 1900 both as Plagiochilabrauniana, Hattori 1952 as Plagiochilion ‘Braunianus’, Miller et al. 1983 as Plagiochilionbraunianum.

Chiastocaulondendroides (Nees) Carl. MOLUCCAS: Jovet-Ast 1951, Grolle 1965. Bacan: Carl 1931a. Ternate/Tidore: New to Ternate: Ternate Piek v. Ternate Oosthelling (Ake abdas), 1951.09.02, D.R. Pleyte 33 (BO11333). Ambon: Stephani 1903a as Plagiochiladendroides, Bonner 1963b, Miller et al. 1983. Seram: Inoue 1986 as Plagiochiladendroides, Akiyama 2009 as Plagiochiladendroides.

Chiastocaulonoppositum (Reinw., Blume et Nees) S.D.F.Patzak, M.A.M.Renner, Schäf.-Verw. et Heinrichs. MOLUCCAS: Ternate/Tidore: LECTOTYPE of Plagiochilaopposita γ filiformis [to be designated], Reinwardt et al. 1824 as Jungermanniaopposita, Lindenberg 1843 as Plagiochilaopposita γ filiformis, Gottsche et al. 1844 as Plagiochilaopposita γ filiformis, Sande Lacoste 1856a as Plagiochilaopposite and Plagiochilaopposita γ filiformis, 1864 as Plagiochilaopposita, Schiffner 1898 as Plagiochilaopposite and Plagiochilaoppositavar.filiformis, Hattori 1942 as Plagiochilaopposita, Inoue 1958 as Noguchiaopposita, Miller et al. 1983 as Plagiochilionoppositum, Zheng et al. 2005 as Plagiochilionoppositum. Ambon: Sande Lacoste 1864 as Plagiochilaopposita, Schiffner 1893b, 1898 both as Plagiochilaopposita, Stephani 1904b as Plagiochilaopposita, Hattori 1942 as Plagiochilaopposita, Inoue 1958 as Noguchiaopposita, Miller et al. 1983 as Plagiochilionoppositum, Bapna and Kachroo 2000b as Plagiochilionoppositum, Zheng et al. 2005 as Plagiochilionoppositum. Seram: Herzog 1921 as Plagiochilaopposita, Carl 1931b as Plagiochilaopposita, Akiyama 1986, Akiyama 2009 both as Plagiochilion ‘oppositus’.

Cladoradula (Spruce) M.A.M.Renner, Gradst., Ilk.-Borg. et F.R.Oliveira-da-Silva

Cladoradulacampanigera (Mont.) M.A.M.Renner, Gradst., Ilk.-Borg. et F.R.Oliveira-da-Silva. MOLUCCAS: Seram: Akiyama 1986, 2009 both as Radulacampanigera, Yamada 1989 as Radulacampanigera.

Cololejeunea (Spruce) Steph.

Cololejeuneaaequabilis (Sande Lac.) Schiffn. MOLUCCAS: Inoue and Miller 1965 as Cololejeuneayulensis, Miller et al. 1983 as Cololejeuneayulensis. Seram: Mizutani 1986a as Cololejeuneayulensis, Akiyama 2009 as Cololejeuneayulensis.

Cololejeuneaangustiflora (Steph.) Mizut. MOLUCCAS: Seram: Mizutani 1986a as Cololejeuneacrenulata and Cololejeuneajavanica, Akiyama 2009 as Cololejeuneajavanica and Cololejeuneacrenulata.

Cololejeuneadozyana (Sande Lac.) Schiffn. MOLUCCAS: Bonner 1963b, Mizutani 1977, Tixier 1980, Pócs and Piippo 2011. Bacan: Benedix 1953. Seram: Mizutani 1986a, Zhu and So 1998b, Zhu and So 1999, Akiyama 2009, Müller et al. 2016.

Cololejeuneaequialbi Tixier. MOLUCCAS: Pócs et al. 2011. Seram: Mizutani 1986a, Pócs et al. 1994, Zhu and So 2001, Eggers 2006, Akiyama 2009 as Cololejeunea ‘equalbi’, Pócs and Piippo 2011, Pócs and Podani 2015.

Cololejeuneafalcata (Horik.) Benedix. MOLUCCAS: Seram: Mizutani 1986a as Cololejeuneafalcatoides, Akiyama 2009 as Cololejeuneafalcatoides.

Cololejeuneafloccosa (Lehm. et Lindenb.) Schiffn. MOLUCCAS: Seram: Mizutani 1986a, Akiyama 2009.

Cololejeuneahamata Steph. MOLUCCAS: Ambon: LECTOTYPE [Bonner 1963b: 830], Stephani 1895, Schiffner 1898, Stephani 1916 as Physocoleahamata, Bonner 1963b.

Cololejeuneahaskarliana (Lehm.) Schiffn. MOLUCCAS: Seram: Mizutani 1986a, Akiyama 2009 as Cololejeunea ‘hasskarliana’.

Cololejeuneainflectens (Mitt.) Benedix. MOLUCCAS: Jovet-Ast and Tixier 1962 as Cololejeuneaciliatilobula, Tixier 1967b, 1971, 1979 all as Cololejeuneaciliatilobula, Mizutani 1975 as Campylolejeuneainflectens, Bapna and Kachroo 2000b as Campylolejeuneainflectens. Ambon: Stephani 1916 as Physocoleaciliatilobula, Benedix 1953 as Cololejeuneaciliatilobula, Bischler 1961 as Campylolejeuneaciliatilobula, Bonner 1963a as Campylolejeuneaciliatilobula, 1963b as Cololejeuneaciliatilobula. Seram: Mizutani 1986a as Cololejeuneapeculiaris, Akiyama 2009 as Cololejeuneapeculiaris.

Cololejeuneakoponenii (Pócs) Pócs. MOLUCCAS: Seram: Bi et al. 2019.

Cololejeunealanciloba Steph. MOLUCCAS: Seram: Mizutani 1986a, Pócs et al. 1994, Akiyama 2009.

Cololejeuneametzgeriopsis (K.I.Goebel) Gradst., R.Wilson, Ilk.-Borg. et Heinrichs. MOLUCCAS: Miller et al. 1983 as Metzgeriopsispusilla. Bacan: Schiffner 1893a as Metzgeriopsispusilla, Schiffner 1893c as Lejeuneapusilla, 1898 as Metzgeriopsispusilla, Stephani 1916 as Metzgeriopsispusilla, Gradstein et al. 2006. Seram: Mizutani 1986a as Metzgeriopsispusilla, Gradstein et al. 2006, Akiyama 2009 as Metzgeriopsispusilla.

Cololejeuneaobliqua (Nees et Mont.) Schiffn. MOLUCCAS: Seram: Mizutani 1986a as Cololejeuneanymannii, Akiyama 2009 as Cololejeunea ‘nymanii’.

Cololejeuneapapillosa (K.I.Goebel) Mizut. MOLUCCAS: Seram: Mizutani 1986a, Akiyama 2009.

Cololejeuneaplagiophylla Benedix. MOLUCCAS: Seram: Mizutani 1986a, Akiyama 2009.

Cololejeuneaplanissima(Mitt.)Abeyw.var.planissima. MOLUCCAS: Seram: Mizutani 1986a as Cololejeuneaplanissima, Akiyama 2009 as Cololejeuneaplanissima.

Cololejeunearaduliloba Steph. MOLUCCAS: Seram: Mizutani 1986a, Akiyama 2009.

Cololejeuneastephanii Schiffn. ex Benedix. MOLUCCAS: Seram: Mizutani 1986a, Akiyama 2009.

Cololejeuneatriapiculata (Herzog) Tixier. MOLUCCAS: Seram: Mizutani 1986a, Akiyama 2009.

Cololejeuneatrichomanis (Gottsche) Besch. MOLUCCAS: Seram: Mizutani 1986a as Cololejeuneagoebelii, Akiyama 2009 as Cololejeuneagoebelii.

Colura (Dumort.) Dumort.

Coluraacroloba (Prantl) Jovet-Ast. MOLUCCAS: Seram: Mizutani 1986a, Eggers et al. 1998, Akiyama 2009.

Coluraamboinensis Steph. MOLUCCAS: Ambon: LECTOTYPE [to be designated], ORIGINAL MATERIAL of Colurolejeuneasuperbavar.typica, Schiffner 1890 as ‘Coluro-Lejeunea’ superba var. typica, Stephani 1916, Bonner 1963b, Miller et al. 1983 as Colurolejeuneasuperba.

Coluraari (Steph.) Steph. MOLUCCAS: Halmahera: Jovet-Ast 1953, Bonner 1963b, Grolle 1965, Pócs et al. 1967, Tixier 1967a, Onraedt 1979, Miller et al. 1983. Seram: Mizutani 1986a, Akiyama 2009.

Coluraconica (Sande Lac.) K.I.Goebel. MOLUCCAS: Seram: Mizutani 1986a, Akiyama 2009.

Coluraimperfecta Steph. MOLUCCAS: Seram: Mizutani 1986a, Akiyama 2009.

Colurakarstenii K.I.Goebel. MOLUCCAS: Ambon: LECTOTYPE [Jovet-Ast 1953: 230, Bonner 1963b: 869], Stephani 1896b as Colurolejeuneakarstenii, 1916, Schiffner 1898 as Colurolejeuneakarstenii, Schiffner and Stephani 1901 as Colurolejeuneakarsteniivar.latifolia, Jovet-Ast 1953, Bonner 1963b, Tixier 1970a, Zhu and So 2001, Pócs 2013, Sangrattanaprasert et al. 2018.

Coluramaxima Jovet-Ast. MOLUCCAS: Halmahera: LECTOTYPE [to be designated], Jovet-Ast 1953, Bonner 1963b.

Coluraornata K.I.Goebel. MOLUCCAS: Pócs and Ninh 2012. Seram: Mizutani 1986a, Pócs et al. 1994, Akiyama 2009.

Colurasuperba (Mont.) Steph. MOLUCCAS: Reiner-Drehwald 1999 as Lejeuneasuperbavar.typica. Ambon: Bonner 1963b as Colurolejeuneasuperba, Geissler and Bischler 1987 as Lejeuneasuperbavar.typica. Seram: Mizutani 1986a, Akiyama 2009.

Conoscyphus Mitt.

Conoscyphustrapezioides (Sande Lac.) Schiffn. MOLUCCAS: Seram: Akiyama 1986, Akiyama 2009.

Cryptolophocolea L.Söderstr., Crand.-Stotl., Stotler et Váňa

Cryptolophocoleacostata (Nees) L.Söderstr. MOLUCCAS: Seram: Akiyama 1986, 2009 both as Lophocoleagiulianettii.

Cuspidatula Steph.

Cuspidatulacontracta (Reinw., Blume et Nees) Steph. MOLUCCAS: Ambon: Schiffner 1893b, 1898 both as Anastrophyllumcontractum, Schiffner 1900 as Anastrophyllumcontractum, Stephani 1901c, Bonner 1962b as Anastrophyllumcontractum, Bonner 1963b, Kitagawa 1970, Grolle 1971, Bizot and Pócs 1974, Miller et al. 1983, Engel and Glenny 2008. Seram: Akiyama 1986 as Jamesoniellacontracta, Akiyama 2009 as Jungermanniacontracta.

Cuspidatulaflaccida (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs. MOLUCCAS: Seram: Akiyama 1986, 2009 both as Anomacaulisflaccidus.

Cuspidatulaflexicaulis (Nees) Váňa et L.Söderstr. MOLUCCAS: Miller et al. 1983 as Jamesoniellaflexicaulis. Ambon: Grolle 1971 as Jamesoniellaflexicaulis. Seram: Akiyama 1986, 2009 both as Jungermanniaflexicaulis.

Denotarisia Grolle

Denotarisialinguifolia (De Not.) Grolle. MOLUCCAS: Ambon: LECTOTYPE of Jungermanniaovifolia [Grolle 1971: 11 as “holotype”], Schiffner 1893b, 1898, 1900 all as Jamesoniellaovifolia, Stephani 1901b as Jamesoniellaovifolia, Grolle 1966a as Jamesoniellaovifolia, 1971, Bonner 1976 as Jungermanniaovifolia, Miller et al. 1983 also as Jamesoniellaovifolia and Jungermanniaovifolia, Váňa and Piippo 1989b, Váňa 1991b. Seram: Akiyama 1986, 2009.

Dinckleria Trevis.

Dinckleriasingularis (Schiffn.) M.A.M.Renner, Schäf.-Verw. et Heinrichs. MOLUCCAS: Ternate/Tidore: Gottsche et al. 1844 as Plagiochilatrapezoidea β tenera, Sande Lacoste 1856a, 1856b both as Plagiochilatrapezoidea β tenera, Schiffner 1898 as Plagiochilatrapezoideavar.tenera. Seram: So 2000a, 2001a, 2001b all as Plagiochilasingularis, Renner et al. 2016.

Diplasiolejeunea (Spruce) Schiffn.

Diplasiolejeuneacavifolia Steph. MOLUCCAS: Seram: Mizutani 1986a, Akiyama 2009.

Diplasiolejeuneapatelligera Herzog. MOLUCCAS: Seram: Mizutani 1986a, Akiyama 2009.

Drepanolejeunea (Spruce) Steph.

DrepanolejeuneablumeiSteph.var.angustistipa Herzog. MOLUCCAS: Bonner 1965. Bacan: Herzog 1936.

Drepanolejeuneadactylophora (Nees, Lindenb. et Gottsche) J.B.Jack et Steph. MOLUCCAS: Mizutani 1961 as Drepanolejeuneadactylophora, Inoue 1965, Kamimura 1974, Mizutani 1975 as Drepanolejeuneadactylophora, Mizutani 1977 as Drepanolejeuneadactylophora, Mizutani 1978 as Drepanolejeuneadactylophora. Bacan: Schiffner 1893c. ?Ambon: Stephani 1913 as Drepanolejeuneadactylophora, Tixier 1962 as Drepanolejeuneadactylophora, 1972, Bonner 1965 as Drepanolejeuneadactylophora, Pócs et al. 1967, Miller et al. 1983 as Drepanolejeuneadactylophora, Zhu and So 2001 as Drepanolejeuneadactylophora. Seram: Mizutani 1986a as Drepanolejeuneadactylophora, Zhu and So 2001 as Drepanolejeuneadactylophora, Akiyama 2009. Note: we are not aware of any first hand report from Ambon.

Drepanolejeuneafoliicola Horik. MOLUCCAS: Seram: Mizutani 1986a as Leptolejeuneafoliicola.

Drepanolejeuneagrandis Herzog. MOLUCCAS: LECTOTYPE [to be designated], Herzog 1939, Bonner 1965, Pócs et al. 1995, Pócs et al. 2019a. Seram: Mizutani 1986a, Akiyama 2009.

Drepanolejeuneahampeana Steph. MOLUCCAS: Ambon: Stephani 1913, Bonner 1965, Miller et al. 1983.

Drepanolejeuneaintermedia Zwickel. MOLUCCAS: O’Shea et al. 1997. Bacan: Grolle 1976. Seram: Mizutani 1986a, Akiyama 2009.

Drepanolejeunealevicornua Steph. MOLUCCAS: Seram: Mizutani 1986a as Drepanolejeunealevicornuavar.levicornua, 1990, Streimann 1991, Szabó 1997, Eggers et al. 1998, Zhu and So 2001, Akiyama 2009 as Drepanolejeunealevicornuavar.levicornua, Siregar et al. 2017, Pócs et al. 2019b, Susilo et al. 2023.

Drepanolejeunealongicornua (Herzog) Mizut. MOLUCCAS: Seram: Mizutani 1986a as Drepanolejeunealevicornuavar.longicornua, 1990, Akiyama 2009 as Drepanolejeunealevicornuavar.longicornua.

Drepanolejeunealongicruris (Steph.) Grolle et R.L.Zhu. MOLUCCAS: Seram: Mizutani 1986a as Rhaphidolejeunealongicruris, Grolle and Zhu 2000, Akiyama 2009 as Rhaphidolejeunealongicruris.

Drepanolejeunealyrata Grolle. MOLUCCAS: Seram: Mizutani 1986a, Akiyama 2009.

Drepanolejeuneamoluccensis Herzog. MOLUCCAS: Bacan: LECTOTYPE [Nadhifah et al. 2021: 6], Herzog 1934, Nadhifah et al. 2021.

Drepanolejeuneaobliqua Steph. MOLUCCAS: Pócs et al. 1967. Bacan: Herzog 1939.

Drepanolejeuneapentadactyla (Mont.) Steph. MOLUCCAS: Jovet-Ast 1958 as Drepanolejeuneamicholitzii, Jovet-Ast and Tixier 1962 as Drepanolejeuneamicholitziivar.genuina, Bonner 1965 as Drepanolejeuneamicholitziivar.genuina, Miller 1968 as Drepanolejeuneamicholitzii, Tixier 1970b as Drepanolejeuneamicholitzii, Kitagawa 1981 as Drepanolejeuneamicholitzii, Miller et al. 1983 as Drepanolejeuneamicholitzii. Bacan: ORIGINAL MATERIAL of Drepanolejeuneamicholitziivar.genuina, Herzog 1934 as Drepanolejeuneamicholitziivar.genuina. Seram: Mizutani 1986a as Drepanolejeuneamicholitzii, Zhu and So 2001, Akiyama 2009 as Drepanolejeuneamicholitzii, Asthana and Shukla 2009, Dey et al. 2013, Susilo et al. 2023.

— var. dactylophoroides (Herzog) Pócs. ?MOLUCCAS: Tixier 1979 as Drepanolejeuneamicholitziivar.dactylophoroides. Note: we are not aware of any first hand report of this variety from Moluccas although it is reported to occur on neighbouring islands.

Drepanolejeuneaspinosocornuta Steph. MOLUCCAS: Seram: Mizutani 1986a, 1990, Akiyama 2009, Pócs et al. 2019a.

Drepanolejeuneatenera K.I.Goebel. MOLUCCAS: Jovet-Ast 1958, Bonner 1965 as Drepanolejeuneateneravar.genuina, Pócs 1969a, Tixier 1974b. Bacan: ORIGINAL MATERIAL of Drepanolejeuneateneravar.genuina, Herzog 1934 as Drepanolejeuneateneravar.genuina. Seram: Mizutani 1986a, Akiyama 2009.

Drepanolejeuneaternatensis (Gottsche) Schiffn. MOLUCCAS: Inoue 1965, Miller 1968, Pócs 1969a, Kamimura 1974, Mizutani 1975, 1977, 1978, Udar and Awasthi 1982 as Drepanolejeuneaternatensisvar.ternatensis, Bapna and Kachroo 2000b as Drepanolejeuneaternatensisvar.ternatensis, Xiong and Cao 2018, Siregar et al. 2020a. Bacan: Herzog 1939 as Drepanolejeuneaternatensisvar.ternatensis as Drepanolejeuneaternatensis. Ternate/Tidore: LECTOTYPE of Lejeuneaternatensis [to be designated], Gottsche et al. 1845b as Lejeuneaternatensis, Sande Lacoste 1856a, 1856b both as Lejeuneaternatensis, Stephani 1890b as Lejeuneaternatensis, Schiffner 1898, Mizutani 1961, Miller et al. 1983 as Lejeuneaternatensis, Microlejeuneaternatensis and Drepanolejeuneaternatensis, Zhu and So 2001, Dey et al. 2013. Seram: Mizutani 1986a as Drepanolejeuneaternatensis, Zhu and So 2001, Akiyama 2009, Dey et al. 2013, Siregar et al. 2017, Susilo et al. 2023.

