Abstract
Rust fungi (Pucciniales) comprise a large group of ecologically and economically important plant pathogens distributed globally where their hosts grow. The first published study of rusts in Indiana was carried out 131 years ago, and a revised checklist of all Pucciniales for Indiana has not been compiled since the works of Jackson from 1917 to 1920. Efforts to compile a checklist five years ago revealed a dire need for revision due to taxonomic and nomenclatural changes. We examined historical records, including online databases and literature, as well as new and historical collections in the Arthur Fungarium (PUR) at Purdue University. We provide an annotated checklist of all known species of rust fungi occurring in 90/92 Indiana counties. Names in prior records have been updated to reflect the current classifications for this group, with cross-reference to older names. A total of 301 Pucciniales taxa across 32 genera and their host species have been verified. This work serves as a resource for statewide plant disease diagnosticians and records the extant rust fungal biodiversity in Indiana to guide research efforts, resource management, and conservation.
Key words: Arthur Fungarium, plant pathogen, Tippecanoe County, uredinology
Introduction
Rust fungi (Pucciniales, formerly Uredinales) represent the largest natural group of phytopathogens and can cause devastating diseases in agricultural, horticultural, and forest habitats, resulting in millions of dollars in damage annually (e.g., rusts of wheat, corn, coffee, and soybeans). There are also some species that have been used as biocontrol against weeds, such as Phragmidium violaceum (Vargas-Gaete et al. 2019) and Puccinia species (Tanner et al. 2015; Bean et al. 2024). They are obligate biotrophs of plants (Arthur 1934; Savile 1971), requiring a living host for development, and have complex life cycles that may require up to seven spore types produced in five different spore stages and alternation between two phylogenetically unrelated host plants to complete their life cycles (Aime et al. 2018). Rusts are distributed across all geographical areas on wild and cultivated plants. It is expected that infectious plant disease ranges will shift under climate change (Caubel et al. 2017; Dudney et al. 2021; Miller et al. 2022), and current lists of species diversity will allow the study of changes in their distribution. Our work seeks to document the current knowledge of rust species present in Indiana.
Rust collection in Indiana began almost 200 years ago. References date back to a specimen labelled Aecidium dircatatum (now known as A. hydnoideum) on Dirca palustris (Thymelaeaceae), collected by Lewis David von Schweinitz during a visit to Hope in 1831 (McCain and Hennen 1982), and records of a rust destroying the foliage of blackberries in Henry and Wayne counties in 1872–1874 (Furnas 1874). However, the first explicit report of a rust fungus in the state was made by John M. Coulter in 1876, who documented a rust on Lespedeza violacea (Fabaceae), presumably near Hanover (Jackson 1915; McCain and Hennen 1982). We must also acknowledge the work and legacy of Joseph Charles Arthur that made uredinology flourish in the state of Indiana. Arthur named 29 genera and 309 species of rust fungi from North America and influenced the work of uredinologists like Frank D. Kern and his successors as directors of the Arthur Fungarium (PUR), George B. Cummins and Joe F. Hennen (McCain and Hennen 1982). Indiana became a state strong in uredinology history, heavily influenced by the PUR, housed at Purdue University. This fungarium represents the major center in the United States for taxonomic studies of rust fungi (McCain and Hennen 1982; Kaishian et al. 2024).
The first account of the rusts of Indiana was presented at the annual meeting of the Indiana Academy of Science in 1890 by E. M. Fisher on the Parasitic Fungi of Indiana, with a considerable number of species mentioned; however, this work was never published (Jackson 1915). A list of those species was obtained by Dr. L. M. Underwood and was included in the first published compilation of Indiana rust fungi by Underwood (1893). It listed 88 rust species observed across 19 counties, with most observations reported from Montgomery, Putnam, Johnson, and Tippecanoe counties (Fig. 1A). Additional lists were later published, including a list of rusts of Tippecanoe County in 1896 by Lillian Snyder and supplemented in 1898 with lists from Madison and Noble counties, as well as lists for Hamilton and Marion counties by G. W. Wilson in 1905, and two complete state lists by J. C. Arthur (Jackson 1915). An updated list was published by Jackson (1915), reporting 141 rusts, 34 of which were new reports for Indiana. The number of rusts for Indiana was later updated to 155 (Jackson 1917) and 165 (Jackson 1920). Since the efforts by Jackson, various rusts and new host records have been reported as part of the sixth annual A. H. Smith Great Lakes Foray (Schoknecht 1981), as specimens evaluated in rust biogeography studies (McCain and Hennen 1981), and studies of rusts on Cyperaceae and Juncaceae in Indiana (McCain 1984). In recent years, efforts have been carried out to document all fungi in the state, including a checklist of Indiana microfungi (Bates et al. 2019). However, the taxonomy for Pucciniales in this list needs considerable revision. Despite the significant efforts to document rusts in Indiana from the past 100 years, a checklist of all Pucciniales in the state has not been compiled since the works of Jackson from 1917 to 1920. This study was undertaken to generate an updated list with a modern re-evaluation.
Figure 1.
