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- Dobzhansky T, Holz A M, Spassky B. Genetics of Natural Populations. VIII. Concealed Variability in the Second and the Fourth Chromosomes of Drosophila Pseudoobscura and Its Bearing on the Problem of Heterosis. Genetics. 1942 Sep;27(5):463–490. doi: 10.1093/genetics/27.5.463. [DOI] [PMC free article] [PubMed] [Google Scholar]