Drepanolejeuneathwaitesiana(Mitt.)Steph.var.thwaitesiana. MOLUCCAS: Seram: Mizutani 1990, Zhu and So 2001, Haerida 2009, Siregar et al. 2017, Susilo et al. 2023.

Drepanolejeuneatricornua Herzog. MOLUCCAS: Seram: LECTOTYPE [Bonner 1965: 105], Herzog 1936, Bonner 1965, Tixier 1979, Mizutani 1990, Pócs et al. 2011, Pócs et al. 2013, Pócs et al. 2014, Siregar et al. 2017, Samarakkody et al. 2018, Kasiani et al. 2019.

Dumortiera Nees

Dumortierahirsuta (Sw.) Nees. MOLUCCAS: Ambon: Sopacua et al. 2020 as Dumortiera ‘hirsute’ [Saparua I]. Note: we are not aware of any first hand report from Moluccas although it is reported to occur on neighbouring islands.

Frullania Raddi

Frullaniaakiyamae S.Hatt. MOLUCCAS: Seram: HOLOTYPE, Hattori 1986b, Engel 1993, Crosby and Engel 2006, Akiyama 2009.

Frullaniaapiculata (Reinw., Blume et Nees) Nees. MOLUCCAS: Verdoorn 1930, Inoue and Miller 1965, Inoue 1965, Pócs et al. 1967, Miller 1968, Swanson and Miller 1969, Tixier 1972, Haerida et al. 2023. Halmahera: Sande Lacoste 1864, Schiffner 1898, Umagap 2019. Ambon: Schiffner 1890, 1898, Sydow 1894. Seram: Sande Lacoste 1864, Schiffner 1898, Hattori 1986b, Akiyama 2009.

— var. apiculata. MOLUCCAS: Ambon: Schiffner 1893b, Miller et al. 1983. Seram: Miller et al. 1983.

— var. goebelii Schiffn. MOLUCCAS: Tixier 1971. Ambon: LECTOTYPE of Frullaniakarstenii [to be designated], Schiffner 1893b, 1898 both as Frullaniakarstenii, Stephani 1911 as Frullaniakarstenii, Bonner 1965 as Frullaniakarstenii.

Frullaniaarmatifolia Verd. MOLUCCAS: Seram: Hattori 1986b, Akiyama 2009.

Frullaniaattenuata Steph. MOLUCCAS: Seram: Hattori 1986b, Akiyama 2009.

Frullaniaberthoumieui Steph. MOLUCCAS: Ambon: Hattori 1975a, 1976, Hattori et al. 1977, Bapna and Kachroo 2000b.

Frullaniaclaviloba Steph. MOLUCCAS: Ambon: LECTOTYPE [Bonner 1965: 266], Stephani 1911, Verdoorn 1930, Bonner 1965, Pócs et al. 1967, Kamimura 1974, Hattori 1975a, 1975d, 1976, 1982, Hattori et al. 1977, Sukkharak 2018.

Frullaniacordistipula (Reinw., Blume et Nees) Nees. MOLUCCAS: Halmahera: Sande Lacoste 1864, Schiffner 1898, Merrill and Merritt 1910, Hattori 1977, Miller et al. 1983.

— var. dentistipula S.Hatt. MOLUCCAS: Seram: HOLOTYPE, Hattori 1986b, Engel 1993, Crosby and Engel 2006, Akiyama 2009.

Frullaniaericoides (Nees) Mont. MOLUCCAS: Verdoorn 1930 as Frullaniasquarrosavar.planescens, Bonner 1965 as Frullaniasquarrosaf.campanuloides, Bai 2002 as Frullaniaericoidesvar.planescens, Haerida et al. 2023. Seram: Sande Lacoste 1864 as Frullaniasquarrosa, Schiffner 1898 as Frullaniasquarrosa, Miller et al. 1983 as Frullaniasquarrosa, Hattori 1986b, Akiyama 2009.

Frullaniafallax Gottsche. MOLUCCAS: Halmahera: Sande Lacoste 1864, Schiffner 1898, Hattori 1975a as Frullaniafallaxvar.fallax.

Frullaniagaudichaudii (Nees et Mont.) Nees et Mont. MOLUCCAS: Ambon: Schiffner 1893b, Schiffner and Stephani 1901, Verdoorn 1930, Bonner 1965, Swanson and Miller 1969, Kamimura 1971, Miller et al. 1983, Haerida et al. 2023. Seram: Hattori 1986b, Akiyama 2009.

Frullaniagracilis (Reinw., Blume et Nees) Nees. MOLUCCAS: Ambon: LECTOTYPE of Frullanialacerifolia [Bonner 1965: 345], Stephani 1911 as Frullanialacerifolia, Verdoorn 1930 as Frullaniagracilisvar.lacerifolia, Bonner 1965 as Frullaniagracilisvar.lacerifolia and Frullanialacerifolia, Kamimura 1974, Miller et al. 1983. Seram: Hattori 1986b.

— var. gracilis. MOLUCCAS: Seram: Akiyama 2009.

— var. vittata S.Hatt. MOLUCCAS: Seram: HOLOTYPE, Hattori 1986b, Engel 1993, Crosby and Engel 2006, Akiyama 2009.

Frullaniagrandistipula Lindenb. MOLUCCAS: Bacan: Verdoorn 1930. Halmahera: Sande Lacoste 1864 as Frullaniarugosa, Schiffner 1898 as Frullaniarugosa, Pócs et al. 1967, Hattori 1974b, 1975a, 1976 as Frullaniagrandistipulavar.grandistipula, 1979, 1982, Miller et al. 1983, Hattori and Piippo 1986.

Frullaniahasskarliana Lindenb. MOLUCCAS: Haerida et al. 2023.

— var. gracilis S.Hatt. MOLUCCAS: Seram: HOLOTYPE, Hattori 1986b, Engel 1993, Crosby and Engel 2006, Akiyama 2009.

— var. hasskarliana. MOLUCCAS: Seram: Hattori 1986b, Akiyama 2009.

— var. parvidentata S.Hatt. MOLUCCAS: Seram: HOLOTYPE, Hattori 1986b, Engel 1993, Crosby and Engel 2006, Akiyama 2009.

Frullaniaheteromorpha Schiffn. MOLUCCAS: Seram: Hattori 1986b, Akiyama 2009.

Frullaniaintegristipula (Nees) Nees

— var. emarginata Verd. MOLUCCAS: Halmahera: Verdoorn 1930, Hattori 1980, Hattori 1982, Hattori and Piippo 1986.

— var. integristipula. MOLUCCAS: Halmahera: Hattori 1980 as Frullaniaintegristipula.

Frullaniaintermedia (Reinw., Blume et Nees) Nees. MOLUCCAS: Inoue and Miller 1965, Bapna and Kachroo 2000b. Seram: Miller et al. 1983.

— subsp. intermedia. MOLUCCAS: Ambon: LECTOTYPE of Frullaniaamboinensis [Bonner 1965: 228], LECTOTYPE of Frullaniabillardiereana [Bonner 1965: 248], Montagne 1843, 1856 both as Frullaniabillardiereana, Gottsche et al. 1845b as Frullaniabillardiereana, Schiffner 1890, 1898 both as Frullaniaamboinensis, 1898 as Frullaniabillardiereana, Sydow 1894 as Frullaniaamboinensis, Schiffner and Stephani 1901 as Frullaniaamboinensis, Stephani 1911 as Frullaniaamboinensis, Verdoorn 1930 as Frullaniaintermedia f. ‘billardieriana’, Verdoorn 1930 as Frullaniaintermediavar.amboinensis, Bonner 1965 as Frullaniaintermedia f. ‘billardierianaa’, Bonner 1965 as Frullaniaintermediavar.amboinensis, Frullaniaamboinensis and Frullaniabillardiereana, Hattori 1976 as Frullaniaintermedia f. ‘billardierianaa’, 1980 as Frullaniabillardiereana, 1982, 1986 both as Frullaniaintermedia f. ‘billardierianaa’, Miller et al. 1983 as Frullaniaintermedia f. ‘billardierana’. Seram: Sande Lacoste 1864 as Frullaniabillardiereana, Schiffner 1898 as Frullaniabillardiereana, Verdoorn 1930 as Frullaniaintermedia f. ‘billardierianaa’, Bonner 1965 as Frullaniaintermedia f. ‘billardierianaa’, Hattori 1976 as Frullaniaintermedia f. ‘billardieriana’, 1980 as Frullaniabillardiereana, 1982, 1986 both as Frullaniaintermedia f. ‘billardieriana’, Miller et al. 1983 as Frullaniaintermedia f. ‘billardierana’.

Frullaniajunghuhniana Gottsche

— var. junghuhniana. MOLUCCAS: Seram: Hattori 1986b, Akiyama 2009.

— var. tenella (Sande Lac.) Grolle et S.Hatt. MOLUCCAS: Seram: Hattori 1986b, Akiyama 2009.

Frullaniamultilacera Steph. MOLUCCAS: Seram: Hattori 1986b, Akiyama 2009.

Frullanianepalensis (Spreng.) Lehm. et Lindenb. MOLUCCAS: Bacan: Verdoorn 1930, Hattori 1952, 1973b, Kamimura 1961, Pócs et al. 1967.

Frullanianigricaulis (Reinw., Blume et Nees) Nees. MOLUCCAS: Halmahera: Verdoorn 1930, Hattori 1974b, 1975c, Hattori et al. 1977.

FrullanianobilisSteph.var.cochleata (Steph.) S.Hatt. MOLUCCAS: Seram: Hattori 1986b, Akiyama 2009.

Frullanianodulosa (Reinw., Blume et Nees) Nees. MOLUCCAS: Verdoorn 1930 as Frullanianodulosa β plana, Frullanianodulosaf.dapitana and Frullanianodulosaf.irreflexa, Kamimura 1943 as Frullanianodulosaf.dapitana, Bonner 1965 as Frullanianodulosaf.dapitana, Bonner 1965 as Frullanianodulosaf.irreflexa, Miller et al. 1983 as Frullaniadapitana and Frullanianodulosaf.irreflexa, Haerida et al. 2023. Morotai: Hattori 1975a, Hattori 1980 as Frullanianodulosavar.nodulosa, Miller et al. 1983. Ambon: LECTOTYPE of Frullanianodulosa β plana [to be designated], Sande Lacoste 1864 also as Frullaniathuillieri, Schiffner 1890 as Frullanianodulosavar.nodulosa and Frullanianodulosa β plana, 1893b, 1898 also as Frullaniareplicate and Frullanianodulosa β plana, Sydow 1894 as Frullanianodulosa β plana, Verdoorn 1930, 1932 as Frullanianodulosaf.irreflexa, Hattori 1951, Bonner 1965 as Frullanianodulosa β plana, Swanson and Miller 1969, Hattori 1975a, 1980 as Frullanianodulosavar.nodulosa, Miller et al. 1983. Seram: Sande Lacoste 1864, Verdoorn 1930, Hattori 1951, Swanson and Miller 1969, Hattori 1980 as Frullanianodulosavar.nodulosa, 1986b, Miller et al. 1983, Akiyama 2009.

Frullaniaorientalis Sande Lac. MOLUCCAS: Bapna and Kachroo 2000b, Haerida et al. 2023. Seram: Hattori 1986b, Akiyama 2009.

Frullaniaornithocephala (Reinw., Blume et Nees) Nees. MOLUCCAS: Haerida 2015. Ambon: Merrill and Merritt 1910, Verdoorn 1930, Bonner 1965, Hattori 1974c, 1975c, 1976, Hattori et al. 1977. Seram: Verdoorn 1930, Bonner 1965, Hattori 1974c, 1975c, 1976, Hattori et al. 1977.

— var. major (Nees) Schiffn. MOLUCCAS: Ambon: Schiffner 1893b, 1898.

— var. ornithocephala. MOLUCCAS: Ambon: Hattori 1974a, 1975a. Seram: Hattori 1974a, 1975a as Frullaniaornithocephala.

Frullaniapauciramea Steph. MOLUCCAS: Seram: Hattori 1986b, Akiyama 2009.

FrullaniapiptophyllaS.Hatt.var.minor S.Hatt. MOLUCCAS: Seram: HOLOTYPE, Hattori 1986b, Engel 1993, Crosby and Engel 2006, Akiyama 2009.

Frullaniapulogensis Steph. MOLUCCAS: Seram: Hattori 1986b, Akiyama 2009.

Frullaniaramuligera (Nees) Mont. MOLUCCAS: Seram: Hattori 1986b, Akiyama 2009.

FrullaniareflexistipulaSande Lac.var.reflexistipulaMOLUCCAS: Haerida et al. 2023. Seram: Sande Lacoste 1864, Schiffner 1898, Verdoorn 1930, Hattori 1973b, 1974a, 1974c, 1975a, 1976, 1982, 1986b, Miller et al. 1983, Hattori and Piippo 1986, Akiyama 2009.

Frullaniarepandistipula Sande Lac. MOLUCCAS: Ambon: Stephani 1911, Verdoorn 1930 [rejecting Stephani’s reports], Bonner 1965, Hattori 1973a [rejecting earlier reports], Bizot et al. 1978. Seram: Hattori 1986b, Akiyama 2009. Note: the only report from Ambon is rejected as Frullaniatricarinata by Verdoorn (1930).

Frullaniaserrata Gottsche. MOLUCCAS: Pócs 1971, Hattori 1974a, Miller et al. 1983. Bacan: Verdoorn 1930.

— var. ceramensis S.Hatt. MOLUCCAS: Seram: HOLOTYPE, Hattori 1986b, Engel 1993, Crosby and Engel 2006, Akiyama 2009.

— var. hamatispina (S.Hatt.) S.Hatt. MOLUCCAS: Seram: Hattori 1986b, Akiyama 2009.

— var. serrata. MOLUCCAS: Seram: Hattori 1986b as Frullaniaserrataf.crispulo-dentata, Akiyama 2009 as Frullaniaserrataf.crispulo-dentata.

Frullaniasubcaduca S.Hatt. MOLUCCAS: Seram: Hattori 1986b, Akiyama 2009.

Frullaniasublignosa Steph. MOLUCCAS: Seram: Hattori 1986b, Akiyama 2009.

FrullaniasubnigricaulisS.Hatt.var.subtruncata S.Hatt. MOLUCCAS: Seram: Hattori 1986b, Akiyama 2009.

Frullaniasubocellata S.Hatt. MOLUCCAS: Seram: HOLOTYPE, Hattori 1986b, 1988, Engel 1993, Crosby and Engel 2006, Akiyama 2009, Pócs et al. 2014.

Frullaniaternatensis Gottsche. MOLUCCAS: Haerida 2015, Haerida et al. 2023. Bacan: Verdoorn 1930, Tixier 1962, Kamimura 1974. Ternate/Tidore: LECTOTYPE [Bonner 1965: 451], Gottsche et al. 1846, Sande Lacoste 1856a, 1856b, Schiffner 1898, Verdoorn 1930, Tixier 1962, 1970a, 1971, 1972, Kamimura 1974, Miller et al. 1983. Ambon: Verdoorn 1932. Seram: Verdoorn 1932, Miller et al. 1983.

— var. non-appendiculata S.Hatt. MOLUCCAS: Seram: Hattori 1986b, Akiyama 2009.

— var. ternatensis. MOLUCCAS: Ternate/Tidore: Hattori 1974a. Seram: Hattori 1975a.

Frullaniatricarinata Sande Lac. MOLUCCAS: Ambon: Schiffner 1893b, 1898, Verdoorn 1930.

Frullaniatrichodes Mitt. MOLUCCAS: Ambon: LECTOTYPE of Frullaniapicta [Bonner 1965: 400; here affirmed], Stephani 1894a as Frullaniapicta, Schiffner 1898 as Frullaniapicta, Verdoorn 1930 as Frullaniapicta, Svihla 1955, 1959 both as Frullaniapicta, Kamimura 1961, 1974 both as Frullaniatenuicaulis, Arnell 1965 as Frullaniapicta, Bonner 1965 as Frullaniapicta, Kitagawa 1969 as Frullaniapicta, Tixier 1973b as Frullaniatenuicaulis, 1974b as Frullaniapicta.

Frullaniavaginata (Sw.) Nees. MOLUCCAS: Ternate/Tidore: Reinwardt et al. 1824 as Jungermanniavaginata, Nees 1830 as Jungermanniavaginata. Ambon: Sande Lacoste 1864 [Saparoea I], Verdoorn 1930, Hattori 1975a, 1975c,1982, Hattori and Piippo 1986.

Gottschelia Grolle

Gottscheliaschizopleura (Spruce) Grolle. MOLUCCAS: Váňa and Piippo 1989b, Váňa 1991b, Schuster 2002, Long and Váňa 2007. Halmahera: Sande Lacoste 1864 as Jungermanniaflexicaulis β microphylla. Ternate/Tidore: Schiffner 1898 as Jamesoniellaflexicaulisvar.microphylla, 1900 as Jamesoniellamicrophylla, Stephani 1901b as Jamesoniellamicrophylla, Bonner 1966 as Jamesoniellamicrophylla, Grolle 1968a, Bizot and Pócs 1974, Miller et al. 1983. Seram: Sande Lacoste 1864 as Jungermanniaflexicaulis β microphylla, Schiffner 1898 as Jamesoniellaflexicaulisvar.microphylla, 1900 as Jamesoniellamicrophylla, Stephani 1901b as Jamesoniellamicrophylla, Bonner 1966 as Jamesoniellamicrophylla, Grolle 1968a, Miller et al. 1983, Bapna and Kachroo 2000b.

Hattoriolejeunea Mizut.

Hattoriolejeuneaakiyamae Mizut. MOLUCCAS: Seram: HOLOTYPE, Mizutani 1986a, Engel 1993, Crosby and Engel 2006, Akiyama 2009.

Herbertus Gray

Herbertuslongifissus Steph. MOLUCCAS: Seram: Akiyama 1986, 2009.

Herbertuslongispinus J.B.Jack et Steph. MOLUCCAS: Ambon: Schiffner 1893b as ‘Herbertalongispina’ [but represents Herbertuspilifer], 1898 as “Herbertalongispina’ [but citing only the type of Herbertuspilifer]. Seram: Akiyama 1986, 2009.

Herbertuspilifer (Steph.) H.A.Mill. MOLUCCAS: Ambon: LECTOTYPE of Schismapilifera [Miller 1965: 327], Stephani 1909c as Schismapilifera, Miller 1965 as ‘Herbertapilifera’, Miller et al. 1983 as Herbertuspilifer and Schismapiligerum, Piippo 1984a as Herbertuspilifer, So 2003b as Herbertuspilifer, Juslén 2006 as Herbertuspilifer.