Distribution of specimens evaluated by Indiana counties A specimens studied by Underwood (1893) per Indiana county (n = 320) B specimens evaluated in this study accessioned in the Arthur Fungarium (PUR) collection (n = 4516; See Suppl. material 1 for detailed metadata for each specimen). Scale bars represent number of specimens studied per Indiana county. Grey color represents no rust reported for that county. Tippecanoe County (filled in black in panel B) was not included in the scale bar representation in panel B for comparison purposes between the two studies, as it was disproportionately represented with 2605 specimens. Individual maps generated with Map charts in Microsoft Excel version 16.88 (Microsoft Corporation, 2024).
Checklists are vital sources of information that set a benchmark for decision-making in research efforts, resource management, and policy (Reyserhove et al. 2020). Checklists have been valuable in monitoring, modeling, and predicting species presence and distribution in diverse groups of organisms. These include pollinators (IPBES 2016), rare species (IUCN 2020), and invasive alien species (Seebens et al. 2017; Reyserhove et al. 2020). They are instrumental in studies related to biodiversity conservation initiatives (Lowman et al. 2018; Parker et al. 2018; Menchetti et al. 2021; Lacher et al. 2022), discovery of new species (Pierson et al. 2014; Cantonwine et al. 2019; Nicolai et al. 2020; Maharani et al. 2022), and changes in response to climate change (Pearman et al. 2024). Furthermore, the use of herbarium records is highly valuable in ecological and epidemiological studies and modeling, such as assisting with the listing of rare taxa (Molano-Flores et al. 2023), providing data for biodiversity and landscape monitoring (Çelekli and Zariç 2023, informing conservation biology (Greve et al. 2016), setting baseline data particularly for poorly inventoried regions (Greve et al. 2016; Park et al. 2022), and supporting the study of phenological (Jones and Daehler 2018; Williams et al. 2021) and global environmental changes (Heberling et al. 2019; Lang et al. 2019). Access to the Arthur Fungarium, in this study, presented a valuable opportunity to assess and document the presence of rusts across Indiana with the purpose of generating a curated dataset to inform future ecological, epidemiological, and conservation work. This project verified records for potential misidentifications, known synonymies, and use of incorrect or obsolete names to reflect the most current classification for these rusts and the report of new records for the state. We provide a complete, up-to-date record of the Pucciniales species and their hosts known from Indiana.
Methods
This checklist was compiled from a variety of sources, including published literature (e.g., Arthur 1934), online databases [MyCoPortal: https://mycoportal.org/portal/collections/list.php (MyCoPortal 2025 and the U.S. National Fungus Collections database: https://nt.ars-grin.gov/fungaldatabases/ (Farr and Rossman 2018) and unpublished herbarium records. Data for the Arthur Fungarium (PUR) are maintained in the open-source software “Specify 7” by the Specify Collections Consortium (https://specifysoftware.org). Sources for each listing are provided within the list. The authors of fungal names follow the conventions in Index Fungorum (http://www.indexfungorum.org/names/AuthorsOfFungalNames.asp), and the current name of host plants follows the conventions used by the PLANTS database (http://plants.usda.gov) and Plants of the World Online (http://www.plantsoftheworldonline.org/). Herbarium specimens were studied to confirm dubious names or records, as well as for species represented by only a single statewide report.
The checklist is arranged alphabetically by genus, species, and subspecies-level taxa. Each species/subspecies is annotated with host family, species, and source of record. The current names of rust species are printed in bold type. Names not printed in bold are synonyms or other non-current names. References were inserted following the host name. The checklist does not include species that were the result of inoculation studies in greenhouses and that are not naturally occurring in the area (i.e., Hemileia vastatrix—PUR66308 and Phakopsora zeae—PUR51735).
We generated a specimen number by county list using location information from verified vouchers in the Arthur Fungarium (Suppl. material 1). Other sources of observations were not included in this analysis, since the level of detail for location varied among specimens. Metadata for records originating from PUR accessions are available in Suppl. material 2.
Maps depicting the number of observations per Indiana county were generated using map charts in Microsoft Excel version 16.88 (Microsoft Corporation, 2024). Additionally, all taxon names on the checklist were cross-checked against MycoBank and Index Fungorum by running a search for each name. Search results accuracy was categorized based on whether the correct current name was present in the search results as the current name or listed as a synonym, if an older name or another name was referenced as the current name in the database, or if a search of the current name or old name returned no results. The results from these searches are available in Suppl. material 3.