Herbertusramosus (Steph.) H.A.Mill. MOLUCCAS: Seram: Akiyama 1986, 2009 both as Herbertusjavanicus.

Herbertussendtneri (Nees) Lindb. MOLUCCAS: Seram: LECTOTYPE of Schismadivaricatum [Miller 1965: 325 as “type”], Herzog 1921, 1926a both as Schismadivaricatum, Miller 1965 as ‘Herbertadivaricata’, Del Rosario 1975a as ‘Herbertadivaricata’, Piippo 1984a as Herbertusarmitanus, Akiyama 1986, 2009 both as Herbertusarmitanus, Gao 2003 as Herbertusdivaricatus, So 2003b as Herbertusarmitanus, Juslén 2006 as Herbertusarmitanus.

Heteroscyphus Schiffn.

Heteroscyphusamboinensis (Schiffn.) Schiffn. MOLUCCAS: Ambon: LECTOTYPE of Chiloscyphusendlicherianus δ amboinensis [to be designated], Schiffner 1890 as Chiloscyphusendlicherianus δ amboinensis, 1898 as Chiloscyphusamboinensis, Stephani 1907 as Chiloscyphusamboinensis, Bonner 1963b as Chiloscyphusdecurrensvar.amboinensis [error for C.endlicherianus δ amboinensis] and Chiloscyphusamboinensis, Miller et al. 1983 as Chiloscyphusendlicherianus δ amboinensis.

Heteroscyphusargutus (Reinw., Blume et Nees) Schiffn. MOLUCCAS: Seram: Akiyama 1986, 2009.

Heteroscyphusaselliformis (Reinw., Blume et Nees) Schiffn. MOLUCCAS: Ambon: Schiffner 1893b, 1898 both as Chiloscyphusaselliformis, Stephani 1907 as Chiloscyphusaselliformis, Bonner 1963b as Chiloscyphusaselliformis and Gamoscyphusaselliformis, Grolle 1965 as Chiloscyphusaselliformis, Inoue 1965, Bonner 1966, 1976 as Jungermanniaaselliformis, Kitagawa 1979a as Chiloscyphusaselliformis, Miller et al. 1983, Piippo 1985, 1989b, Yamada and Hayashi 2003. Seram: Akiyama 1986, 2009.

Heteroscyphuscoalitus(Hook.)Schiffn.var.coalitusMOLUCCAS: Ambon: Sande Lacoste 1864 as Chiloscyphuscoalitus, Schiffner 1890, 1898 both as Chiloscyphuscoalitus, Sydow 1894 as Chiloscyphuscoalitus, Inoue and Miller 1965 as Heteroscyphus ‘coaltitus’, Miller et al. 1983, Piippo 1985, 1989b, 1993b. Seram: Akiyama 1986, 2009.

Heteroscyphussplendens (Lehm. et Lindenb.) Grolle. MOLUCCAS: Halmahera: Sande Lacoste 1864 as Chiloscyphusdecurrens, De Notaris 1874 as Chiloscyphusdecurrens, Schiffner 1898 as Chiloscyphusdecurrens, Pócs 1971 as Heteroscyphusdecurrens, Miller et al. 1983 as Heteroscyphusdecurrens. Ambon: Schiffner 1893b, 1898 both as Chiloscyphusdecurrens, Grolle 1965 as Chiloscyphusdecurrens, Pócs 1971 as Heteroscyphusdecurrens, Kitagawa 1979a as Chiloscyphusdecurrens, Miller et al. 1983 as Heteroscyphusdecurrens, Piippo 1985, 1989b, 1993b, Piippo and Tan 1992, Bapna and Kachroo 2000b as Chiloscyphusdecurrens, Yamada and Hayashi 2003, Hayashi and Yamada 2004. Seram: Sande Lacoste 1864 as Chiloscyphusdecurrens, Schiffner 1898 as Chiloscyphusdecurrens, Herzog 1921 as Chiloscyphusdecurrens, Grolle 1965 as Chiloscyphusdecurrens, Pócs 1971 as Heteroscyphusdecurrens, Kitagawa 1979a as Chiloscyphusdecurrens, Miller et al. 1983 as Heteroscyphusdecurrens, Piippo 1985, Akiyama 1986, Piippo 1989b, 1993b, Piippo and Tan 1992, 1993b, Yamada and Hayashi 2003, Hayashi and Yamada 2004, Akiyama 2009.

Heteroscyphuszollingeri (Gottsche) Schiffn. MOLUCCAS: Seram: Akiyama 1986, 2009.

Kurzia G.Martens

Kurziageniculata Mizut. MOLUCCAS: Seram: Akiyama 1986, 2009.

Kurziagonyotricha (Sande Lac.) Grolle. MOLUCCAS: Del Rosario 1971, 1975b both as Microlepidoziagonyotricha, Pócs et al. 2011. Seram: Akiyama 1986, 2009.

Leiomitra Lindb.

Leiomitrabreviseta (Steph.) R.M.Schust. MOLUCCAS: Seram: Akiyama 1986, 2009 both as Trichocoleabreviseta.

Lejeunea Lib.

Lejeuneaalbescens (Steph.) Mizut. MOLUCCAS: Miller et al. 1983. Bacan: Eifrig 1936 as Taxilejeuneaalbescens. Seram: Mizutani 1986a, Akiyama 2009, Lee 2013.

Lejeuneaanisophylla Mont. MOLUCCAS: Ambon: Pócs 2010, Lee 2013. Seram: Mizutani 1986a as Lejeuneacatanduana, Akiyama 2009 as Lejeuneacatanduana.

Lejeuneadiscreta Lindenb. MOLUCCAS: Mizutani 1971, Tixier 1973a, 1979 both as Hygrolejeuneadiscreta, Mizutani 1975, 1977, Miller et al. 1983, Long and Grolle 1990, Parihar et al. 1994, Zhu and So 1999, Bapna and Kachroo 2000b, Chantanaorrapint et al. 2003, 2004, Alam and Srivastava 2009, Haerida 2009, Lee et al. 2010. Bacan: Eifrig 1936 as Hygrolejeuneadiscreta.

Lejeuneaexilis (Reinw., Blume et Nees) Grolle. MOLUCCAS: Pócs et al. 2011. Ambon: LECTOTYPE of Harpalejeuneaexigua [Grolle & Reiner-Drehwald 1999: 41 as “holotype”], Stephani 1913 as Harpalejeuneaexigua, Grolle 1988 as Lejeuneamicrostipula, Grolle and Reiner-Drehwald 1999 as Microlejeuneamicrostipula. Seram: Mizutani 1986a, Akiyama 2009.

Lejeuneafissistipula (Steph.) Steph. MOLUCCAS: Ambon: LECTOTYPE of Eulejeuneafissistipula [Bonner 1965: 138], Stephani 1896b as Eulejeuneafissistipula, 1915, Schiffner 1898 as Eulejeuneafissistipula, Bonner 1965 as Eulejeuneafissistipula, Miller et al. 1983 also as Eulejeuneafissistipula. Seram: Mizutani 1986a, Akiyama 2009.

Lejeuneaflava(Sw.)Neessubsp.orientalis R.M.Schust. MOLUCCAS: LECTOTYPE of Jungermanniaindica [to be designated]. Ternate/Tidore: Reinwardt and Nees 1823 as Jungermanniaindica.

Lejeunealumbricoides (Nees) Nees. MOLUCCAS: Ambon: LECTOTYPE of Taxilejeuneakarstenii [to be designated], Schiffner 1893b, 1898 as Taxilejeunealumbricoides, Stephani 1914 as Taxilejeuneakarstenii, Eifrig 1936 as Taxilejeunealumbricoides, Kamimura 1974, Lee et al. 2014. Seram: Mizutani 1986a, Akiyama 2009, Lee 2013.

Lejeuneamicroloba Taylor. MOLUCCAS: Seram: Mizutani 1986a, Zhu and Reiner-Drehwald 2004 as Lejeuneachalmersii, Akiyama 2009, Lee 2013.

Lejeuneasordida (Nees) Nees. MOLUCCAS: Mizutani 1977, Miller et al. 1983 also as Hygrolejeuneaparkinsonii, Pócs et al. 2011. Ambon: LECTOTYPE of Hygrolejeuneaamboinensis [to be designated], Schiffner 1890, 1898 both as Hygrolejeuneaamboinensis, Sydow 1894 as Hygrolejeuneaamboinensis, Stephani 1914 as Hygrolejeuneaamboinensis, Eifrig 1936 as Taxilejeunealaxiretis and Taxilejeuneasordida, Miller et al. 1963, 1968 both as Hygrolejeuneasordida, Mizutani 1964 as Taxilejeuneasordida, Inoue and Miller 1965 as Hygrolejeuneasordida, Swanson and Miller 1969 as Hygrolejeuneasordida, Miller et al. 1983 also as Hygrolejeuneasordida and Hygrolejeuneaamboinensis, Thiers 1986, Lee 2013. Seram: Mizutani 1986a, Akiyama 2009.

Lejeuneastenodentata M.A.M.Renner et Pócs. MOLUCCAS: Mizutani 1977 as Drepanolejeuneadentata, Thouvenot 2015, Pócs et al. 2019a. Ambon: LECTOTYPE of Drepanolejeuneadentata [Bonner 1965: 82], Stephani 1896b, 1913 both as Drepanolejeuneadentata, Schiffner 1898 as Drepanolejeuneadentata, Bonner 1965 as Drepanolejeuneadentata, Mizutani 1972 as Drepanolejeuneadentata, Pócs et al. 1995 as Stenolejeuneadentata, Schuster 2000b as Stenolejeuneadentata, Renner et al. 2013.

Lejeuneaumbilicata (Nees) Nees. MOLUCCAS: Ternate/Tidore: Sande Lacoste 1856a, 1956b both as Omphalanthusumbilicatus, Schiffner 1898 as Hygrolejeuneaumbilicata.

Lepicolea Dumort.

Lepicolearara (Steph.) Grolle. MOLUCCAS: Pócs et al. 2011. Seram: Akiyama 1986, 2009.

Lepicoleayakusimensis (S.Hatt.) S.Hatt. MOLUCCAS: Seram: Akiyama 1986, 2009.

Lepidolejeunea R.M.Schust.

Lepidolejeuneabidentula (Steph.) R.M.Schust. MOLUCCAS: Geissler and Bischler 1989 as Pycnolejeuneadecurvifolia. Ambon: LECTOTYPE of Hygrolejeuneadecurvifolia [Mizutani 1978: 133], Stephani 1896b, 1910a both as Hygrolejeuneadecurvifolia, Hoffmann 1935 as Pycnolejeuneadecurvifolia, Bonner 1966 as Hygrolejeuneadecurvifolia, Mizutani 1978 as Pycnolejeuneabadia, Miller et al. 1983 as Hygrolejeuneadecurvifolia. Seram: Mizutani 1986a, Akiyama 2009.

Lepidozia (Dumort.) Dumort.

Lepidoziaborneensis Steph. MOLUCCAS: Seram: Akiyama 1986, 2009, Doei 1987b.

Lepidoziabrotheri Steph. MOLUCCAS: Seram: Doei 1987b, Akiyama 2009.

Lepidoziaceramensis Herzog. MOLUCCAS: Seram: LECTOTYPE [to be designated], Herzog 1926b, Doei 1987b.

Lepidoziacladorhiza (Reinw., Blume et Nees) Nees. MOLUCCAS: Seram: Akiyama 1986, 2009, Doei 1987b.

Lepidoziaferdinandii-muelleri Steph. MOLUCCAS: Halmahera: Engel 1987. Seram: Herzog 1926a as Lepidozia ‘Friderici Mülleri’, 1926b as Lepidozia ‘Ferdinandi Mülleri’, Grolle 1968b as Lepidozia ‘fernandi-muelleri’], Mizutani 1968b, 1974, Miller et al. 1983, Piippo 1984b, Akiyama 1986, 2009 as Lepidozia ‘ferdinandii-muelleri’, Doei 1987b, Enroth 1991, Schuster 2000a.

Lepidoziagriseola Herzog. MOLUCCAS: Seram: LECTOTYPE [to be designated], Herzog 1926b, Doei 1987b.

Lepidoziahaskarliana (Gottsche, Lindenb. et Nees) Steph. MOLUCCAS: Siregar et al. 2018. Seram: Akiyama 1986, 2009 as Lepidozia ‘hasskarliana’, Doei 1987b as Lepidozia ‘hasskarliana’, von Konrat et al. 2014, Siregar and Pasaribu 2019.

Lepidoziaintegrifolia Doei. MOLUCCAS: Seram: HOLOTYPE, Doei 1987a, 1987b, Engel 1993, Crosby and Engel 2006, Akiyama 2009.

Lepidozialongifolia Steph. MOLUCCAS: Ambon: LECTOTYPE [here designated], Stephani 1909a, Miller et al. 1983.

Lepidoziamassartiana Schiffn. ex Steph. MOLUCCAS: Ambon: Stephani 1909a, Kitagawa 1973, Miller et al. 1983, Geissler and Bischler 1985.

Lepidoziasubintegra Lindenb. MOLUCCAS: Inoue and Miller 1965, Mizutani 1968a, 1976, Del Rosario 1975b, Miller et al. 1983, Bapna and Kachroo 2000a, Bakalin et al. 2021. Seram: Akiyama 1986, 2009 [misidentification of Lepidoziasubtrichodes fide Doei (1987b: 525).].

Lepidoziasubtrichodes Steph. MOLUCCAS: Seram: Herzog 1926b, 2009, Doei 1987b.

Lepidoziatrichodes (Reinw., Blume et Nees) Nees. MOLUCCAS: Ambon: Schiffner 1893b, 1898, Mizutani 1968a, 1974, Pócs 1968, Del Rosario 1975b, Kamimura 1975 as Lepidozia ‘tricodes’], Miller et al. 1983, Piippo 1984b, Doei 1987b, Siregar et al. 2018. Seram: Akiyama 1986, 2009, Doei 1987b.

Leptolejeunea (Spruce) Steph.

Leptolejeuneadentistipula Steph. MOLUCCAS: Seram: Mizutani 1986a, Akiyama 2009 as ‘Lepidolejeunea’ dentistipula.

Leptolejeuneaelliptica (Lehm. et Lindenb.) Besch. MOLUCCAS: Dey et al. 2009. Seram: Mizutani 1986a, Akiyama 2009 as ‘Lepidolejeunea’ elliptica.

Leptolejeuneaepiphylla (Mitt.) Steph. MOLUCCAS: Bapna and Kachroo 2000b. Seram: Mizutani 1986a, Akiyama 2009 as ‘Lepidolejeunea’ epiphylla.

Leptolejeuneafoliicola Steph. MOLUCCAS: Tixier 1974b, Parihar et al. 1994, Bapna and Kachroo 2000b. Seram: Akiyama 2009 as ‘Lepidolejeunea’ foliicola.

Leptolejeuneamaculata (Mitt.) Schiffn. MOLUCCAS: Vanden Berghen 1977, Haerida 2009, Siregar et al. 2020a. Seram: Mizutani 1986a, Akiyama 2009 as ‘Lepidolejeunea’ maculata.

Leptolejeuneaschiffneri (Steph. ex Schiffn.) Steph. MOLUCCAS: Tixier 1972 as Leptolejeuneaschiffneri, Mizutani 1975 as Leptolejeuneaschiffneri, Miller et al. 1983 as Leptolejeuneaschiffneri, Geissler and Bischler 1985 as Leptolejeuneaschiffnerif.angustifolia, Bapna and Kachroo 2000b. Bacan: Herzog 1942 as Leptolejeuneaschiffnerivar.genuina.

Leptolejeuneavitrea (Nees) Schiffn. MOLUCCAS: Jovet-Ast 1958, Jovet-Ast and Tixier 1962, Tixier 1970a, 1970b, 1971, 1973a, 1977, 1979, 1980, Miller et al. 1983, Pócs et al. 1994, Eggers 2006, Siregar et al. 2020a. Bacan: Herzog 1942.

Lobatiriccardia (Mizut. et S.Hatt.) Furuki

Lobatiriccardiacoronopus (De Not.) Furuki. MOLUCCAS: Seram: Furuki 1996 as Lobatiriccardialobata, Furuki 2006.

— subsp. coronopus. MOLUCCAS: Seram: Preussing et al. 2010.

Lophocolea (Dumort.) Dumort.

Lophocoleadeningeri Herzog. MOLUCCAS: Buru: LECTOTYPE [here designated], Herzog 1921.

Lophocoleakurzii Sande Lac. MOLUCCAS: Seram: Akiyama 1986, 2009.

Lopholejeunea (Spruce) Steph.

Lopholejeuneaapplanata (Reinw., Blume et Nees) Schiffn. MOLUCCAS: Ternate/Tidore: Sande Lacoste 1864 as Lejeunea ‘adplanata’, Schiffner 1898. Ambon: Sande Lacoste 1864 as Lejeunea ‘adplanata’, Miller et al. 1983. Seram: Sande Lacoste 1864 as Lejeunea ‘adplanata’, Schiffner 1898, Miller et al. 1983.

Lopholejeuneaeulopha (Taylor) Schiffn. MOLUCCAS: Haerida et al. 2010, Siregar et al. 2020a. Halmahera: Verdoorn 1934a, 1934b, Hattori 1951, Kamimura 1974, Miller et al. 1983. Ternate/Tidore: Mizutani 1979, Miller et al. 1983, Zhu and Gradstein 2005. Ambon: Schiffner 1890 as Lopholejeuneaeulopha, Sydow 1894, Verdoorn 1934a, 1934b, Hattori 1951, Kamimura 1974, Miller et al. 1983. Seram: Mizutani 1979, 1986a, Miller et al. 1983, Akiyama 2009.

Lopholejeuneahorticola Schiffn. MOLUCCAS: Haerida et al. 2010, Siregar et al. 2014, Siregar 2015. Seram: Mizutani 1986a, Akiyama 2009.

Lopholejeuneanigricans (Lindenb.) Schiffn. MOLUCCAS: Mizutani 1994, Haerida et al. 2010, Siregar et al. 2014, Siregar et al. 2020a. Ternate/Tidore: Verdoorn 1934b. Seram: Mizutani 1986a, Akiyama 2009 also as Lopholejeuneajavanica.

Lopholejeuneasubfusca (Nees) Schiffn. MOLUCCAS: Mizutani 1994, Haerida 2009, Haerida et al. 2010, Siregar et al. 2014, 2020a, Siregar 2015, Rosyanti et al. 2018. Ambon: Schiffner 1893b as Lejeunea ‘Sagraeana’, 1898 as Lopholejeuneasagranavar.subfusca, Stephani 1912b as Lopholejeuneapyriflora, Verdoorn 1934a, 1934b, Hattori 1951, Jovet-Ast 1958, Inoue and Miller 1965, Tixier 1966, 1967a, 1967b, Pócs et al. 1967, Swanson and Miller 1969, Miller et al. 1983 also as Lopholejeuneapyriflora and Lopholejeuneasagrana, Geissler and Bischler 1985 as Lopholejeuneapyriflora, Bapna and Kachroo 2000b, Gradstein et al. 2002, Zhu and Gradstein 2005. Banda: Verdoorn 1934b, Jovet-Ast 1958, Tixier 1966, 1967a, Swanson and Miller 1969. Seram: Mizutani 1986a, Zhu and Gradstein 2005, Akiyama 2009.

Lopholejeuneawiltensii Steph. MOLUCCAS: Haerida et al. 2010, Siregar 2015. Halmahera: Umagap 2019. Ternate/Tidore: Mizutani 1979, Zhu and Gradstein 2005. Seram: Mizutani 1979, Zhu and Gradstein 2005, Siregar et al. 2014, Siregar 2015.

Marchantia L.

Marchantiaemarginata Reinw., Blume et Nees. MOLUCCAS: Siregar et al. 2013, Siregar 2015, Ginting and Batubara 2019, Siregar et al. 2020b, Ruklani and Rubasinghe 2022. Ternate/Tidore: Bischler and Piippo 1991. Seram: Bischler and Piippo 1991.

— subsp. emarginata. MOLUCCAS: Bischler 1989, Singh and Singh 2013. Ternate/Tidore: Bischler-Causse 1989. Ambon: LECTOTYPE of Marchantiaamboinensis [Bischler-Causse 1989: 183 as “type”, second-step lectotypification needed], Montagne 1838, 1856 both as Marchantiaamboinensis, Gottsche et al. 1846 as Marchantiaamboinensis, Schiffner 1898 as Marchantiaamboinensis, Stephani 1899a as Marchantiaamboinensis, Miller et al. 1983 as Marchantiaamboinensis, Bischler-Causse 1989, Bischler and Piippo 1991. Seram: Bischler-Causse 1989.

Marchantiageminata Reinw., Blume et Nees. MOLUCCAS: Halmahera: Umagap 2019.

Marchantiapolymorpha L. MOLUCCAS: Ambon: Sopacua et al. 2020 [Saparua I].

Marchantiarubribarba Steph. MOLUCCAS: Ambon: Bischler-Causse 1989, Bischler and Piippo 1991.

Mastigophora Nees

Mastigophoradiclados (Brid. ex F.Weber) Nees. MOLUCCAS: Buru: Herzog 1921. Halmahera: LECTOTYPE of Sendtneravrieseana [to be designated], Sande Lacoste 1864 as Sendtneradiclados β calcarata and Sendtneravrieseana, Schiffner 1898 also as Mastigophoravrieseana, Stephani 1909c as Mastigophoravrieseana, Inoue and Miller 1965, Miller et al. 1983. Ternate/Tidore: Sande Lacoste 1864 as Sendtneradiclados α scorpioides, Schiffner 1898 as Mastigophoradicladosvar.scorpioides, Inoue and Miller 1965, Miller et al. 1983. Ambon: Schiffner 1898, Inoue and Miller 1965, Miller et al. 1983. Seram: Herzog 1921, Akiyama 1986, 2009.

— var. borneensis (De Not.) Schiffn. MOLUCCAS: Ambon: Schiffner 1893b, 1898.

— var. diclados. MOLUCCAS: Ambon: Schiffner 1893b as Mastigophoradiclados.

Metalejeunea Grolle

Metalejeuneacucullata (Reinw., Blume et Nees) Grolle. MOLUCCAS: Grolle 1995. Ambon: SYNTYPE of Microlejeuneaparallela [lectotype to be designated], Schiffner 1890 as Microlejeuneaparallela, 1898 as Eulejeuneaparallela, Sydow 1894 as Microlejeuneaparallela, Stephani 1915 as Microlejeuneaparallela. Seram: Mizutani 1986a as Lejeuneacucullata, Akiyama 2009 as Lejeuneacucullata.

Metzgeria Raddi

Metzgeriaciliata Raddi. MOLUCCAS: Halmahera: Umagap 2019 as Metzgeriadecipiens.

Metzgeriafoliicola Schiffn. MOLUCCAS: Ternate/Tidore: So 2003c. Seram: Akiyama 1986, 2009.

Metzgeriafurcata (L.) Corda. MOLUCCAS: Halmahera: Umagap 2019.

MetzgerialeptoneuraSprucevar.leptoneuraMOLUCCAS: Engel 1976, Schuster 1983. Buru: So 2003c. Ambon: LECTOTYPE of Metzgeriahamatiformis [to be designated], Schiffner 1893b, 1898 both as Metzgeriahamatiformis, Stephani 1899d as Metzgeriahamatiformis, Kuwahara 1966 as Metzgeriahamata, 1984, Schuster 1992, Engel 1978, Miller et al. 1983, So 2003c, Costa 2008. Seram: Akiyama 1986, 2009.

Microlejeunea (Spruce) Steph.

Microlejeuneafilicuspis (Steph.) Heinrichs, Schäf.-Verw., Pócs et S.Dong. MOLUCCAS: Grolle 1979a as Harpalejeuneafilicuspis, Miller et al. 1983 as Drepanolejeuneafilicuspis. Ambon: Grolle 1979a as Harpalejeuneafilicuspis.

Mnioloma Herzog

Mniolomafuscum (Lehm.) R.M.Schust. MOLUCCAS: Seram: Akiyama 1986, 2009 both as Calypogeiafusca.

Neolepidozia Fulford et J.Taylor

Neolepidoziacuneifolia (Steph.) Fulford et J.Taylor. MOLUCCAS: Ambon: Engel and Smith Merrill 2004 as Telaraneacuneifolia, von Konrat et al. 2014.

Neolepidoziapapulosa (Steph.) Fulford et J.Taylor. MOLUCCAS: Seram: Akiyama 1986, 2009 both as Lepidoziapapulosa, Doei 1987b as Lepidoziapapulosa.

Neolepidoziawallichiana (Gottsche) Fulford et J.Taylor. MOLUCCAS: Ambon: Sande Lacoste 1864 as Lepidoziawallichiana, De Notaris 1874 as Lepidoziawallichiana, Schiffner 1893b, 1898 both as Lepidoziawallichiana, Hattori and Mizutani 1958 as Lepidoziawallichiana, Inoue and Miller 1965 as Lepidoziawallichiana, Swanson and Miller 1969, Mizutani 1974, 1976 both as Lepidoziawallichiana, Del Rosario 1975b as Lepidoziawallichiana, Kamimura 1975 as Lepidoziawallichiana, Miller et al. 1983, Piippo 1984b as Lepidoziawallichiana, Doei 1987b as Lepidoziawallichiana, Sharma and Srivastava 1993 as Lepidoziawallichiana, Bapna and Kachroo 2000a as Lepidoziawallichiana, Engel and Smith Merrill 2004 as Telaraneawallichiana. Seram: Doei 1987b as Lepidoziawallichiana, Akiyama 2009 as Lepidoziawallichiana.

Pallavicinia Gray

Pallaviciniaambigua (Mitt.) Steph. MOLUCCAS: Seram: Grolle and Piippo 1986, Koponen et al. 2000, Furuki 2002.

Pallavicinialyellii (Hook.) Gray. MOLUCCAS: Inoue and Miller 1965, Swanson and Miller 1969, Miller et al. 1983, Bapna and Kachroo 2000b, Manju et al. 2015. Halmahera: Umagap 2019.

Plagiochila (Dumort.) Dumort.

Plagiochilaabietina (Nees) Mont. et Nees. MOLUCCAS: Ambon: Stephani 1903b, Bonner 1962a, Miller et al. 1983.

Plagiochilaakiyamae Inoue. MOLUCCAS: Seram: HOLOTYPE, Inoue 1986, Engel 1993, Grolle and So 1998, Grolle and So 1999, So and Grolle 2000, So 2001b, Crosby and Engel 2006, Akiyama 2009, Singh and Rawat 2020.

Plagiochilaamboynensis Taylor. MOLUCCAS: Geissler and Bischler 1989 [type, Amboina], So and Grolle 2000 [type, Amboina]. Ambon: LECTOTYPE [So and Grolle 2000: 201 as “holotype”], Taylor 1846, Gottsche et al. 1847 as Plagiochila ‘amboinensis’, Schiffner 1898 as Plagiochila ‘amboinensis’, Stephani 1904a as Plagiochila ‘amboinensis’, Bonner 1962a, Inoue 1984, So 2001c, Schuster 2021.

Plagiochilaarbuscula (Brid. ex Lehm. et Lindenb.) Lindenb. MOLUCCAS: Halmahera: Sande Lacoste 1864 as Plagiochilabelangeriana, Schiffner 1898 as Plagiochilabelangeriana, Miller et al. 1983. Ternate/Tidore: Sande Lacoste 1864 as Plagiochilabelangeriana, Miller et al. 1983. Ambon: Sande Lacoste 1864 as Plagiochilabelangeriana [Saparoea I], Schiffner 1893b, 1898 both as Plagiochilabelangeriana, Miller et al. 1983. Seram: Inoue 1986, So 2001b, Akiyama 2009.

Plagiochilabantamensis (Reinw., Blume et Nees) Mont. MOLUCCAS: Seram: Inoue 1986, Akiyama 2009.

Plagiochilabicornuta Steph. MOLUCCAS: Pócs et al. 2011. Ambon: Stephani 1903a, Bonner 1962a, Inoue 1981, Miller et al. 1983, So 2000a, 2001a, Schuster 2021. Seram: So 2000a, 2001a.

Plagiochilablepharophora (Nees) Lindenb. MOLUCCAS: Ternate/Tidore: Schiffner 1898, 1900, Stephani 1903c, Miller et al. 1983.

Plagiochilachauviniana Mont. MOLUCCAS: Seram: Inoue 1986 as Plagiochilanovae-guineae, Akiyama 2009 as Plagiochilanovae-guineae.

Plagiochilafrondescens (Nees) Lindenb. MOLUCCAS: Ambon: Sande Lacoste 1864 as Plagiochilafrondescens γ rigida, Schiffner 1893b, 1898 1900 all as Plagiochilafrondescensvar.tenerrima, 1898 as Plagiochilafrondescens γ rigida, Stephani 1903a, Swanson and Miller 1969, Miller et al. 1983, Piippo 1989a. Seram: Inoue 1986, Akiyama 2009.

Plagiochilagymnoclada Sande Lac. MOLUCCAS: Singh Deo and Singh 2014. Buru: Herzog 1921, Carl 1931b, Inoue 1984, Inoue 1989. Ternate/Tidore: Sande Lacoste 1864, Schiffner 1898, 1900, Stephani 1903b. Seram: Inoue 1986 as Plagiochilapseudaberrans, Akiyama 2009 as Plagiochilapseudaberrans.

Plagiochilahampeana Gottsche. MOLUCCAS: Seram: Inoue 1986 as Plagiochilagedeana, So and Grolle 2001, Akiyama 2009 as Plagiochilagedeana.

Plagiochilajunghuhniana Sande Lac. MOLUCCAS: Ambon: So 2000b, 2001b.

Plagiochilakuhliana Sande Lac. MOLUCCAS: Seram: Sande Lacoste 1864, Schiffner 1898, 1900, Stephani 1903d, Bonner 1962a, Geissler and Bischler 1989.

Plagiochilakurzii Steph. MOLUCCAS: Seram: Sande Lacoste 1864 as Plagiochilabantamensisvar.denticulata, Schiffner 1898 as Plagiochilabantamensisvar.denticulata, Bonner 1962a as Plagiochilabantamensisvar.denticulata.

Plagiochilamassalongoana Schiffn. MOLUCCAS: Seram: Inoue 1986, Akiyama 2009.

Plagiochilapulvinata Steph. MOLUCCAS: So and Grolle 1999. Seram: HOLOTYPE of Plagiochilaceramica, Inoue 1986 as Plagiochilaceramica, Engel 1993 as Plagiochilaceramica, So and Grolle 2000, Crosby and Engel 2006 as Plagiochilaceramica, Akiyama 2009 as Plagiochilaceramica.

Plagiochilarenitens (Nees) Lindenb. MOLUCCAS: ?Halmahera: Schiffner 1900, Stephani 1903b. Seram: Inoue 1984, 1986, 1989, Piippo 1989a, Akiyama 2009, Schuster 2021. Note: we are not aware of any first hand report from Halmahera.

Plagiochilasalacensis Gottsche. MOLUCCAS: Seram: Inoue 1986, Akiyama 2009.

Plagiochilasandei Dozy. MOLUCCAS: So and Grolle 1999. Ternate/Tidore: Schiffner and Stephani 1901 as Plagiochilablepharophoravar.major. Ambon: Sande Lacoste 1864 [Saparoea I], Schiffner 1898, 1900 [both Saparoea I], Stephani 1903d [Saparoea I], Grolle 1968b as Plagiochilaseemannii, Miller et al. 1983 as Plagiochilaseemannii, Inoue 1986 as Plagiochilaseemannii, Piippo 1989a as Plagiochilaseemannii. Seram: Sande Lacoste 1864, Stephani 1903d, Inoue 1986 also as Plagiochilaseemannii, 1989, Müller and Schäfer-Verwimp 1999, Akiyama 2009 also as Plagiochilaseemannii.

Plagiochilasciophila Nees. MOLUCCAS: ?Ambon: Pócs 1971 as Plagiochilaacanthophylla. Seram: Inoue 1986 also as Plagiochiladecidua, 1989, Akiyama 2009 also as Plagiochiladecidua. Note: we aree not aware of any first hand report from Ambon.

Plagiochilateysmannii Sande Lac. MOLUCCAS: So and Grolle 1999. Bacan: LECTOTYPE of Plagiochilabatjanensis [Inoue 1984: 90 as “type”], Herzog 1938 as Plagiochilabatjanensis, Bonner 1962a as Plagiochilabatjanensis, Inoue 1984, Geissler and Bischler 1989 as Plagiochilabatjanensis, Piippo 1989a, So and Grolle 2000. Ambon: Schiffner 1893b, 1898 both as Plagiochilasemialata. Seram: Inoue 1986, 1989, Akiyama 2009 as Plagiochila ‘teysmanii’.

Plagiochilatrabeculata Steph. MOLUCCAS: Seram: So 2001b.

Pleurozia Dumort.

Pleuroziaconchifolia (Hook. et Arn.) Austin. MOLUCCAS: Pócs et al. 2011. Ambon: Schiffner 1898Pleurozia ‘conchaefolia’, Miller et al. 1983.

Pleuroziagigantea (F.Weber) Lindb. MOLUCCAS: Ambon: Schiffner 1893b, 1898, Horikawa 1935, Pócs et al. 1967, Tixier 1973b, 1974b, Bizot and Pócs 1974, Miller et al. 1983, Thiers 1993. Seram: Akiyama 1986, 2009.

Plicanthus R.M.Schust.

Plicanthushirtellus (F.Weber) R.M.Schust. MOLUCCAS: Seram: LECTOTYPE of Chandonanthusgracilis [here designated], Herzog 1921 as Chandonanthushirtellus, 1926a as Chandonanthusgracilis, Herzog 1926b, 1926b both as Chandonanthushirtellus, Bonner 1963b as Chandonanthusgracilis, Akiyama 1986, 2009 both as Chandonanthushirtellus, Váňa et al. 2013.

Podomitrium Mitt.

Podomitriummalaccense (Steph.) Campb. MOLUCCAS: Ambon: Troll 1930 as Hymenophytonmalaccense, Grolle 1966a, Schuster 1969 as Podomitrium ‘mallacense’, Miller et al. 1983, Grolle and Piippo 1986. Seram: Akiyama 1986, 2009.

Porella L.

Porellaacutifolia(Lehm. et Lindenb.)Trevis.var.acutifoliaMOLUCCAS: Halmahera: Sande Lacoste 1864 as Madothecaacutifolia, Schiffner 1898 as Madothecaacutifolia, Hattori 1970 as Porellaacutifoliavar.subligulifera.

Porellageheebii (Steph.) S.Hatt. MOLUCCAS: Seram: Bi et al. 2019.

Porellajavanica (Gottsche) Inoue. MOLUCCAS: Hattori 1978, Miller et al. 1983. Buru: LECTOTYPE of Madothecacrenilobula [here designated], Herzog 1921, 1926a both as Madothecacrenilobula.

Psiloclada Mitt.

PsilocladaclandestinaMitt.subsp.clandestinaMOLUCCAS: Ambon: LECTOTYPE of Psilocladaunguligera [to be designated], Schiffner 1893a, 1893b, 1898 all as Psilocladaunguligera, Stephani 1909a, Herzog 1953, Arnell 1955, Fulford and Taylor 1959 as Psilocladaunguligera, Arnell 1963, Mizutani 1974, 1976, Schuster 1980, Hürlimann 1983, Miller et al. 1983. Seram: Akiyama 1986, 2009.

— subsp. melanesica R.M.Schust. ?MOLUCCAS: Ambon: Miller 1985, Schuster 2000a with a “?”], Engel and Glenny 2008. Note: we are not aware of any positively identified specimen of this subspecies from Moluccas.

Ptychanthus Nees

Ptychanthusstriatus(Lehm. et Lindenb.)Neesvar.striatus. MOLUCCAS: Mizutani 1977, Miller et al. 1983 as Ptychanthusmoluccensis, Haerida et al. 2010, Siregar et al. 2014, Siregar 2015, Siregar et al. 2020a. Buru: Herzog 1921, 1926a both as Ptychanthusmoluccensis. Halmahera: Sande Lacoste 1864 as Ptychanthusjavanicus, Schiffner 1898 as Ptychanthusjavanicus, Verdoorn 1934a, 1934b, Umagap 2019. Ambon: LECTOTYPE of Ptychanthussquarrosus [to be designated], Lehmann 1844 as Ptychanthussquarrosus, Gottsche et al. 1845a as Ptychanthussquarrosus, Montagne 1856 as Ptychanthussquarrosus, Stephani 1890a, Schiffner 1898, Verdoorn 1933 as Ptychanthusmoluccensis, 1934a, Mizutani 1961, Tixier 1971, Kamimura 1974, Miller et al. 1983, Bapna and Kachroo 2000b. Seram: SYNTYPE of Ptychanthusmoluccensis [lectotype to be designated], Sande Lacoste 1864 as Ptychanthusmoluccensis, Schiffner 1898 as Ptychanthusmoluccensis, Verdoorn 1933 as Ptychanthusmoluccensis, 1934b, Miller et al. 1983 as Ptychanthusstriatus, Mizutani 1986a, Akiyama 2009.

Pycnolejeunea (Spruce) Schiffn.

Pycnolejeuneacavistipula (Steph.) Mizut. MOLUCCAS: Ambon: LECTOTYPE of Strepsilejeuneacavistipula [Mizutani 1972: 161 as “type”], Stephani 1896b as Strepsilejeuneacavistipula, 1913 as Trachylejeuneacavistipula, Schiffner 1898 as Strepsilejeuneacavistipula, Mizutani 1972, Thiers 1986.

Pycnolejeuneaconnivens Schiffn. ex P.Syd. MOLUCCAS: Ambon: LECTOTYPE of Pycnolejeuneaconnivens [to be designated], Schiffner 1890 as Pycnolejeuneaconnivens, Sydow 1894.

Radula Dumort.

Radulaanceps Sande Lac. MOLUCCAS: Seram: Akiyama 1986, 2009, Yamada 1989, Bapna and Kachroo 2000b.