Results and discussion
Our checklist of rust fungi in Indiana includes a total of 301 current names, 339 synonyms and non-current names, and 13 ambiguous names. It documents 301 Pucciniales taxa (275 species, 21 varieties, 4 subspecies, and 1 forma specialis) distributed on 900+ host taxa. Most of the specimens from Indiana housed in the Arthur Fungarium are from Tippecanoe County (57.6%), with 2,605 out of 4,516 specimens collected in the same county where the fungarium is located. The next most sampled counties show a markedly smaller percentage, with Jasper (2.45%; 111 specimens), Montgomery (2.24%; 101 specimens), Hamilton (1.97%; 89 specimens), Putnam (1.97%; 89 specimens), Franklin (1.77%; 80 specimens), Fayette (1.64%; 74 specimens), Lawrence (1.64%; 74 specimens), and Kosciusko (1.44%; 65 specimens) counties. A total of 28% of counties (19/90) have between 20 and 53 specimens vouchered in the PUR (from highest specimen number to lowest: Porter, Marion, LaGrange, Brown, St. Joseph, Steuben, White, Warren, LaPorte, Fountain, Spencer, Lake, Carroll, Madison, Vigo, Henry, Posey, Orange, Parke, Wayne, Noble, Warrick, Marshall, Crawford, and Johnson). While another 28% of counties (19/90) have between 10 and 19 specimens accessioned per county (from highest specimen number to lowest: Jefferson, Owen, Starke, Jackson, Newton, Martin, Washington, Gibson, Clark, Harrison, Perry, Pulaski, Clinton, Knox, Monroe, Benton, Cass, Fulton, and Vanderburgh), 41% of the Indiana counties represented in the PUR collection have less than 10 specimens vouchered in the PUR per county (Fig. 1B). A detailed list of all counties and their respective number of specimens in the PUR is provided in Suppl. material 2.
The recorded distribution of rust fungi in Indiana has been attributed to the distribution of the collectors and is likely representative of changes in land use (e.g., conversion to agricultural land in a large part of the state). McCain and Hennen (1981) pointed out that over 70% of Indiana rust fungi at the time occurred within easy collecting distance of West Lafayette, where the PUR is located (e.g., Fig. 1A). The distribution data was skewed away from eastern and southern Indiana, with information at the time only for 44 of the 92 Indiana counties, evidently exemplifying the distribution of collectors rather than of taxa (McCain 1984). Although this disproportionate representation from Tippecanoe County persists—an issue that continues to this day—the PUR currently houses specimens for 90/92 counties in Indiana, only lacking representatives from Hancock and Ohio counties. This highlights the advances in Pucciniales knowledge for Indiana in the past 40+ years and the efforts to conduct fieldwork across the state. Yet, it is still clear that systematic statewide specimen collection is lacking to generate valid biogeographical or phenological hypotheses (McCain 1984).
Our checklist includes a total of 339 Pucciniales taxon names that had been previously reported for Indiana that are currently obsolete names, as well as 13 ambiguous taxon names. Taxonomic confusion has characterized rust fungi due to the development of multiple classification schemes, homoplasy in diagnostic characters, and the use of separate naming systems for sexual and asexual morphs under prior nomenclatural codes. Additionally, more than 200 generic names have been described in Pucciniales, but only about one-third of those names are accepted (Aime and McTaggart 2021). Taxonomy and nomenclature of many of the rusts reported in Indiana have changed over time and not always kept up to date in records. This became evident in the rusts listed in the recent checklist of Indiana microfungi (Bates et al. 2019). This study provides updated names with the most correct identification based on a re-evaluation of the dataset considering current taxonomy.
Index Fungorum (https://www.indexfungorum.org/) and MycoBank (https://www.mycobank.org/) are regularly consulted by mycologists and plant pathologists to assign current taxonomy to their rust fungi samples. Nevertheless, we note that these public name repositories may still contain inaccurate names (Suppl. material 3). Both repositories seem to follow similar patterns of accuracy/inaccuracy (Fig. 2). Of the 653 names listed on our checklist (301 current names, 339 synonyms and non-current names, and 13 ambiguous names), only 58.5% (MycoBank) and 63% (Index Fungorum) of these names directly refer to the correct name when conducting a search in each database. The misidentifications and use of obsolete names are usually not congruent between the two databases. Discrepancy issues between these repositories are known and extend to disagreements in terms of placing genera to family- and order-level, based on differential updates and taxonomic opinion (Durkin et al. 2020). Our goal is that this resource, with the most up-to-date taxonomy, serves as a reference point for correct identification of rusts from Indiana to plant diagnosticians and researchers.
Figure 2.
MycoBank and Index Fungorum taxonomic accuracy in comparison to the curated list of taxa in our Indiana rust fungi checklist (n = 653) after conducting a search for each taxon name in our list in both repositories. Search results accuracy was categorized based on whether the correct current name was present in the search results as the current name or listed as a synonym, if an older name or another name was referenced as the current name in the database, or if a search of the current name or old name returned no results.
Indiana checklist of rust fungi (Pucciniales)
In this checklist, 301 rust taxa from Indiana are reported. A list of host species, taxonomic/nomenclatural synonyms of each rust taxon, and references are also included.
Aecidium albiperidium Arthur
See: Puccinia caricina DC., s.lat.
Aecidium aquilegiae Pers.
See: Puccinia persistens Plowr. s.lat.
Aecidium asperifolii Pers.
See: Puccinia recondita Roberge ex Desm.
Aecidium asterum Schwein.
See: Puccinia dioicae Magnus, s.lat.
Aecidium berberidis Pers.
See: Puccinia graminis Pers.
Aecidium boehmeriae Arthur
Boehmeria cylindrica (L.) Sw. (Urticaceae) (Jackson 1915; Anderson and Anderson 1919; USDA 1960) PUR: 42992
Aecidium compositarum Mart.
Eupatorium perfoliatum L. (Asteraceae) (Snyder 1896)
Note: Probably aecial state of Puccinia eleocharidis Arthur.
Aecidium cyparissiae DC.
Euphorbia commutata Engelm. ex A. Gray (Euphorbiaceae) (Underwood 1896)
Note: Probably refers to Aecidium tithymali Arthur.
Aecidium dakotense Griffiths
Thalictrum dioicum L. (Ranunculaceae) (Jackson 1920) PUR: 6760
Aecidium dicentrae Trel.
See: Aplopsora dicentrae (Mains & H.W. Anderson) Buriticá & J.F. Hennen
Aecidium dracontii Schwein. [as ‘dracontinatum’ in protologue]
See: Uromyces ari-triphylli (Schwein.) Seeler
Aecidium erigeronatum Schwein.
See: Puccinia dioicae Magnus, s.lat.
Aecidium eupatorii Dietel
See: Puccinia eleocharidis Arthur
Aecidium euphorbiae Pers. ex J.F. Gmel.
See: Uromyces proëminens (DC.) Lév.
Aecidium geranii DC.
See: Uromyces geranii (DC.) Lév.
Aecidium grossulariae (J.F. Gmel.) DC.
See: Puccinia caricina DC., s.lat.
Aecidium hepaticatum Schwein.
See: Tranzschelia arthurii Tranzschel & M.A. Litv.
Aecidium hydnoideum Berk. & M.A. Curtis
See: Puccinia dioicae Magnus, s.lat.
Aecidium impatientis Schwein.
See: Puccinia impatientis (Schwein.) Arthur
Aecidium lycopi W.R. Gerard
See: Puccinia angustata Peck
Aecidium mariae-wilsoniae Peck [as ‘mariae-wilsoni’]
See: Puccinia ellisiana Thüm.
Aecidium oenotherae Peck
See: Puccinia dioicae Magnus, s.lat.
Aecidium onobrychidis Burrill
See: Puccinia andropogonis Schwein.
Aecidium pedatatum Schwein
See: Uromyces andropogonis Tracy
Aecidium pentastemonis Schwein.
See: Puccinia andropogonis Schwein.
Aecidium physalidis Burrill
See: Puccinia kansensis Ellis & Barthol.
Aecidium pteleae Berk. & M.A. Curtis
See: Puccinia windsoriae Schwein.
Aecidium pustulatum M.A. Curtis
See: Puccinia andropogonis Schwein.
Aecidium ranunculacearum DC.
Anemone pensylvanica L. (Ranunculaceae) (Snyder 1896)
Notes: A. ranunculacearum is an ambiguous name and may refer to Puccinia magnusiana, P. recondita, or Uromyces dactylidis.
Aecidium ranunculi Schwein.
See: Puccinia eatoniae Arthur
Aecidium sambuci Schwein.
See: Puccinia sambuci (Schwein.) Arthur
Aecidium tithymali Arthur
Euphorbia commutata Engelm. ex A. Gray (=Tithymalus commutatus (Engelm. ex A. Gray) Klotzsch & Garcke (Euphorbiaceae) (Jackson 1917; Arthur 1918; Anderson and Anderson 1919; USDA 1960) PUR: 43105, 43110, 43111, 43112, 43113, 43114, 47301, N6674.
Aecidium trillii Burrill
See: Uromyces halstedii De Toni
Aecidium verbenicola Ellis & Kellerm. (Also reported as Aecidium verbenae Speg. [author as ‘Spreng.’]
See: Puccinia vilfae Arthur & Holw.
Allodus ambigua (Alb. & Schwein.) Arthur
See: Puccinia difformis Kunze
Allodus claytoniata (Schwein.) Arthur
See: Puccinia mariae-wilsoniae G.W. Clinton
Allodus podophylli (Schwein.) Arthur
Podophyllum peltatum L. (Berberidaceae) (Underwood 1893; Snyder 1896, 1898; Arthur 1899; Jackson 1915; Anderson and Anderson 1919) PUR: 35717, 35720, 35721, 35727, 35731, 35732, 35733, 35734, 35735, 35736, 35737, 35741, 35742, 35743, 35744, 35750, 35751, 35753, 35754, 35756, 35764, 35767, 35768, 35770, 35771, 35773, 35777, 35780, 35781, 35794, 35795, 35796, 35797, 35803, 35804, 35805, 35807, 35812, 35813, 35814, 35816, 35818, 46794, 46795, 46810, 65820, 88839, 89279, N617, N2151, N2152, N2153, N4516, N4530, N4668, N5872, N5873, N5874, N5875, N5876, N5877, N5878
Host not reported (Van Hook 1910)
Allodus tenuis (Schwein.) Arthur
See: Puccinia tenuis Burrill
Aplopsora dicentrae (Mains & H.W. Anderson) Buriticá & J.F. Hennen
Dicentra cucullaria (L.) Bernh. (Fumariaceae) (Underwood 1893; Anderson and Anderson 1919; Buriticá 1998); (=Bicuculla cucullaria (L.) Millsp.) (Jackson 1917)
Laportea canadensis (L.) Weddell (Urticaceae) (Buriticá 1998)
Aplopsora tanakae (S. Ito) Buriticá & J.F. Hennen
Amphicarpaea bracteata (L.) Fernald (Fabaceae) (Emmons et al. 1960; Cummins 1962; Cooke 1975; Buriticá 1998)
Aregma disciflora (Tode) Arthur
See: Phragmidium mucronatum (Pers.) Schltdl.