Raduladensifolia Castle. MOLUCCAS: Seram: Akiyama 1986, 2009, Yamada and Piippo 1989, Yamada 1989, Yamada 2002.

Radulaformosa (C.F.W.Meissn. ex Spreng.) Nees. MOLUCCAS: Ambon: LECTOTYPE of Radulapycnolejeuneoides [Castle 1950: 267; second-step lectotypification needed], Schiffner 1893b, 1898 both as Radulapycnolejeuneoides, Stephani 1910b also as Radulapycnolejeuneoides, Jovet-Ast 1949, Castle 1950, 1968, Steere 1953, Inoue and Miller 1965, Grolle 1966a, Kitagawa 1979a as Radulaformosavar.pycnolejeuneoides, Miller et al. 1983 also as Radulapycnolejeuneoides, So 2006. ?Seram: Yamada 1979, 1989, Miller et al. 1983, Siregar 2015, Ginting and Batubara 2019. Note: we are not aware of any first hand report from Seram.

Radulajavanica Gottsche. MOLUCCAS: Buru: New to Moluccas: Buru, E hoe, 1921.09, Toxopens 299a, det. H. Castle (BO4137]. Seram: LECTOTYPE of Radulaceramensis [to be designated], LECTOTYPE of Radulaovalifolia [Castle 1966: 68; second-step lectotypification needed)], Stephani 1884 as Radulaovalifolia and Radulaceramensis, Schiffner 1898 as Radulaceramensis and Radulaovalifolia, Stephani 1910b, 1924 both as Radulaceramensis, Castle 1966 as Radulawallichiana, Miller et al. 1983 as Radulaceramensis, Akiyama 1986, 2009, Yamada 1989.

Radulamultiflora Gottsche ex Schiffn. MOLUCCAS: Seram: Akiyama 1986.

Radulanymannii Steph. MOLUCCAS: Ambon: Swanson and Miller 1969, Miller et al. 1983. Seram: Akiyama 1986, 2009 as Radula ‘nymanii’, Yamada 1989 as Radula ‘nymanii’.

Radulareflexa Nees et Mont. MOLUCCAS: Ambon: LECTOTYPE [Castle 1965: 374], Montagne 1843, 1846, 1856, Gottsche et al. 1845a, Schiffner 1898, Castle 1965, 1968, Yamada 1976, 1979, 2002, Miller et al. 1983, So 2005, 2006. Seram: LECTOTYPE of Radulasubsimilis [Castle 1965: 376 as “type”], Sande Lacoste 1864, Stephani 1884 as Radulasubsimilis, Schiffner 1898 also as Radulasubsimilis, Stephani 1910b as Radulasubsimilis, Castle 1965, 1968 both as Radulasubsimilis, Yamada 1979 as Radulasubsimilis, 1987, Miller et al. 1983 also as Radulasubsimilis, Yamada and Piippo 1989, Streimann 1991 as Radulasubsimilis, Piippo et al. 2002, So 2006, Siregar 2015.

Radulasumatrana Steph. MOLUCCAS: Seram: Akiyama 1986, 2009, Yamada 1989.

Radulatjibodensis K.I.Goebel. MOLUCCAS: Seram: Akiyama 1986, 2009, Yamada 1989, Zhu 1999, Zhu and So 2001.

Radulavan-zantenii K.Yamada. MOLUCCAS: Seram: Akiyama 1986, 2009 as Radula ‘vanzantenii’, Yamada and Piippo 1989.

Radulavrieseana Sande Lac. MOLUCCAS: Siregar 2015. Obi: Yamada 1979. Banda: Sande Lacoste 1864, Schiffner 1898. Seram: LECTOTYPE [Castle 1965: 372], Sande Lacoste 1864, Schiffner 1898, Stephani 1910b as Radula ‘vriesei’, Castle 1965, 1968 both as Radula ‘Vriesei’, Yamada 1974 as Radula ‘vriesei’, 1979, Akiyama 1986, 1989, 2009, Yamada and Piippo 1989, Bapna and Kachroo 2000b, Siregar 2015.

Riccardia Gray

Riccardiaalbomarginata (Steph.) Schiffn. MOLUCCAS: Ambon: LECTOTYPE of Aneuraalbomarginata [Bonner 1962b: 85, cf. also Furuki 1995: 112 as “holotype”], Stephani 1893a, 1893b, 1899c all as Aneura ‘albo-marginata’, Schiffner 1898, Bonner 1962b as Aneuraalbomarginata, Grolle 1967, 1968b, Miller et al. 1983 also as Aneuraalbomarginata, Furuki 1995, Thouvenot et al. 2018.

Riccardiaaspera (Steph.) Grolle. MOLUCCAS: Seram: Furuki 1998.

Riccardiakarstenii (Steph.) Schiffn. MOLUCCAS: Ambon: LECTOTYPE of Aneurakarstenii [Bonner 1962b: 114], Stephani 1893a, 1893b, 1899c all as Aneurakarstenii, Schiffner 1898, Bonner 1962b as Aneurakarstenii.

Riccia L.

Ricciaamboinensis Schiffn. MOLUCCAS: Ambon: LECTOTYPE [to be designated], Schiffner 1890, 1898, Sydow 1894, Stephani 1898, Campbell 1915 as Riccia ‘amboiniana’.

Ricciabillardierei Mont. et Nees. MOLUCCAS: Bapna and Kachroo 2000b, Chaudhary et al. 2006. Halmahera: Meijer 1958, Jovet-Ast 2000, 2003. Banda: Jovet-Ast 2003.

Ricciajunghuhniana Nees et Lindenb. MOLUCCAS: Ambon: Xiang et al. 2022.

Saccogynidium Grolle

Saccogynidiummuricellum (De Not.) Grolle. MOLUCCAS: Miller et al. 1983, Enroth 1991. Bacan: Grolle and Schultze-Motel 1972. ?Ambon: Piippo 1985, 1989b, Gao et al. 2001. Seram: Akiyama 1986, 2009. Note: we are not aware of any first hand report from Ambon.

Saccogynidiumrigidulum (Nees) Grolle. MOLUCCAS: Inoue and Miller 1965, Grolle and Schultze-Motel 1972, Miller et al. 1983, Enroth 1991. Ambon: LECTOTYPE of Chiloscyphusgranulatus [to be designated], Schiffner 1893b, 1898 as Chiloscyphusgranulatus, Stephani 1908a as Saccogynarigidula, Hattori 1951 as Saccogynarigidula, Bonner 1963b as Chiloscyphusgranulatus, Inoue and Miller 1965, Grolle 1968b as Saccogynidiumjugatum, Miller et al. 1983 also as Chiloscyphusgranulatus, Piippo 1985, 1989b, Gao et al. 2001.

Sandeothallus R.M.Schust.

Sandeothallusradiculosus (Schiffn.) R.M.Schust. MOLUCCAS: Ambon: Akiyama 2009. Seram: Akiyama 1986.

Schiffneria Steph.

Schiffneriahyalina Steph. MOLUCCAS: Engel 1987, Long and Grolle 1990, Piippo and Tan 1992, Váňa 1993, O’Shea et al. 1997, Bapna and Kachroo 2000a, Schuster 2002. Halmahera: Bacan: LECTOTYPE [Grolle and Piippo 1984: 305 ad “holotype”], Stephani 1894b, 1908a, Schiffner 1898, Kitagawa 1973, Grolle and Piippo 1984, Asthana et al. 1994, Váňa et al. 2014.

Schiffneriolejeunea Verd.

Schiffneriolejeuneacumingiana (Mont.) Gradst. MOLUCCAS: Ambon: Verdoorn 1934a, 1934b both as Ptychocoleuscumingianus, Tixier 1966, 1967b, 1972 all as Ptychocoleuscumingianus, Miller 1968 as Ptychocoleuscumingianus, Swanson and Miller 1969 as Ptychocoleuscumingianus, Kachroo 1970 as Ptychocoleuscumingianus, Miller et al. 1983, Bapna and Kachroo 2000b. Seram: Verdoorn 1934a, 1934b both as Ptychocoleuscumingianus, Miller 1968 as Ptychocoleuscumingianus, Swanson and Miller 1969 as Ptychocoleuscumingianus, Gradstein and Terken 1981, Miller et al. 1983.

Schiffneriolejeuneapulopenangensis (Gottsche) Gradst. MOLUCCAS: Bapna and Kachroo 2000b, Siregar et al. 2020a. Ambon: ORIGINAL MATERIAL of Acrolejeuneadensifolia, Schiffner 1890 as Acrolejeuneadensifolia, 1898 as Acrolejeuneapulopenangensis, Sydow 1894 as Acrolejeuneadensifolia, Stephani 1912b as Ptychocoleusdensifolius, Verdoorn 1934b as Ptychocoleuspulopenangensis, Bonner 1962b as Acrolejeuneadensifolia, Mizutani 1964 as Ptychocoleuspulopenangensis, Miller et al. 1983 also as Acrolejeuneadensifolia, Hürlimann 1989.

Schiffneriolejeuneatumida (Nees) Gradst. MOLUCCAS: Ambon: Gradstein and Terken 1981. Seram: Mizutani 1986a, Akiyama 2009.

— var. haskarliana (Gottsche) Gradst. et Terken. MOLUCCAS: Mizutani 1977, 1978 both as Schiffneriolejeunea ‘hasskarliana’. Ambon: Schiffner 1893b as Lejeuneahaskarliana, 1898 as Acrolejeuneahaskarliana, Verdoorn 1934a, 1934b as Ptychocoleus ‘Hasskarlianus’, Amakawa 1960 as Ptychocoleushaskarlianus, Miller et al. 1983 as Acrolejeuneahaskarliana and Schiffneriolejeuneahaskarliana.

Schistochila Dumort.

Schistochilaacuminata Steph. MOLUCCAS: Seram: LECTOTYPE of Schistochilapurpurascens [here designated, cf. So 2003a: 80 as “holotype”)], Herzog 1926a, 1926b both as Schistochilapurpurascens, Grolle 1966b as Schistochilapurpurascens, Hässel 1976 as Schistochilapurpurascens, So 2003a.

Schistochilaaligera (Nees et Blume) J.B.Jack et Steph. MOLUCCAS: Pócs 1971, Tixier 1972. Buru: Herzog 1921, 1926a also as Schistochilagraeffeana, Miller et al. 1983 as Schistochilastergraeffeanus and Schistochilagraeffeana, So 2003a. Bacan: Buch 1939. ?Halmahera: Miller et al. 1983 as Schistochilasteraliger. Ambon: ORIGINAL MATERIAL of Schistochilaphilippinensisvar.transiens, Schiffner 1893b as ‘Schistocheila’ aligera, 1898, Stephani 1896a as ‘Schistocheila’ philippinensis, 1909c, Buch 1939 also as Schistochilaphilippinensis, Bonner 1966 as Gottscheaaligera, Grolle 1966b also as Schistochilaphilippinensis, Grolle 1966b, Pócs 1968, 1971, Tixier 1972, Miller et al. 1983 as Gottscheaaligera, Schistochilasteraliger and as Schistochilasterphilippinensis, Piippo 1984a also Schistochilaphilippinensis, Bapna and Kachroo 2000b, So 2003a. Seram: Akiyama 1986, 2009 both also as Schistochilaphilippinensis and Schistochilarecurvata.

Schistochilabeccariana (De Not.) Trevis. MOLUCCAS: Ambon: LECTOTYPE of Schistochilaamboinensis [here designated, cf. So 2003a: 87 as “holotype”], Stephani 1909c as Schistochilaamboinensis, So 2003a.

Schistochilablumei (Nees) Trevis. MOLUCCAS: Tixier 1974a. Halmahera: Sande Lacoste 1864 as Gottscheablumei, Schiffner 1898 as Schistochila ‘blumii’, Buch 1939, Grolle 1966b, Pócs 1971, Kitagawa 1973, 1978, Miller et al. 1983, Piippo 1984a, Inoue 1985, Enroth 1991 as Schistochila ‘blumii’. Ambon: So 2003a. Seram: Akiyama 1986, 2009 both as Schistochilaformosana.

Schistochiladoriae (De Not.) Trevis. MOLUCCAS: Ambon: Schiffner 1893b as ‘Schistocheila’ doriae, 1898.

Schistochilareinwardtii (Nees) Schiffn. MOLUCCAS: Ambon: So 2003a.

Schistochilasciurea(Nees)Schiffn.var.sciurea. MOLUCCAS: Ambon: Schiffner 1893b as ‘Schistocheila’ sciurea f. robustior, 1898, Verdoorn 1937, Hattori 1968, Miller et al. 1983, So 2003a. Seram: LECTOTYPE of Schistochilainversa [here designated, cf. So 2003a: 98 as “holotype”], Herzog 1926b as Schistochilainversa, Grolle 1966b as Schistochilainversa, Hässel 1976 as Schistochilainversa, Akiyama 1986, 2009, So 2003a.

Schizophyllopsis Váňa et L.Söderstr.

Schizophyllopsisbidens (Reinw., Blume et Nees) Váňa et L.Söderstr. MOLUCCAS: Seram: Akiyama 1986, 2009 both as Anastrophyllumbidens, Váňa and Piippo 1989b as Anastrophyllumbidensvar.bidens as Anastrophyllumbidens, Váňa 1991b as Anastrophyllumbidens, Schill and Long 2003 as Anastrophyllumbidens, Eggers 2006 as Anastrophyllumbidens.

Solenostoma Mitt.

Solenostomaariadne (Taylor) R.M.Schust. ex Váňa et D.G.Long. MOLUCCAS: Ambon: Sande Lacoste 1864 as Jungermanniaariadne, Schiffner 1898, 1900 both as Nardiaariadne, Stephani 1901a as Jungermanniaariadne, Amakawa 1968 as Jungermanniaariadne, Swanson and Miller 1969 as Jungermanniaariadne, Váňa 1972 as Jungermanniaariadne [rejecting report by Sande Lacoste 1864 as J.tetragona], Bonner 1976 as Jungermanniaariadne, Miller et al. 1983 as Plectocoleaariadne, Váňa and Piippo 1989a as Jungermanniaariadne. Seram: Akiyama 1986, 2009 both as Jungermanniaariadne, Váňa and Piippo 1989a as Jungermanniaariadne, Váňa 1991a as Jungermanniaariadne, Bapna and Kachroo 2000a as Jungermanniaariadne.

Solenostomacomatum(Nees)C.Gaovar.comatum. MOLUCCAS: Ternate/Tidore: Sande Lacoste 1856b as Plagiochilacomata, Schiffner 1898, 1900 both as Nardiacomata, Stephani 1901a as Jungermanniacomata, Hattori 1951 as Plectocoleacomata, Bonner 1965 as Eucalyx ‘comatus’, Váňa 1972 as Jungermanniacomata, 1973 as Jungermanniacomata [with a ‘?’], Miller et al. 1983 as Jungermanniacomata, Váňa and Piippo 1989a as Jungermanniacomata.

Solenostomatetragonum (Lindenb.) R.M.Schust. ex Váňa et D.G.Long. MOLUCCAS: Ambon: Váňa 1972, 1973, 1975, 1991a all both as Jungermanniatetragona, Hattori 1975b as Jungermanniatetragona, Miller et al. 1983 as Jungermanniatetragona, Váňa and Piippo 1989a as Jungermanniatetragona, Bapna and Kachroo 2000a as Jungermanniatetragona, Srivastava and Sharma 2000 as Jungermanniatetragona, Easa 2003 as Jungermanniatetragona. Seram: Akiyama 1986, 2009 both as Jungermanniatetragona, Váňa and Piippo 1989a as Jungermanniatetragona, Váňa 1991a as Jungermanniatetragona.

Solenostomatruncatum (Nees) R.M.Schust. ex Váňa et D.G.Long. MOLUCCAS: Halmahera: Váňa and Piippo 1989a as Jungermanniatruncata, Váňa 1991a as Jungermanniatruncata. Ambon: Váňa 1973, 1991a as Jungermanniatruncata, Váňa and Piippo 1989a as Jungermanniatruncata, Bapna and Kachroo 2000a as Jungermanniatruncata, Akiyama 2009 as Jungermanniatruncata. Seram: Akiyama 1986, 2009 both as Jungermanniatruncata, Váňa and Piippo 1989a as Jungermanniatruncata, Váňa 1991a as Jungermanniatruncata.

Spruceanthus Verd.

Spruceanthuspolymorphus (Sande Lac.) Verd. MOLUCCAS: Halmahera: Umagap 2019. Seram: Mizutani 1986a, Akiyama 2009.

Symphyogynopsis Grolle

Symphyogynopsisgottscheana (Mont. et Nees) Grolle. MOLUCCAS: Wigginton and Porley 2001, Schaumann et al. 2002. Seram: Grolle and Piippo 1986 as Symphyogynopsisfilicum, Grolle 1987, Enroth 1991, Furuki 2002 as Symphyogynopsisfilicum.

Syzygiella Spruce

Syzygiellaovalifolia Inoue. MOLUCCAS: Seram: Akiyama 1986, 2009, So and Grolle 2003.

Syzygiellasecurifolia (Nees) Inoue. MOLUCCAS: Halmahera: Sande Lacoste 1864 as Plagiochilavariegata, Schiffner 1898, 1900 as Syzygiellavariegata, Miller et al. 1983 as Syzygiellavariegata. Seram: Akiyama 1986, 2009.

Telaranea Spruce ex Schiffn.

Telaraneamajor (Herzog) J.J.Engel et G.L.Merr. MOLUCCAS: Schuster 2000a as Arachniopsismajor. Ambon: Grolle 1968b as Arachniopsismajor. Seram: Akiyama 1986, 2009 as Arachniopsismajor, von Konrat et al. 2014.

Telaraneatrisetosa (Steph.) Grolle. MOLUCCAS: Seram: Akiyama 1986, 2009.

Temnoma Mitt.

Temnomasetigerum (Lindenb.) R.M.Schust. MOLUCCAS: Seram: Grolle 1967, Akiyama 1986, 2009, Long and Grolle 1990, Bapna and Kachroo 2000a.

Thysananthus Lindenb.

Thysananthusappendiculatus Steph. MOLUCCAS: Ambon: Sukkharak 2015. Seram: Mizutani 1986a, Akiyama 2009.

ThysananthusconvolutusLindenb.var.convolutus. MOLUCCAS: Haerida et al. 2010, Siregar et al. 2014, 2017, 2020a, Siregar 2015, Ginting and Batubara 2019. Buru: Verdoorn 1934b, Eggers and Schäfer-Verwimp 1987. Ambon: Sukkharak 2011. Seram: Mizutani 1986a, Akiyama 2009.

Thysananthusfrauenfeldii Reichardt. MOLUCCAS: Thiers and Gradstein 1989 as Mastigolejeuneaundulata, Gradstein et al. 2002 as Mastigolejeuneaundulata, Sass-Gyarmati 2003 as Mastigolejeuneaundulata, Sukkharak 2011 as Thysananthusundulatus. Seram: Mizutani 1986a, 1986b both as Mastigolejeuneaundulata, Akiyama 2009 as Mastigolejeuneaundulata.