Aregma fragariae (DC.) Arthur
See: Phragmidium tormentillae Fuckel
Aregma speciosum Fr. [as ‘speciosa’]
See: Phragmidium speciosum (Fr.) Cooke
Bubakia crotonis (Cooke) Arthur
Croton monanthogynus Michx. (Euphorbiaceae) (Jackson 1915)
Croton sp. PUR: N2586
Bullaria bardanae (Corda) Arthur
See: Puccinia bardanae (Wallr.) Corda
Bullaria bullata (Pers.) Arthur
See: Puccinia angelicae (Schumach.) Fuckel
Bullaria chrysanthemi (Roze) Arthur
See: Puccinia tanaceti var. tanaceti DC.
Bullaria globosipes (Peck) H.S. Jacks.
See: Puccinia globosipes Peck
Bullaria hieracii (Schumach.) Arthur
See: Puccinia hieracii (Röhl.) H. Mart.
Bullaria kuhniae (Schwein.) Kern
See: Puccinia kuhniae Schwein.
Bullaria kuhniae (Schwein.) H.S. Jacks. (nomen invalidum)
See: Puccinia kuhniae Schwein.
Bullaria taraxaci Arthur
See: Puccinia hieracii (Röhl.) H. Mart.
Caeoma agrimoniae Schwein.
See: Pucciniastrum agrimoniae (Dietel) Tranzschel
Caeoma nitens Schwein.
See: Gymnoconia nitens (Schwein.) Arthur
Caeomurus caladii (Schwein.) Kuntze
See: Uromyces ari-triphylli (Schwein.) Seeler
Caeomurus caryophyllinus (Schrank) Kuntze
See: Uromyces dianthi (Pers.) Niessl
Caeomurus euphorbiae (Cooke & Peck) Kuntze
See: Uromyces proëminens (DC.) Lév.
Caeomurus gaurinus (Peck) Arthur
See: Uromyces plumbarius Peck
Caeomurus graminicola (Burrill) Kuntze [as ‘graminicolus’]
See: Puccinia graminicola (Burrill) Demers & Castl.
Caeomurus howei (Peck) Kuntze
See: Uromyces asclepiadis Cooke
Caeomurus hedysari-paniculati (Schwein.) Arthur
See: Uromyces hedysari-paniculati (Schwein.) Farl.
Caeomurus hyperici-frondosi (Schwein.) Arthur
See: Uromyces triquetrus Cooke
Caeomurus junci (Tul. & C. Tul.) Kuntze
See: Uromyces junci (Desm.) Tul. & C. Tul.
Caeomurus lespedezae-procumbentis (Schwein.) Arthur
See: Uromyces lespedezae-procumbentis (Schwein.) M.A. Curtis
Caeomurus peryginus (Halst.) Kuntze [as ‘perigynius’]
See: Uromyces perigynius Halst.
Caeomurus phaseoli (Pers.) Arthur
See: Uromyces appendiculatus var. appendiculatus (Pers.) Unger
Caeomurus pisi (Pers.) Gray
This is an illegitimate name (Art. 53.1)
See: Uromyces viciae-fabae (Pers.) J. Schröt.
Caeomurus polygoni (Pers.) Kuntze
See: Uromyces polygoni-avicularis (Pers.) G.H. Otth
Caeomurus rudbeckiae (Arthur & Holw.) Kuntze
See: Uromyces rudbeckiae Arthur & Holw
Caeomurus terebinthi (DC.) Kuntze
See: Pileolaria brevipes Berk. & Ravenel
Caeomurus trifolii (R. Hedw.) Gray, nom. ambig.
See: Uromyces trifolii; U. trifolii-repentis var. trifolii-repentis and U. trifolii-repentis var. fallens
Trifolium medium L. (Fabaceae) (Arthur 1899) [Doubtful record]
Calyptospora columnaris (Alb. & Schwein.) J.G. Kühn
Abies balsamea (L.) Mill. (Pinaceae)
Cerotelium tanakae S. Ito
See: Aplopsora tanakae (S. Ito) Buriticá & J.F. Hennen
Chrysomyxa albida J.G. Kühn
See: Kuehneola uredinis (Link) Arthur
Chrysomyxa ledicola (Peck) Lagerh.
Picea mariana (Mill.) Britton, Sterns & Poggenb. (Pinaceae) PUR: 4729
Coleosporium apocynaceum Cooke
Amsonia cv. ‘Blue Ice’ (Apocynaceae) (Abbasi et al. 2016a)
Coleosporium asterum (Dietel) Syd. & P. Syd.
Aster salicifolius Lam. (Asteraceae) (Jackson 1915)
A. shortii Hook (Jackson 1915)
Callistephus chinensis (L.) Nees (=Callistephus hortensis Cass.) (Asteraceae)
C. hortensis Cass. (Jackson 1915; Pipal 1915; Anderson and Anderson 1919)
Pinus echinata Mill. (Pinaceae)
Pinus sp.