Thysananthusfruticosus (Lindenb. et Gottsche) Schiffn. MOLUCCAS: Mizutani 1977. Obi: Verdoorn 1934b. Banda: Sande Lacoste 1864 as Bryopterisfruticosa, Schiffner 1898, Hattori 1951. Seram: Sande Lacoste 1864 as Bryopterisfruticosa, Verdoorn 1934a, 1934b, Hattori 1951, Miller et al. 1983, Mizutani 1986a, 1987, Akiyama 2009.

Thysananthushumilis (Gottsche) Sukkharak et Gradst. MOLUCCAS: Mizutani 1977 as Mastigolejeuneahumilis. Ambon: Stephani 1912a as Mastigolejeuneahumilis, Verdoorn 1934a, 1934b both as Mastigolejeuneahumilis, Miller et al. 1983 as Mastigolejeuneahumilis, Awasthi and Udar 1984 as Mastigolejeuneahumilis, Bapna and Kachroo 2000b as Mastigolejeuneahumilisvar.humilis. Banda: Verdoorn 1934b as Mastigolejeuneahumilis, Bonner 1963a as Brachiolejeuneamolukkensis, Tixier 1966 as Mastigolejeuneahumilis, Bapna and Kachroo 2000b as Mastigolejeuneahumilis. Seram: LECTOTYPE of Brachiolejeuneamolukkensis [Bonner 1963a: 456], Awasthi and Udar 1984 as Mastigolejeuneahumilis, Mizutani 1986a, 1986b both as Mastigolejeuneahumilis, Bapna and Kachroo 2000b as Mastigolejeuneahumilis, Akiyama 2009 as Mastigolejeuneahumilis.

Thysananthusligulatus (Lehm. et Lindenb.) Sukkharak et Gradst. MOLUCCAS: Buru: Verdoorn 1934b as Mastigolejeunealigulata, Swanson and Miller 1969 as Mastigolejeunealigulata, Miller et al. 1983 as Mastigolejeunealigulata. Ambon: Mizutani 1986b as Mastigolejeunealigulata. Seram: Sande Lacoste 1864 as Phragmicomaligulata, Schiffner 1898 as Mastigolejeunealigulata, Swanson and Miller 1969 as Mastigolejeunealigulata, Miller et al. 1983 as Mastigolejeunealigulata, Mizutani 1986a, 1986b both as Mastigolejeunealigulata, Akiyama 2009 as Mastigolejeunealigulata, Sukkharak and Gradstein 2014 as Mastigolejeunealigulata.

Thysananthusrepletus (Taylor) Sukkharak et Gradst. MOLUCCAS: Ambon: Verdoorn 1934b as Mastigolejeuneaatypos, Grolle 1968b as Mastigolejeuneaatypos, Miller et al. 1983 as Mastigolejeuneaatypos. Seram: Verdoorn 1934b as Mastigolejeuneaatypos, Grolle 1968b as Mastigolejeuneaatypos, Miller et al. 1983 as Mastigolejeuneaatypos.

Thysananthusspathulistipus (Reinw., Blume et Nees) Lindenb. MOLUCCAS: Mizutani 1977, Haerida et al. 2010, Siregar et al. 2014, Siregar 2015, Siregar et al. 2017. Ambon: LECTOTYPE of Mastigolejeuneaamboinensis [to be designated], LECTOTYPE of Mastigolejeuneaamboinensisvar.paucidentata [to be designated], Sande Lacoste 1864 [Saparoea I], Schiffner 1890 as Mastigolejeuneaamboinensis, Schiffner 1890 as Mastigolejeuneaamboinensisvar.paucidentata, 1898, Sydow 1894 as Mastigolejeuneaamboinensis, Verdoorn 1934b [also Saparoea I], Swanson and Miller 1969, Miller et al. 1983, Bapna and Kachroo 2000b. Seram: Sande Lacoste 1864, Schiffner 1898, Verdoorn 1934a, 1934b, Grolle 1965, Swanson and Miller 1969, Mizutani 1986a, Akiyama 2009 as Thysananthus ‘spatulistipus’.

Thysananthustruncatus (Mizut.) Sukkharak et Gradst. MOLUCCAS: Seram: Mizutani 1986a, 1986b both as Mastigolejeuneatruncata, Pócs et al. 1994 as Mastigolejeuneatruncata, Akiyama 2009 as Mastigolejeuneatruncata, Sukkharak and Gradstein 2014 as Mastigolejeuneatruncata.

Thysananthusvirens Ångstr. MOLUCCAS: Haerida et al. 2010 as Mastigolejeuneavirens, Siregar et al. 2014, 2020a both as Mastigolejeuneavirens, Siregar 2015 as Mastigolejeuneavirens, Ginting and Batubara 2019 as Mastigolejeuneavirens, Siregar et al. 2020a as Mastigolejeuneavirens. Seram: Mizutani 1986a as Mastigolejeuneavirens, Mizutani 1986b as Mastigolejeuneavirens, Akiyama 2009 as Mastigolejeuneavirens.

Treubia K.I.Goebel

Treubiainsignis K.I.Goebel. MOLUCCAS: Seram: Akiyama 1986, 2009.

Trichocolea Dumort.

Trichocoleapluma (Reinw., Blume et Nees) Mont. MOLUCCAS: Buru: Herzog 1921 as Trichocoleastriolata. Ambon: Schiffner 1893b, 1898 both as Trichocoleatomentellavar.javanica, Katagiri et al. 2013. Seram: Akiyama 1986, 2009.

Trichocoleatomentella (Ehrh.) Dumort. MOLUCCAS: Morotai: New to Moluccas: Morotai, Sangowo, Exp. Kostermans) (BO13087)]. Ambon: Miller et al. 1983 as Trichocoleatomentella, Bapna and Kachroo 2000a. Note: we are not aware of any first hand report from Ambon but it is here confirmed for Moluccas.

Tricholepidozia (R.M.Schust.) E.D.Cooper

Tricholepidoziakogiana (Steph.) E.D.Cooper. MOLUCCAS: Seram: Akiyama 1986, 2009 both as Telaraneakogiana.

Tricholepidozianeesii (Lindenb.) E.D.Cooper. MOLUCCAS: Halmahera: Grolle 1966c as Telaraneaneesii, Pócs 1971 as Telaraneaneesii, Mizutani 1974, 1976 both as Telaraneaneesii, Engel 1978 as Telaraneaneesii, Kitagawa 1978 as Telaraneaneesii, Yamaguchi 1983 as Telaraneaneesii, Miller 1986 as Telaraneaneesii, Enroth 1991 as Telaraneaneesii, Engel and Smith Merrill 2004 as Telaraneaneesii. Seram: Piippo 1984b as Telaraneaneesii, Akiyama 1986, 2009 both as Telaraneaneesii.

Tuyamaella S.Hatt.

Tuyamaellaserratistipa S.Hatt. MOLUCCAS: Seram: Mizutani 1986a, Zhu and So 1998a, 2000, Akiyama 2009.

Wettsteinia Schiffn.

Wettsteiniainversa (Sande Lac.) Schiffn. MOLUCCAS: Seram: Akiyama 1986, 2009.

Zantenia (S.Hatt.) Váňa et J.J.Engel

Zanteniaborneensis (Herzog) Váňa et J.J.Engel. MOLUCCAS: Seram: Váňa and Piippo 1989b as Anastrophyllumborneense, Váňa 1991b as Anastrophyllumborneense.

Zanteniakarstenii (Schiffn.) Váňa et J.J.Engel. MOLUCCAS: Herzog 1926a as Anastrophyllumkarstenii. Ambon: LECTOTYPE of Anastrophyllumkarstenii [Bonner 1962b: 73], Schiffner 1893a, 1893b, 1898 all as Anastrophyllumkarstenii, Stephani 1901c as Anastrophyllumkarstenii, Bonner 1962b as Anastrophyllumkarstenii, Hattori 1966 as Anastrophyllumkarstenii.

Zoopsis Hook.f. ex Gottsche, Lindenb. et Nees

Zoopsisliukiuensis Horik. MOLUCCAS: Grolle and Piippo 1984. Ambon: Grolle 1968b, Schuster 1969, Miller et al. 1983, Schuster 1999, Engel and Glenny 2008. Seram: Akiyama 1986, 2009.

Zoopsissetigera K.I.Goebel. MOLUCCAS: Ambon: LECTOTYPE [Grolle and Piippo 1984: 303 as “neotype”], Grolle and Piippo 1984, Schuster 1999, Engel and Glenny 2008. Seram: Akiyama 1986, 2009.

Names not referred to any accepted taxon

We are not able to refer the following published names to any accepted taxon.

Phaeoceros Prosk.

Phaeocerosvelutinus J.Haseg. ex H.Akiyama nom. inval.MOLUCCAS: Ambon: ORIGINAL MATERIAL. Seram: Akiyama 1986. Note: we have not found the name Phaeocerosvelutinus anywhere else than in Akiyama (1986) listing it from Moluccas, and we do not know where to refer it.

Jungermannia L.

Jungermanniagigantea β laxior (Nees) Lindenb. MOLUCCAS: Ternate/Tidore: SYNTYPE of Jungermanniagigantea β laxior [lectotype to be designated], ORIGINAL MATERIAL of Plagiochilatrapezoidea γ major, Nees 1830, 1831 both as Jungermanniagigantea β laxior, Lindenberg 1840 as Plagiochilatrapezoidea γ ‘maior’, Schiffner 1898, 1900 as Plagiochilatrapezoideavar.major, Geissler and Bischler 1987 as Jungermanniagigantea β laxior. Note: this name does have a confusing history and before someone really select and identify a lectotype, it is impossible to know what taxon it represents

Jungermanniatamarisci L. var. β minus-ramosa Schwägr. nom. inval.MOLUCCAS: ORIGINAL MATERIAL, Gaudichaud 1828. Note: Frullaniatamarisci is a mainly Eurasian taxon and this name must refer to a tropical taxon

Plagiochila (Dumort.) Dumort.

Plagiochilaopposita (Reinw., Blume et Nees) Lindenb. β falcata (Nees) Schiffn. MOLUCCAS: Ternate/Tidore: Sande Lacoste 1856b, Schiffner 1898. Note: Plagiochilaopposita is now Chiastocaulonoppositum but we are not sure if unranked β falcata also belong to that species and is worth recognizing.

Taxa doubtfully occurring in Moluccas

We doubt the identification of the following taxa or we are not aware of any first-hand report from Moluccas.

Acromastigum A.Evans

Acromastigumechinatum (Gottsche) A.Evans. MOLUCCAS: Ambon: Stephani 1909a as Mastigobryumechinatum, Bonner 1962b, Swanson and Miller 1969, Miller et al. 1983. Note: we are not aware of any first hand report from Moluccas although it is reported to occur on neighbouring islands.

Bazzania Gray

Bazzaniaceylanica (Mitt.) Steph. MOLUCCAS: Ambon: Stephani 1902 as Mastigobryumceylanicum. Note: we are not aware of any first hand report from Moluccas although it is reported to occur on neighbouring islands.

Bazzaniajaponica (Sande Lac.) Lindb. MOLUCCAS: Siregar and Pasaribu 2019. Note: we are not aware of any first hand report from Moluccas although it is reported to occur on neighbouring islands.

Calypogeia Raddi

Calypogeiatosana (Steph.) Steph. MOLUCCAS: Halmahera: Umagap 2019. Note: a Sino-Himalayan taxon that seems unlikely to occur in the tropics.

Chiloscyphus Corda

Chiloscyphusernstianus Steph. MOLUCCAS: Seram: Akiyama 1986, 2009 both as Lophocoleaaff.ernstiana. Note: we are not aware of any positively identified report from Moluccas. It is otherwise only known from the type from Sumatra and if worth recognizing it certainly belongs to some other genus.

Cololejeunea (Spruce) Steph.

Cololejeuneacordiflora Steph. MOLUCCAS: Ambon: Tixier 1962 as Cololejeuneakarstenii. Note: we are not aware of any first hand report from Moluccas although it is reported to occur on neighbouring islands.

Cololejeuneaverrucosa Steph. MOLUCCAS: Miller 1968, Pócs and Ninh 2012, Pócs 2012. Ambon: Miller et al. 1983. Note: we are not aware of any first hand report from Moluccas although it is reported to occur on neighbouring islands.

Cryptolophocolea L.Söderstr., Crand.-Stotl., Stotler et Váňa

Cryptolophocoleaciliolata (Nees) L.Söderstr., Crand.-Stotl., Stotler et Váňa. MOLUCCAS: Seram: Akiyama 1986, 2009 both as Lophocoleaaff.ciliolata. Note: we are not aware of any positively identified report from Moluccas although it is reported to occur on neighbouring islands.

Leptolejeunea (Spruce) Steph.

Leptolejeuneabalansae Steph. MOLUCCAS: Ambon: Jovet-Ast 1958. Note: we are not aware of any first hand report from Moluccas although it is reported to occur on neighbouring islands.

Leptolejeuneasubacuta Steph. ex A.Evans. MOLUCCAS: Inoue and Miller 1965, Miller 1968, Swanson and Miller 1969, Miller et al. 1983, Bapna and Kachroo 2000b. Note: we are not aware of any first hand report from Moluccas although it is reported to occur on neighbouring islands.

Mastigophora Nees

Mastigophorawoodsii (Hook.) Nees. MOLUCCAS: Seram: LECTOTYPE of Mastigophoraramentifissa [to be designated], Herzog 1926a, 1926b as Mastigophoraramentifissa, Inoue 1971. Note: Inoue (1971) synonymized Mastigophoraramentifissa with Mastigophorawoodsii but doubted the origin of the type of the former as Mastigophorawoodsii is unlikely to occur in the tropics.

Plagiochila (Dumort.) Dumort.

Plagiochilagracilis Lindenb. et Gottsche. MOLUCCAS: Miller et al. 1983. Note: we are not aware of any first hand report from Moluccas although it is reported to occur on neighbouring islands.

Radula Dumort.

Radulaamentulosa Mitt. MOLUCCAS: Seram: Yamada 1999. Note: we are not aware of any first-hand report from Moluccas.

Schistochila Dumort.

Schistochilaschultzei Steph. MOLUCCAS: Seram: Akiyama 1986, 2009 both as Schistochilaaff.schultzei. Note: we are not aware of any positively identified specimen from Moluccas although it is reported from neighboring islands.

Spruceanthus Verd.

Spruceanthussemirepandus (Nees) Verd. MOLUCCAS: Haerida et al. 2010. Note: we are not aware of any positively identified specimen from Moluccas although it is reported from neighboring islands.

Thysananthus Lindenb.

Thysananthusauriculatus (Wilson et Hook.) Sukkharak et Gradst. MOLUCCAS: Haerida et al. 2010 as Mastigolejeuneaauriculata, Singh et al. 2010 as Mastigolejeuneaauriculata. Note: we are not aware of any positively identified specimen from Moluccas although it is reported from neighboring islands.

Taxa reported but rejected from Moluccas

The following taxa are reported to occur in Moluccas but the reports are rejected either earlier or here as misidentified or highly unlikely to occur in the area.

Cheilolejeunea (Spruce) Steph.

Cheilolejeuneamariana (Gottsche) B.M.Thiers et Gradst. MOLUCCAS: Seram: Mizutani 1986a as Spruceanthusmarianus, Akiyama 2009 as Spruceanthusmarianus. Note: a Pacific species frequently, but erroneously, reported from SE Asia and many other areas. Reports should mainly be referred to Archilejeuneaplaniuscula.

Chiastocaulon Carl

Chiastocaulonfimbriatum (Mitt.) S.D.F.Patzak, M.A.M.Renner, Schäf.-Verw. et Heinrichs. MOLUCCAS: Seram: Akiyama 1986, 2009 both as Plagiochilionaff.fimbriatum. Note: a Sino-Himalayan taxon very unlikely to occur in the tropics.

Conocephalum Hill

Conocephalumconicum (L.) Dumort. MOLUCCAS: Ambon: Sopacua et al. 2020 as ‘Conocephelum’ conicum [Saparua I]. Note: presumably a mainly European taxon reported sensu lato from many other places. It is unlikely that the genus Conocephalum occurs in the tropics, but if it is a Conocephalum sp. it is probably more related to Conocephalumsalebrosum than to Conocephalumconicum.

Cryptolophocolea L.Söderstr., Crand.-Stotl., Stotler et Váňa

Cryptolophocoleaconnata (Sw.) L.Söderstr. et Váňa. MOLUCCAS: Gaudichaud 1828 as Jungermanniaconnata. Note: an American species and this old record must be rejected as either mis-identified or mis-labelled.

Dendroceros Nees

Dendrocerosexalatus Steph. MOLUCCAS: Ambon: LECTOTYPE [Hasegawa 1980: 302], Stephani 1909b, 1917, Bonner 1965. Note: Stephani erroneously reported the type locality as “Amboina” but as this is a Brazilian species that must be due to mislabeling (cf. Hasegawa 1980).

Heteroscyphus Schiffn.

Heteroscyphuscaledonicus (Steph.) Schiffn. MOLUCCAS: Ambon: Miller et al. 1983. Note a Pacific species possibly endemic to New Caledonia.

Lepicolea Dumort.

Lepicoleaochroleuca (Spreng.) Spruce. MOLUCCAS: Seram: Herzog 1921. Note: an Afro-American species only once reported from SE Asia. This report must be an error.

Lepicoleascolopendra (Hook.) Dumort. ex Trevis. MOLUCCAS: Seram: Herzog 1926b. Note: an Australian/New Zealandian species frequently, but erroneously, reported from many other places.

Lepidozia (Dumort.) Dumort.

Lepidoziamicrophylla (Hook.) Lindenb. MOLUCCAS: Seram: Siregar and Pasaribu 2019. Note: a species endemic to New Zealand (cf. Engel and Schuster 2001) and all reports from other areas must be rejected.

Lepidoziarigida Steph. MOLUCCAS: Buru: ORIGINAL MATERIAL of Lepidoziarigidaf.minor, Herzog 1921 as Lepidoziarigidaf.minor, Herzog 1926a as Lepidoziarigidavar.minor. Note: a species from Vanuatu. Herzog’s material must belong to another species.

Plagiochila (Dumort.) Dumort.

Plagiochilagigantea Lindenb. MOLUCCAS: Ternate/Tidore: Nees 1830 as Jungermanniagigantea. Note: a New Zealand taxon.

Plagiochilatrapezoidea Lindenb. MOLUCCAS: Ternate/Tidore: Reinwardt et al. 1824 as Jungermanniasimplex Web, Schiffner 1900, Bapna and Kachroo 2000b. Note: Inoue (1984) designated the only specimen from Java of Jungermanniasimplex Web. that he could locate as a neotype of Plagiochilatrapezoidea and Plagiochilatrapezoidea β tenera is described as based on Jungermanniasimplex Web. However, Inoue (1984) reject the report of β tenera in Gottsche et al. (1844; all other reports probably based on this) as Plagiochilaciliata. The occurrence of Plagiochilatrapezoidea in Moluccas is therefore not confirmed and reports should be rejected following Inoue, until confirmed, although it occurs in most neighbouring islands.

Podomitrium Mitt.

Podomitriumphyllanthus (Hook.) Mitt. MOLUCCAS: Ambon: Schiffner 1890, 1898 as ‘Hymenophytum’ phyllanthus, Sydow 1894. Note: this is an Australian/New Zealandian species.

Porella L.

Porellanitens (Steph.) S.Hatt. MOLUCCAS: Long and Grolle 1990. Note: the second-hand reports from Malesia in Long and Grolle (1990) of this Sino-Himalayan species is due to supposed, but rejected, synonymy with Porellajavanica (cf. also Hattori 1978).

Pycnolejeunea (Spruce) Schiffn.

Pycnolejeuneaschwaneckei (Steph.) Schiffn. ex P.Syd. MOLUCCAS: Ambon: Schiffner 1890, Sydow 1894 as Pycnolejeunea ‘schwaneckii’. Note: this is a neotropical species.

Radula Dumort.

Radulapinnulata Mitt. MOLUCCAS: Miller et al. 1983. Note: Yamada (2002) rejected all reports from Moluccas.

Riccardia Gray

Riccardialatifrons (Lindb.) Lindb. MOLUCCAS: Ambon: Schiffner 1890 as Aneuralatifrons, Sydow 1894 as Aneuralatifrons. Note: a mainly boreal species and all reports from tropical areas should be rejected.

Schistochila Dumort.

Schistochilaneesii (Mont.) Lindb. MOLUCCAS: Ambon: Schiffner 1893b, 1898, Buch 1939, Hässel 1976 as Paraschistochilaneesii. Note: an African species rejected from Moluccas by So (2003a).

Symbiezidium Trevis.

Symbiezidiumtransversale (Sw.) Trevis. MOLUCCAS: Gaudichaud 1828 as Jungermanniatransversalis. Note: an American species.

Zoopsis Hook.f. ex Gottsche, Lindenb. et Nees

Zoopsisargentea (Hook.f. et Taylor) Gottsche, Lindenb. et Nees. MOLUCCAS: Ambon: Stephani 1908a, Del Rosario 1975b, Miller et al. 1983. Note: an Australian/New Zealandian species.

Zoopsissetulosa Leitg. MOLUCCAS: Miller et al. 1983 as Zoopsisrigida. Ambon: Schiffner 1893b as Cephaloziasetulosa, 1898, Stephani 1908b, Hattori 1951, Hodgson 1965, Schuster 1969. Note: an Australian/New Zealandian/ New Caledonian species rejected from Asia as Zoopsissetigera by Grolle and Piippo (1984).

Synonyms

Acrolejeuneadensifolia (Schiffn.) Schiffn. ex P.Syd. nom. inval. = Schiffneriolejeuneapulopenangensis

Acrolejeuneadensifolia Schiffn. nom. inval. = Schiffneriolejeuneapulopenangensis

Acrolejeuneahaskarliana (Gottsche) Schiffn. ≡ Schiffneriolejeuneatumidavar.haskarliana

Acrolejeuneapulopenangensis (Gottsche) Schiffn. ≡ Schiffneriolejeuneapulopenangensis

Acrolejeunearostrata Schiffn. nom. inval. = Acrolejeuneapycnocladasubsp.pycnoclada

Acrolejeunearostrata var. α minor Schiffn. nom. inval. = Acrolejeuneapycnocladasubsp.pycnoclada

Acrolejeunearostrata var. β major Schiffn. nom. inval. = Acrolejeuneapycnocladasubsp.pycnoclada

Acrolejeuneasecurifolia (Nees) Steph. nom. inval. ≡ Acrolejeuneasecurifolia

Acromastigumbidenticulatum A.Evans = Acromastigumbancanum

Anastrophyllumbidens (Reinw., Blume et Nees) Steph. ≡ Schizophyllopsisbidens

Anastrophyllumborneense Herzog ≡ Zanteniaborneensis

Anastrophyllumcontractum (Reinw., Blume et Nees) Steph. ≡ Cuspidatulacontracta

Anastrophyllumkarstenii Schiffn. ≡ Zanteniakarstenii

Anastrophyllumrevolutum Steph. ≡ Anastrophyllopsisrevoluta

Aneuraalbomarginata Steph. ≡ Riccardiaalbomarginata

Aneurakarstenii Steph. ≡ Riccardiakarstenii

Aneuralatifrons Lindb. ≡ Riccardialatifrons

Aneurapinguisf.normalis Schiffn. nom. inval. = Aneurapinguis

Anomacaulisflaccidus (Steph.) Grolle ≡ Cuspidatulaflaccida

Anthocerosamboinensis Schiffn. ≡ Foliocerosamboinensis

Anthocerosappendiculatus Steph. ≡ Foliocerosappendiculatus

Anthocerosglandulosus Lehm. et Lindenb. ≡ Foliocerosglandulosus

Anthocerosgrandis Ångstr. = Megacerosflagellaris

Arachniopsismajor Herzog ≡ Telaraneamajor

Aspiromitusamboinensis (Schiffn.) Steph. ≡ Foliocerosamboinensis

Bazzaniaaustralis (Mont.) Trevis. = Bazzaniatridens

Bazzaniaconcinna (De Not.) Trevis. = Bazzaniaintermedia

Bazzaniadivaricata (Nees) Trevis. = Acromastigumdivaricatum

Bazzaniaechinatiformis (De Not.) Trevis. ≡ Acromastigumechinatiforme

Bazzaniaintermedia auct. = Bazzaniatridensvar.tridens

Bazzanialongidens (Steph.) Schiffn. = Bazzaniaconophylla

Bazzaniaoshimensis (Steph.) Horik. = Bazzaniatridens

Brachiolejeuneamolukkensis Steph. = Thysananthushumilis

Bryopterisfruticosa Lindenb. et Gottsche ≡ Thysananthusfruticosus

Calypogeiafusca (Lehm.) Steph. ≡ Mniolomafuscum

Campylolejeuneaciliatilobula (Schiffn.) S.Hatt. = Cololejeuneainflectens

Campylolejeuneainflectens (Mitt.) Mizut. ≡ Cololejeuneainflectens

Caudalejeuneacircinata Steph. = Caudalejeuneacristiloba

Caudalejeunearecurvistipula (Gottsche) Schiffn. = Caudalejeuneareniloba

Cephaloziasetulosa (Leitg.) Spruce ≡ Zoopsissetulosa

Chandonanthusgracilis Herzog = Plicanthushirtellus

Chandonanthushirtellus (F.Weber) Mitt. ≡ Plicanthushirtellus

Cheilolejeuneaimbricata (Nees) S.Hatt. = Cheilolejeuneatrapezia

Cheilolejeuneameyeniana (Nees, Lindenb. et Gottsche) R.M.Schust. et Kachroo = Cheilolejeuneatrapezia

Chiloscyphusamboinensis (Schiffn.) Schiffn. ≡ Heteroscyphusamboinensis

Chiloscyphusaselliformis (Reinw., Blume et Nees) Nees ≡ Heteroscyphusaselliformis

Chiloscyphuscoalitus (Hook.) Nees ≡ Heteroscyphuscoalitus

Chiloscyphusdecurrens (Reinw., Blume et Nees) Nees = Heteroscyphussplendens

Chiloscyphusdecurrensvar.amboinensis (Schiffn.) Schiffn. ex Bonner nom. inval. ≡ Heteroscyphusamboinensis

Chiloscyphusendlicherianus δ amboinensis Schiffn. ≡ Heteroscyphusamboinensis

Chiloscyphusfalcifolius Steph. = Acroscyphellatjiwideiensis

Chiloscyphusgranulatus Schiffn. = Saccogynidiumrigidulum

Clasmatocoleatjiwideiensis (Sande Lac.) Grolle ≡ Acroscyphellatjiwideiensis

Cololejeuneaciliatilobula (Schiffn.) Schiffn. = Cololejeuneainflectens

Cololejeuneacrenulata (Herzog) Benedix nom. inval. = Cololejeuneaangustiflora

Cololejeuneafalcatoides Benedix = Cololejeuneafalcata

Cololejeuneagoebelii (Gottsche ex Schiffn.) Schiffn. = Cololejeuneatrichomanis

Cololejeuneajavanica (Steph.) Mizut. = Cololejeuneaangustiflora

Cololejeuneakarstenii (Steph.) Benedix = Cololejeuneacordiflora

Cololejeuneanymannii (Steph.) Benedix = Cololejeuneaobliqua

Cololejeuneapeculiaris (Herzog) Benedix = Cololejeuneainflectens

Cololejeuneayulensis (Steph.) Benedix = Cololejeuneaaequabilis

Coluraparadoxa (Schiffn.) Steph. ≡ Calatholejeuneaparadoxa

Colurolejeuneakarstenii (K.I.Goebel) Steph. ≡ Colurakarstenii

Colurolejeuneakarsteniivar.latifolia (Schiffn.) Schiffn. nom. inval. = Colurakarstenii

Colurolejeuneaparadoxa (Schiffn.) Schiffn. ≡ Calatholejeuneaparadoxa

Colurolejeuneasuperba (Mont.) Steph. ≡ Colurasuperba

Colurolejeuneasuperba Schiffn. nom. inval. = Coluraamboinensis

Colurolejeuneasuperba var. α typica Schiffn. nom. inval. = Coluraamboinensis

Dendroceroskarstenii Schiffn. ex Steph. = Dendrocerosacutilobus

Drepanolejeuneadactylophora (Nees, Lindenb. et Gottsche) Schiffn. ≡ Drepanolejeuneadactylophora

Drepanolejeuneadentata Steph. ≡ Lejeuneastenodentata

Drepanolejeuneafilicuspis Steph. ≡ Microlejeuneafilicuspis

Drepanolejeunealevicornuavar.longicornua Herzog ≡ Drepanolejeunealongicornua

Drepanolejeuneamicholitzii Steph. = Drepanolejeuneapentadactyla

Drepanolejeuneamicholitziivar.dactylophoroides Herzog ≡ Drepanolejeuneapentadactylavar.dactylophoroides

Drepanolejeuneamicholitziivar.genuina Herzog nom. illeg. = Drepanolejeuneapentadactyla

Drepanolejeuneateneravar.genuina Herzog nom. illeg. = Drepanolejeuneatenera

Drepanolejeuneaternatensis (Gottsche) Steph. nom. inval. ≡ Drepanolejeuneaternatensis

Eucalyxcomata (Nees) Verd. ≡ Solenostomacomatum

Eulejeuneafissistipula Steph. ≡ Lejeuneafissistipula

Eulejeuneaparallela (Schiffn.) Schiffn. = Metalejeuneacucullata

Euosmolejeuneaheterophylla (Sande Lac.) Schiffn. = Cheilolejeuneatrifaria

Euosmolejeuneaintegristipula Steph. = Cheilolejeunealindenbergii

Euosmolejeuneatrifaria (Reinw., Blume et Nees) Steph. ≡ Cheilolejeuneatrifaria

Frullaniaamboinensis Schiffn. = Frullaniaintermediasubsp.intermedia

Frullaniabillardiereana Nees et Mont. = Frullaniaintermediasubsp.intermedia

Frullaniadapitana Steph. = Frullanianodulosa

Frullaniaericoidesvar.planescens (Verd.) S.Hatt. = Frullaniaericoides

Frullaniagracilisvar.lacerifolia (Steph.) Verd. = Frullaniagracilis

Frullaniaintermediaf.billardiereana (Nees et Mont.) Verd. = Frullaniaintermediasubsp.intermedia

Frullaniaintermediavar.amboinensis (Schiffn.) Verd. = Frullaniaintermediasubsp.intermedia

Frullaniakarstenii Schiffn. = Frullaniaapiculata var. goebelii

Frullanialacerifolia Steph. = Frullaniagracilis

Frullanianodulosaf.dapitana (Steph.) Verd. = Frullanianodulosa

Frullanianodulosaf.irreflexa Verd. = Frullanianodulosa

Frullanianodulosa β plana Schiffn. = Frullanianodulosa

Frullaniapicta Steph. = Frullaniatrichodes

Frullaniareplicata (Nees) Mont. = Frullanianodulosa

Frullaniarugosa Mitt. = Frullaniagrandistipula

Frullaniaserrataf.crispulo-dentata Verd. = Frullaniaserratavar.serrata

Frullaniasquarrosa (Mont.) Nees = Frullaniaericoides

Frullaniasquarrosaf.campanuloides Verd. = Frullaniaericoides

Frullaniasquarrosavar.planescens Verd. = Frullaniaericoides

Frullaniatenuicaulis Mitt. = Frullaniatrichodes

Frullaniathuillieri Nees = Frullanianodulosa

Gamoscyphusaselliformis (Reinw., Blume et Nees) Trevis. ≡ Heteroscyphusaselliformis

Gottscheaaligera (Nees et Blume) Nees ≡ Schistochilaaligera

Gottscheablumei (Nees) Nees ≡ Schistochilablumei

Harpalejeuneaexigua Steph. = Lejeuneaexilis

Harpalejeuneafilicuspis (Steph.) Mizut. ≡ Microlejeuneafilicuspis

Herbertusarmitanus (Steph.) H.A.Mill. = Herbertussendtneri

Herbertusdivaricatus (Herzog) H.A.Mill. = Herbertussendtneri

Herbertusjavanicus (Steph.) H.A.Mill. = Herbertusramosus

Herbertusjavanicus (Steph.) Schiffn. nom. inval. = Herbertusramosus

Heteroscyphusdecurrens (Nees) Schiffn. = Heteroscyphussplendens

Hygrolejeuneaamboinensis (Schiffn.) Schiffn. nom. inval. = Lejeuneasordida

Hygrolejeuneaamboinensis (Schiffn.) Schiffn. ex P.Syd. = Lejeuneasordida

Hygrolejeuneadecurvifolia Steph. = Lepidolejeuneabidentula

Hygrolejeuneadiscreta (Lindenb.) Schiffn. ≡ Lejeuneadiscreta

Hygrolejeuneadiscreta (Lindenb.) Steph. nom. inval. ≡ Lejeuneadiscreta

Hygrolejeuneaparkinsonii Steph. = Lejeuneasordida

Hygrolejeuneasordida (Nees) Schiffn. ≡ Lejeuneasordida

Hygrolejeuneasordida (Nees) Steph. nom. inval. ≡ Lejeuneasordida

Hygrolejeuneaumbilicata (Nees) Steph. nom. inval. ≡ Lejeuneaumbilicata

Hymenophytonmalaccense Steph. ≡ Podomitriummalaccense

Hymenophytonphyllanthus (Hook.) Steph. ≡ Podomitriumphyllanthus

Jamesoniellacontracta (Reinw., Blume et Nees) N.Kitag. ≡ Cuspidatulacontracta

Jamesoniellaflexicaulis (Nees) Schiffn. ≡ Cuspidatulaflexicaulis

Jamesoniellaflexicaulis var. β microphylla (Nees) Schiffn. = Gottscheliaschizopleura

Jamesoniellamicrophylla (Nees) Schiffn. = Gottscheliaschizopleura

Jamesoniellaovifolia (Schiffn.) Schiffn. = Denotarisialinguifolia

Jungermanniaariadne Taylor ≡ Solenostomaariadne

Jungermanniaaselliformis Reinw., Blume et Nees ≡ Heteroscyphusaselliformis

Jungermanniacomata Nees ≡ Solenostomacomatum

Jungermanniaconnata Sw. ≡ Cryptolophocoleaconnata

Jungermanniacontracta Reinw., Blume et Nees ≡ Cuspidatulacontracta

Jungermanniaflexicaulis Nees ≡ Cuspidatulaflexicaulis

Jungermanniaflexicaulis β microphylla Gottsche, Lindenb. et Nees = Gottscheliaschizopleura

Jungermanniagigantea Hook. nom. illeg. ≡ Plagiochilagigantea

Jungermanniagigantea β laxior Nees ≡ Plagiochilagigantealaxior

Jungermanniaindica Nees = Lejeuneaflavasubsp.orientalis

Jungermanniaopposita Reinw., Blume et Nees ≡ Chiastocaulonoppositum

Jungermanniaovifolia Schiffn. = Denotarisialinguifolia

Jungermanniasimplex Weber nom. illeg. = Plagiochilatrapezoidea

Jungermanniatetragona Lindenb. ≡ Solenostomatetragonum

Jungermanniatransversalis Sw. ≡ Symbiezidiumtransversale

Jungermanniatruncata Nees ≡ Solenostomatruncatum

Jungermanniavaginata Sw. ≡ Frullaniavaginata

Lejeuneaapplanata (Reinw., Blume et Nees) Nees ≡ Lopholejeuneaapplanata

Lejeuneacatanduana (Steph.) H.A.Mill., Bonner et Bischl. = Lejeuneaanisophylla

Lejeuneachalmersii (Steph.) Mizut. = Lejeuneamicroloba

Lejeuneaconnivens Gottsche = Cheilolejeuneaceylanica

Lejeuneacucullata (Reinw., Blume et Nees) Nees ≡ Metalejeuneacucullata

Lejeuneahaskarliana (Gottsche) Spruce nom. illeg. ≡ Schiffneriolejeuneatumidavar.haskarliana

Lejeuneaheterophylla Sande Lac. = Cheilolejeuneatrifaria

Lejeuneamicrostipula (Steph.) Prantl = Lejeuneaexilis

Lejeuneaparadoxa Schiffn. ≡ Calatholejeuneaparadoxa

Lejeuneapusilla (K.I.Goebel) Schiffn. nom. illeg. = Cololejeuneametzgeriopsis

Lejeunearecurvistipula Gottsche = Caudalejeuneareniloba

Lejeunearostrata var. α minor Schiffn. ex P.Geissler et Bischl. nom. inval. = Acrolejeuneapycnocladasubsp.pycnoclada

Lejeuneasagrana (Mont.) Gottsche, Lindenb. et Nees = Lopholejeuneasubfusca

Lejeuneasagranavar.dentistipula Schiffn. = Caudalejeuneareniloba

Lejeuneasuperba var. α typica Schiffn. nom. inval. = Colurasuperba

Lejeuneaternatensis Gottsche ≡ Drepanolejeuneaternatensis

Lejeuneatrapezia (Nees) Nees ≡ Cheilolejeuneatrapezia

Lejeuneatrifaria (Reinw., Blume et Nees) Nees ≡ Cheilolejeuneatrifaria

Lepidoziapapulosa Steph. ≡ Neolepidoziapapulosa

Lepidoziarigidaf.minor (Steph.) Herzog nom. inval. = Lepidoziarigida

Lepidoziarigidavar.minor (Steph.) Herzog nom. inval. = Lepidoziarigida

Lepidoziawallichiana Gottsche ≡ Neolepidoziawallichiana

Leptolejeuneafoliicola (Horik.) R.M.Schust. nom. illeg. ≡ Drepanolejeuneafoliicola

Leptolejeuneaschiffnerif.angustifolia Herzog nom. inval. = Leptolejeuneaschiffneri