P. taeda L.
Solidago altissima L. (Asteraceae) (Anderson and Anderson 1919; Emmons et al. 1960; Schoknecht 1981)
S. arguta Aiton (Underwood 1893; Arthur 1899; Jackson 1915; Anderson and Anderson 1919)
S. bicolor L. (Anderson and Anderson 1919; Jackson 1920)
S. caesia L. (Underwood 1893; Arthur 1899; Jackson 1915; Anderson and Anderson 1919; Emmons et al. 1960; Schoknecht 1981)
S. canadensis (cf.) L. (Arthur 1899; Anderson and Anderson 1919)
S. canadensis L. (Underwood 1893; Jackson 1915)
S. erecta Pursh (Jackson 1920)
S. flexicaulis L. (Arthur 1899; Jackson 1915); (=S. latifolia L.) (Underwood 1893; Anderson and Anderson 1919)
S. gigantea Aiton (=S. gigantea subsp. serotina (Kuntze) McNeill) (McCain et al. 1990); (=S. serotina Aiton) (Underwood 1893; Arthur 1899; Jackson 1915)
S. juncea Aiton (Jackson 1920; Cooke 1967)
S. nemoralis Aiton (McCain et al. 1990)
S. patula Muhl. ex Willd. (Underwood 1893; Arthur 1899; Jackson 1915; Anderson and Anderson 1919)
S. rugosa Mill. (Underwood 1893; Arthur 1899; Jackson 1915)
Solidago sp. (Snyder 1896)
S. uliginosa Nutt.
S. ulmifolia Muhl. ex Willd. (Jackson 1915; Anderson and Anderson 1919; Emmons et al. 1960)
Symphyotrichum cordifolium (L.) G.L. Nesom (=Aster cordifolius L. and A. sagittifolius Wedemeyer ex Willd.) (Asteraceae) (Underwood 1893; Arthur 1899; Jackson 1915; Anderson and Anderson 1919)
S. drummondii (Lindl.) G.L. Nesom; var. drummondii (Lindl.) G.L. Nesom (=A. drummondii Lindl.) (Jackson 1915)
S. ericoides (L.) G.L. Nesom; var. ericoides (=A. ericoides L.) (Jackson 1915; Anderson and Anderson 1919)
S. laeve (L.) A. Löve & D. Löve
S. lanceolatum (Willd.) G.L. Nesom; ssp. lanceolatum var. lanceolatum (=A. paniculatus Lam.) (Underwood 1893; Arthur 1899; Jackson 1915; Anderson and Anderson 1919)
S. lateriflorum (L.) A. Löve & D. Löve
S. novae-angliae (L.) G.L. Nesom; (=A. novae-angliae L.) (Underwood 1893; Arthur 1899; Jackson 1915; Anderson and Anderson 1919)
S. novi-belgii (L.) G.L. Nesom var. novi-belgii (=A. longifolius Lam.) (Jackson 1915)
S. oolentangiense (Riddell) G.L. Nesom var. oolentangiense (=A. azureus Lindl.) (Underwood 1893; Arthur 1899; Jackson 1915; Anderson and Anderson 1919)
S. pilosum var. pilosum (Willd.) G.L. Nesom (=A. pilosus Willd.)
S. praealtum (Poir.) G.L. Nesom (=A. salicifolius Aiton) (Underwood 1893); var. praealtum (Poir.) G.L. Nesom (=A. praealtus Poir.) (McCain et al. 1990)
S. puniceum (L.) Á. Löve & D. Löve; var. puniceum (L.) Á. Löve & D. Löve (=A. firmus Nees) (McCain et al. 1990); (=A. puniceus L.) (Underwood 1893; Arthur 1899; Jackson 1915); (=S. firmum (Nees) G.L. Nesom)
S. shortii (Lindl.) G.L. Nesom (=A. shortii Lindl.) (Underwood 1893; Anderson and Anderson 1919)
Symphyotrichum sp. (=Aster sp.) (Snyder 1898)
S. tradescantii (L.) G.L. Nesom (=A. tradescantii L.) (Underwood 1893; Arthur 1899; Jackson 1915)
Coleosporium campanulae (Pers.) Tul.
Campanula rapunculoides L. (Campanulaceae) (McCain and Hennen 1981; McCain et al. 1990)
Campanulastrum americanum (L.) Small (=Campanula americana L.) (Campanulaceae) (Jackson 1915; Anderson and Anderson 1919; Cooke 1967)
Coleosporium delicatulum (Arthur & F. Kern) Hedgc. & Long
Euthamia graminifolia (L.) Nutt. (Asteraceae) (Jackson 1915); var. graminifolia (L.) Nutt. (=Solidago graminifolia (L.) Salisb.) (Emmons et al. 1960)
Solidago lanceolata L. (Asteraceae) (Anderson and Anderson 1919)
Coleosporium elephantopi Thüm. [as ‘ elephantopodis ’]
Elephantopus carolinianus Willd. (Asteraceae) (Jackson 1915)
Coleosporium helianthi (Schweinitz) Arthur
Helianthus decapetalus L. (Asteraceae) (Jackson 1915)
H. microcephalus Torr. & A. Gray (McCain et al. 1990)
H. giganteus L. (USDA 1960)
H. hirsutus Raf. (Jackson 1915)
Silphium integrifolium Michx. (Asteraceae) (Jackson 1915)
S. perfoliatum L.
S. terebinthinaceum Jacq. (Jackson 1915)
Coleosporium hydrangeae (Berk. & M.A. Curtis) L. Snyder
See: Pucciniastrum hydrangeae (Magnus) Arthur
Coleosporium ipomoeae (Schwein.) Burrill
Ipomoea pandurata (L.) G. Mey. (Convolvulaceae) (Underwood 1896; Arthur 1899; Jackson 1915; Anderson and Anderson 1919)
Ipomoea sp. (Snyder 1896)
Coleosporium montanum (Arthur & F. Kern) McTaggart & Aime
Solidago sp. (Asteraceae) (McTaggart and Aime 2018) PUR: N11689, N21953
Coleosporium pinicola (Arthur) Arthur
Pinus virginiana Mill. (Pinaceae) (USDA 1960)
Coleosporium rubi Ellis & Holw.
See: Kuehneola uredinis (Link) Arthur
Coleosporium solidaginis (Schwein.) Thüm.
Solidago canadensis (Asteraceae) PUR: N23076, N23056
Coleosporium sonchi-arvensis (Pers.) Lév.
Hieracium sp. (Asteraceae) (Underwood 1893) [Doubtful record]
See: Coleosporium asterum (Dietel) Syd. & P. Syd. and C. vernoniae Berk. & M.A. Curtis
Coleosporium terebinthinaceae (Schwein.) Arthur
See: Coleosporium helianthi (Schweinitz) Arthur
Coleosporium tussilaginis (Pers.) Lév.
Packera aurea (L.) Á. Löve & D. Löve (=Senecio aureus L.) (Asteraceae) (McCain et al. 1990)
Senecio sp. (Asteraceae) PUR: 89361
Coleosporium vernoniae Berk. & M.A. Curtis
Elephantopus carolinianus Raeusch. (Asteraceae)
Pinus echinata Mill. (Pinaceae)
P. nigra Arnold (USDA 1960)
P. rigida Mill.
P. virginiana Mill.
Vernonia arkansana DC. (Asteraceae)
V. fasciculata Michx. (Underwood 1893; Arthur 1899; Jackson 1915; Anderson and Anderson 1919)
V. gigantea (Walter) Trel. ; subsp. gigantea (=V. altissima Nutt.) (Jackson 1915; Anderson and Anderson 1919; Emmons et al. 1960; Schoknecht 1981)
Vernonia × illinoensis Gleason [gigantea × missurica] (Jackson 1920)
V. missurica Raf.
V. noveboracensis (L.) Michx. (Underwood 1893; Arthur 1899; Jackson 1915; Van Hook 1927)
Vernonia sp. (Cooke 1967)
Coleosporium viburni Arthur
Viburnum lentago L. (Caprifoliaceae)
Viburnum sp. (Ruhl et al. 1982)
Coleosporium sp.
Pinus sp. (Pinaceae) (Ruhl et al. 1983)
Cronartium comandrae Peck
Comandra umbellata (L.) Nutt. (Santalaceae)
Cronartium fusiforme Hedgc. & N.R. Hunt ex Cummins
Pinus sp. (Pinaceae) (Ruhl et al. 1982)
Cronartium harknessii E. Meinecke
Pinus sp. (Pinaceae) (Ruhl et al. 1983)
Cronartium quercuum (Berk.) Miyabe ex Shirai
Quercus alba L. (Fagaceae)
Q. bicolor Willd.
Q. macrocarpa Michx.
Q. michauxii Nutt.
Q. muehlenbergii Engelm.
Q. pagoda Raf.
Q. phellos L.
Q. rubra L.
Cronartium ribicola J.C. Fisch.
Ribes cynosbati L. (Grossulariaceae) (Emmons et al. 1960)
Ribes sp.
R. aureum var. villosum DC. (=R. odoratum H.L. Wendl.) (Wilson et al. 2014)
Dasyspora anemones-virginianae (Schwein.) Arthur
See: Puccinia anemones-virginianae Schwein.
Dasyspora asteris (Duby) Arthur
See: Puccinia cnici-oleracei Pers. ex Desm.
Dasyspora circaeae (Pers.) Arthur
See: Puccinia circaeae Pers.
Dasyspora dayi (Clinton) Arthur
See: Puccinia dayi Clinton
Dasyspora glechomatis (DC.) Arthur [as gleocomatis]
See: Puccinia glechomatis DC.
Dasyspora lobeliae (W.R. Gerard) Arthur
See: Puccinia lobeliae W.R. Gerard
Dasyspora malvacearum (Bertero) Arthur
See: Puccinia malvacearum Bertero ex Mont.
Dasyspora physostegiae (Peck & Clinton) H.S. Jacks.