Leptolejeuneaschiffnerivar.genuina Herzog nom. inval. = Leptolejeuneaschiffneri

Leptolejeuneaschiffneri (Steph. ex Schiffn.) Steph. = Leptolejeuneaschiffneri

Lobatiriccardialobata (Schiffn.) Furuki = Lobatiriccardiacoronopus

Lophocoleaciliolata (Nees) Gottsche ≡ Cryptolophocoleaciliolata

Lophocoleaernstiana (Steph.) N.Kitag. ≡ Chiloscyphusernstianus

Lophocoleagiulianettii Steph. = Cryptolophocoleacostata

Lopholejeuneadentistipula (Schiffn.) Schiffn. = Caudalejeuneareniloba

Lopholejeuneaeulopha (Taylor) Steph. nom. inval. ≡ Lopholejeuneaeulopha

Lopholejeuneajavanica (Nees) Schiffn. = Lopholejeuneanigricans

Lopholejeuneapyriflora Steph. = Lopholejeuneasubfusca

Lopholejeuneasagrana (Mont.) Steph. = Lopholejeuneasubfusca

Lopholejeuneasagranavar.dentistipula (Schiffn.) Schiffn. ex P.Syd. = Caudalejeuneareniloba

Lopholejeuneasagranavar.dentistipula Schiffn. nom. inval. = Caudalejeuneareniloba

Lopholejeuneasagrana var. β subfusca (Nees) Schiffn. ≡ Lopholejeuneasubfusca

Madothecaacutifolia Lehm. et Lindenb. ≡ Porellaacutifolia

Madothecacrenilobula Herzog = Porellajavanica

Marchantiaamboinensis Nees et Mont. = Marchantiaemarginatasubsp.emarginata

Mastigobryumamboinense Steph. = Bazzaniatridens

Mastigobryumasymmetricum Steph. ≡ Bazzaniaasymmetrica

Mastigobryumceylanicum Mitt. ≡ Bazzaniaceylanica

Mastigobryumconcinnum De Not. = Bazzaniaintermedia

Mastigobryumcrassitextum (Steph.) Steph. ≡ Bazzaniacrassitexta

Mastigobryumdeningeri Herzog = Bazzanialongicaulis

Mastigobryumechinatiforme De Not. ≡ Acromastigumechinatiforme

Mastigobryumechinatum Gottsche ≡ Acromastigumechinatum

Mastigobryumfallax Sande Lac. ≡ Bazzaniafallax

Mastigobryumhorridulum (Schiffn.) Steph. ≡ Bazzaniahorridula

Mastigobryuminsigne De Not. ≡ Bazzaniainsignis

Mastigobryumintermedium Gottsche et Lindenb. ≡ Bazzaniaintermedia

Mastigobryumirregulare Steph. ≡ Bazzaniairregularis

Mastigobryumjavanicum Sande Lac. ≡ Bazzaniajavanica

Mastigobryumlongidens Steph. = Bazzaniaconophylla

Mastigobryummerrillanum Steph. ≡ Bazzaniamerrillana

Mastigobryumpectinatum (Nees) Lindenb. et Gottsche = Bazzaniapectinata

Mastigobryumstresemannii Herzog ≡ Bazzaniastresemannii

Mastigobryumsubtile Sande Lac. ≡ Bazzaniasubtilis

Mastigobryumtridens (Reinw., Blume et Nees) Dumort. ≡ Bazzaniatridens

Mastigobryumuncigerum (Reinw., Blume et Nees) Nees ≡ Bazzaniauncigera

Mastigobryumvittatum Gottsche ≡ Bazzaniavittata

Mastigobryumwallichianum Lindenb. ≡ Bazzaniawallichiana

Mastigolejeuneaamboinensis (Schiffn.) Schiffn. ex P.Syd. = Thysananthusspathulistipus

Mastigolejeuneaamboinensis Schiffn. nom. inval. = Thysananthusspathulistipus

Mastigolejeuneaamboinensis var. β paucidentata Schiffn. nom. inval. = Thysananthusspathulistipus

Mastigolejeuneaatypos Schiffn. ex Sydow = Thysananthusrepletus

Mastigolejeuneaauriculata (Wilson et Hook.) Steph. ≡ Thysananthusauriculatus

Mastigolejeuneahumilis (Gottsche) Schiffn. ≡ Thysananthushumilis

Mastigolejeunealigulata (Lehm. et Lindenb.) Schiffn. ≡ Thysananthusligulatus

Mastigolejeuneatruncata Mizut. ≡ Thysananthustruncatus

Mastigolejeuneaundulata Gradst. et Grolle = Thysananthusfrauenfeldii

Mastigolejeuneavirens (Ångstr.) Steph. ≡ Thysananthusvirens

Mastigophoradiclados var. β scorpioides (Reinw., Blume et Nees) Schiffn. = Mastigophoradiclados

Mastigophoraramentifissa Herzog = Mastigophorawoodsii

Mastigophoravrieseana (Sande Lac.) Schiffn. = Mastigophoradiclados

Megacerosgrandis (Ångstr.) Steph. = Megacerosflagellaris

Metahygrobiellastolonacea (Herzog) R.M.Schust. ≡ Cephaloziastolonacea

Metzgeriadecipiens (C.Massal.) Schiffn. = Metzgeriaciliata

Metzgeriahamatiformis Schiffn. = Metzgerialeptoneuravar.leptoneura

Metzgeriopsispusilla K.I.Goebel = Cololejeuneametzgeriopsis

Microlejeuneamicrostipula Steph. = Lejeuneaexilis

Microlejeuneaparallela Schiffn. = Metalejeuneacucullata

Microlejeuneaternatensis (Gottsche) Herzog nom. inval. ≡ Drepanolejeuneaternatensis

Microlepidoziagonyotricha (Sande Lac.) Del Ros. ≡ Kurziagonyotricha

Nardiaariadne (Taylor) Schiffn. ≡ Solenostomaariadne

Nardiacomata (Nees) Schiffn. ≡ Solenostomacomatum

Noguchiaopposita (Reinw., Blume et Nees) Inoue ≡ Chiastocaulonoppositum

Omphalanthusumbilicatus (Nees) Nees ≡ Lejeuneaumbilicata

Otolejeuneasemperiana (Steph.) Grolle ≡ Allorgellasemperiana

Paraschistochilaneesii (Mont.) R.M.Schust. ≡ Schistochilaneesii

Phaeoceroshirticalyx (Steph.) J.Haseg. ≡ Phaeomegaceroshirticalyx

Phaeoceroslaevissubsp.carolinianus (Michx.) Prosk. ≡ Phaeoceroscarolinianus

Phaeocerospolyandrus (Steph.) J.Haseg. = Phaeomegaceroshirticalyx

Phragmicomafertilis (Reinw., Blume et Nees) Nees ex Mont. ≡ Acrolejeuneafertilis

Phragmicomaligulata (Lehm. et Lindenb.) Gottsche, Lindenb. et Nees ≡ Thysananthusligulatus

Phragmicomareniloba Gottsche ≡ Caudalejeuneareniloba

Physocoleaciliatilobula (Schiffn.) Steph. = Cololejeuneainflectens

Physocoleahamata (Steph.) Steph. ≡ Cololejeuneahamata

Plagiochilaacanthophylla Gottsche = Plagiochilasciophila

Plagiochilabantamensisvar. γdenticulata Sande Lac. = Plagiochilakurzii

Plagiochilabatjanensis Schiffn. ex Herzog = Plagiochilateysmannii

Plagiochilabelangeriana Lindenb. = Plagiochilaarbuscula

Plagiochilablepharophoravar.major (Schiffn.) Bonner = Plagiochilasandei

Plagiochilabrauniana (Nees) Lindenb. ≡ Chiastocaulonbraunianum

Plagiochilaceramica Inoue = Plagiochilapulvinata

Plagiochilacomata (Nees) Lindenb. ≡ Solenostomacomatum

Plagiochiladecidua Inoue et Grolle = Plagiochilasciophila

Plagiochiladendroides (Nees) Lindenb. ≡ Chiastocaulondendroides

Plagiochilafrondescensvar.tenerrima (Nees) Schiffn. = Plagiochilafrondescens

Plagiochilafrondescens α tenerrima (Nees) Lindenb. = Plagiochilafrondescens

Plagiochilafrondescens γ rigida (Nees) Lindenb. = Plagiochilafrondescens

Plagiochilagedeana Schiffn. = Plagiochilahampeana

Plagiochilanovae-guineae Sande Lac. = Plagiochilachauviniana

Plagiochilaopposita (Reinw., Blume et Nees) Lindenb. ≡ Chiastocaulonoppositum

Plagiochilaopposita var. β falcata (Nees) Schiffn. ≡ Plagiochilaconjugatavar.falcata

Plagiochilaoppositavar. γfiliformis (Lindenb.) Schiffn. = Chiastocaulonoppositum

Plagiochilaopposita β falcata (Nees) Lindenb. ≡ Plagiochilaconjugatavar.falcata

Plagiochilaopposita γ filiformis Lindenb. nom. inval. = Chiastocaulonoppositum

Plagiochilapseudaberrans Inoue et Grolle = Plagiochilagymnoclada

Plagiochilaseemannii Mitt. = Plagiochilasandei

Plagiochilasemialata Sande Lac. = Plagiochilateysmannii

Plagiochilasingularis Schiffn. ≡ Dinckleriasingularis

Plagiochilatrapezoidea var. β tenera (Nees) Schiffn. = Dinckleriasingularis

Plagiochilatrapezoideavar. γmajor (Lindenb.) Schiffn. = Plagiochilagigantealaxior

Plagiochilatrapezoidea β tenera (Nees) Lindenb. = Dinckleriasingularis

Plagiochilatrapezoidea γ major Lindenb. nom. illeg. = Plagiochilagigantealaxior

Plagiochilavariegata Lindenb. = Syzygiellasecurifolia

Plagiochilionbraunianum (Nees) S.Hatt. ≡ Chiastocaulonbraunianum

Plagiochilionfimbriatum (Mitt.) Inoue ≡ Chiastocaulonfimbriatum

Plagiochilionoppositum (Reinw., Blume et Nees) S.Hatt. ≡ Chiastocaulonoppositum

Plectocoleaariadne (Taylor) Mitt. ≡ Solenostomaariadne

Plectocoleacomata (Nees) S.Hatt. ≡ Solenostomacomatum

Porellaacutifoliavar.subligulifera S.Hatt. = Porellaacutifoliavar.acutifolia

Psilocladaunguligera Schiffn. = Psilocladaclandestinasubsp.clandestina

Ptychanthusjavanicus Nees = Ptychanthusstriatusvar.striatus

Ptychanthusmoluccensis Sande Lac. = Ptychanthusstriatusvar.striatus

Ptychanthussquarrosus Mont. = Ptychanthusstriatus

Ptychocoleusbrunneus Steph. = Acrolejeuneapycnocladasubsp.pycnoclada

Ptychocoleuscumingianus (Mont.) Trevis. ≡ Schiffneriolejeuneacumingiana

Ptychocoleusdensifolius (Schiffn. ex P.Syd.) Steph. nom. illeg. = Schiffneriolejeuneapulopenangensis

Ptychocoleushaskarlianus (Gottsche) Steph. ≡ Schiffneriolejeuneatumidavar.haskarliana

Ptychocoleuspulopenangensis (Gottsche) Trevis. ≡ Schiffneriolejeuneapulopenangensis

Ptychocoleuspycnocladus (Taylor) Steph. ≡ Acrolejeuneapycnoclada

Ptychocoleustener Steph. = Acrolejeuneafertilis

Pycnolejeuneabadia Steph. = Lepidolejeuneabidentula

Pycnolejeuneaceylanica (Gottsche) Schiffn. ≡ Cheilolejeuneaceylanica

Pycnolejeuneaceylanica (Gottsche) Steph. nom. inval. ≡ Cheilolejeuneaceylanica

Pycnolejeuneaconnivens (Gottsche) Gottsche ex Schiffn. nom. inval. = Pycnolejeuneaconnivens

Pycnolejeuneadecurvifolia (Steph.) Steph. = Lepidolejeuneabidentula

Pycnolejeuneademissa Steph. ex Kachroo et R.M.Schust. nom. inval. = Cheilolejeunealongidens

Pycnolejeuneafalsinervis (Sande Lac.) Schiffn. ≡ Cheilolejeuneafalsinervis

Pycnolejeuneagigantea Steph. ≡ Cheilolejeuneagigantea

Pycnolejeuneaincisa (Gottsche) Schiffn. ≡ Cheilolejeuneaincisa

Pycnolejeuneameyeniana (Nees, Lindenb. et Gottsche) Steph. nom. inval. = Cheilolejeuneatrapezia

Pycnolejeuneaschwaneckei Steph. nom. inval. = Pycnolejeuneaschwaneckei

Pycnolejeuneatrapezia (Nees) Schiffn. ≡ Cheilolejeuneatrapezia

Pycnolejeuneaventricosa (Schiffn.) Schiffn. ex P.Syd. = Cheilolejeuneaventricosa

Pycnolejeuneaventricosa Schiffn. nom. inval. = Cheilolejeuneaventricosa

Radulacampanigera Mont. ≡ Cladoradulacampanigera

Radulaceramensis Steph. = Radulajavanica

Radulaformosavar.pycnolejeuneoides (Schiffn.) N.Kitag. = Radulaformosa

Radulaovalifolia Steph. = Radulajavanica

Radulapycnolejeuneoides Schiffn. = Radulaformosa

Radulasubsimilis Steph. = Radulareflexa

Radulawallichiana Lehm. = Radulajavanica

Rhaphidolejeunealongicruris (Steph.) Herzog ≡ Drepanolejeunealongicruris

Riccardiapinguis (L.) Gray ≡ Aneurapinguis

Saccogynarigidula (Nees) Schiffn. ≡ Saccogynidiumrigidulum

Saccogynidiumjugatum (Mitt.) Grolle = Saccogynidiumrigidulum

Schiffneriolejeuneahaskarliana (Gottsche) Gradst. ≡ Schiffneriolejeuneatumidavar.haskarliana

Schismadivaricatum Herzog = Herbertussendtneri

Schismapilifera (Schiffn.) Steph. = Herbertuspilifer

Schismapiligerum Steph. nom. inval. = Herbertuspilifer

Schistochilaamboinensis Steph. = Schistochilabeccariana

Schistochilaformosana Horik. = Schistochilablumei

Schistochilagraeffeana J.B.Jack et Steph. = Schistochilaaligera

Schistochilainversa Herzog = Schistochilasciurea

Schistochilaphilippinensis (Mont.) J.B.Jack et Steph. nom. inval. = Schistochilaaligera

Schistochilaphilippinensis (Mont.) Steph. = Schistochilaaligera

Schistochilaphilippinensisvar.transiens H.Buch nom. inval. = Schistochilaaligera

Schistochilapurpurascens Herzog = Schistochilaacuminata

Schistochilarecurvata H.Buch nom. inval. = Schistochilaaligera

Schistochilasciureaf.robustior Schiffn. = Schistochilasciurea

Schistochilasteraliger (Nees et Blume) H.A.Mill. ≡ Schistochilaaligera

Schistochilastergraeffeanus (J.B.Jack et Steph.) H.A.Mill. = Schistochilaaligera

Schistochilasterphilippinensis (Mont.) H.A.Mill. = Schistochilaaligera

Sendtneradiclados α scorpioides (Reinw., Blume et Nees) Nees = Mastigophoradiclados

Sendtneradiclados β calcarata (Reinw., Blume et Nees) Gottsche, Lindenb. et Nees nom. inval. = Mastigophoradiclados

Sendtneravrieseana Sande Lac. = Mastigophoradiclados

Spruceanthusmarianus (Gottsche) Mizut. ≡ Cheilolejeuneamariana

Stenolejeuneadentata (Steph.) Pócs ≡ Lejeuneastenodentata

Strepsilejeuneacavistipula Steph. ≡ Pycnolejeuneacavistipula

Symphyogynopsisfilicum (Nadeaud) Grolle = Symphyogynopsisgottscheana

Syzygiellavariegata (Lindenb.) Spruce = Syzygiellasecurifolia

Taxilejeuneaalbescens Steph. ≡ Lejeuneaalbescens

Taxilejeuneakarstenii Steph. = Lejeunealumbricoides

Taxilejeunealaxiretis (Steph.) Eifrig = Lejeuneasordida

Taxilejeunealumbricoides (Nees) Steph. ≡ Lejeunealumbricoides

Taxilejeuneasordida (Nees) Eifrig ≡ Lejeuneasordida

Telaraneacuneifolia (Steph.) J.J.Engel et G.L.Merr. ≡ Neolepidoziacuneifolia

Telaraneakogiana (Steph.) Grolle ≡ Tricholepidoziakogiana

Telaraneaneesii (Lindenb.) Fulford ≡ Tricholepidozianeesii

Telaraneawallichiana (Gottsche) R.M.Schust. ≡ Neolepidoziawallichiana

Thysananthusrenilobus (Gottsche) Schiffn. ≡ Caudalejeuneareniloba

Thysananthusundulatus (Gradst. et Grolle) Sukkharak nom. inval. = Thysananthusfrauenfeldii

Trachylejeuneacavistipula (Steph.) Steph. ≡ Pycnolejeuneacavistipula

Trichocoleabreviseta Steph. ≡ Leiomitrabreviseta

Trichocoleastriolata Steph. = Trichocoleapluma

Trichocoleatomentellavar.javanica (Reinw., Blume et Nees) Schiffn. = Trichocoleapluma

Xenolejeuneaceylanica (Gottsche) Tixier nom. illeg. ≡ Cheilolejeuneaceylanica

Zoopsisrigida Pearson = Zoopsissetulosa

Citation

Nadhifah A, Haerida I, Fastanti FS, Söderström L, Hagborg A, von Konrat M (2024) Beyond nutmeg, mace, and cloves: Checklist of the liverworts and hornworts of Maluku Islands (Moluccas), Indonesia. PhytoKeys 239: 107–193. https://doi.org/10.3897/phytokeys.239.116679

Funding Statement

Indonesia Endowment Fund for Education (LPDP), the Ministry of Finance, Republic of Indonesia

Additional information

Conflict of interest

The authors have declared that no competing interests exist.

Ethical statement

No ethical statement was reported.

Funding

This study was supported by the National Science Foundation (Award No. 2001509), the Indonesia Endowment Fund for Education (LPDP), the Ministry of Finance, Republic of Indonesia (NIB 20230722199822), and Deputy for Facilitation of Research and Innovation, National Research and Innovation Agency of Indonesia (BRIN) (2860/II.7/HK.01.00/8/2022 No. 33).

Author contributions

Conceptualization: MK, AN. Data curation: AH, LS. Formal analysis: LS. Methodology: MK. Resources: FSF, AN, IH. Visualization: LS. Writing - original draft: AN, MK. Writing - review and editing: AN, IH, AH, FSF, LS.

Author ORCIDs

Ainun Nadhifah https://orcid.org/0000-0003-0575-4306

Ida Haerida https://orcid.org/0000-0001-5769-9383

Fandri Sofiana Fastanti https://orcid.org/0000-0002-0925-7270

Lars Söderström https://orcid.org/0000-0002-9315-4978

Matt von Konrat https://orcid.org/0000-0001-9579-5325

Data availability

All of the data that support the findings of this study are available in the main text.