See: Puccinia physostegiae Peck & Clinton
Dasyspora ranunculi (Kuntze) Arthur
See: Puccinia andina Dietel & Neger
Dasyspora saxifragae (Schltdl.) Arthur
See: Puccinia heucherae (Schwein.) Dietel
Dasyspora seymeriae (Burrill) Arthur [as ‘seymouriae’ by Arthur (1906)]
See: Puccinia seymeriae Burrill
Dasyspora silphii (Schwein.) Arthur
See: Puccinia silphii Schwein.
Dasyspora xanthii (Schwein.) Arthur
See: Puccinia xanthii Schwein.
Dicaeoma aletridis (Berk. & M.A. Curtis) Kuntze
See: Puccinia aletridis Berk. & M.A. Curtis
Dicaeoma andropogi (Schwein.) Kuntze
See: Puccinia andropogonis Schwein.
Dicaeoma andropogonis (Schwein.) Kuntze
See: Puccinia andropogonis Schwein.
Dicaeoma anemones (Pers.) Arthur
See: Tranzschelia pseudofusca M. Scholler & M. Abbasi
Dicaeoma anemones-virginianae (Schwein.) Arthur
See: Puccinia anemones-virginianae Schwein.
Dicaeoma angustatum (Peck) Kuntze
See: Puccinia angustata Peck
Dicaeoma antirrhini (Dietel & Holw.) H.S. Jacks.
See: Puccinia antirrhini Dietel & Holw.
Dicaeoma apocryptum (Ellis & Tracy) Kuntze
See: Puccinia apocrypta Ellis & Tracy
Dicaeoma argentatum (Schultz) Kuntze
See: Puccinia argentata (Schultz) G. Winter
Dicaeoma asparagi (DC.) Kuntze
See: Puccinia asparagi DC.
Dicaeoma asperifolii (Pers.) Kuntze
See: Puccinia recondita Roberge ex Desm. s.str.
Dicaeoma asperifolii (Pers.) Kuntze, sensu Arthur (1899), nom. ambig.
See: Puccinia impatientis (Schwein.) Arthur and P. recondita Roberge ex Desm. s.str.
Avena sativa L (Poaceae) (Arthur 1899) [Doubtful record]
Dicaeoma asteris (Duby) Kuntze
See: Puccinia cnici-oleracei Pers. ex Desm.
Dicaeoma bolleyanum (Sacc.) Kuntze
See: Puccinia sambuci (Schwein.) Arthur
Dicaeoma calthae (Link) Kuntze
See: Puccinia calthae Link
Dicaeoma canaliculatum (Schwein.) Kuntze
See: Puccinia canaliculata (Schwein.) Lagerh.
Dicaeoma cephalanthi (Seym.) H.S. Jacks.
See: Puccinia seymouriana Arthur
Dicaeoma circaeae (Pers.) Kuntze
See: Puccinia circaeae Pers.
Dicaeoma cnici H. (Mart.) Arthur
See: Puccinia cnici H. Mart.
Dicaeoma clematidis (DC.) Arthur, nom. ambig.
This is an ambiguous name applied by different authors to different taxa.
Aquilegia sp. (Ranunculaceae) (Jackson 1915)
Clematis virginiana L. (Ranunculaceae) (Jackson 1915)
Thalictrum thalictroides (L.) Eames & B. Boivin (=Syndesmon thalictroides (L.) Hoffmanns. ex Britton (Ranunculaceae) (Jackson 1915)
Elymus repens (L.) Gould (=Agropyron repens (L.) P. Beauv.) (Poaceae) (Jackson 1915)
Bromus ciliatus L. (Poaceae) (Jackson 1915)
B. arvensis L. (=Bromus japonicus Thunb.) (Jackson 1915)
B. ciliatus var. purgans (L.) A. Gray (=B. purgans L.) (Jackson 1915)
B. secalinus L. (Jackson 1915)
Hordeum jubatum L. (Poaceae) (Jackson 1915)
Glyceria grandis S. Watson var. grandis (=Panicularia grandis (S. Watson) Nash (Poaceae) (Jackson 1915)
Dicaeoma conoclinii (Seym.) Kuntze
See: Puccinia conoclinii Seym. ex Burrill
Dicaeoma convolvuli (Pers.) Kuntze
See: Puccinia convolvuli (Pers.) Castagne
Dicaeoma cyperi (Arthur) Kuntze
See: Puccinia cyperi Arthur
Dicaeoma cypripedii (Arthur) Arthur
See: Puccinia cypripedii Arthur
Dicaeoma dayi (Clinton) Kuntze
See: Puccinia dayi Clinton
Dicaeoma dochmia (Berk. & M.A. Curtis) Kuntze
See: Puccinia dochmia Berk. & M.A. Curtis
Dicaeoma eatoniae Arthur
See: Puccinia eatoniae Arthur
Dicaeoma eleocharidis (Arthur) Kuntze
See: Puccinia eleocharidis Arthur
Dicaeoma ellisianum (Thüm.) Kuntze
See: Puccinia ellisiana Thüm.
Dicaeoma emaculatum (Schwein.) Kuntze
See: Puccinia emaculata Schwein.
Dicaeoma epiphyllum (L.) Kuntze
See: Puccinia poarum Nielsen
[Jackson’s (1920) report on Alopecurus geniculatus L. is Puccinia poae-nemoralis G.H. Otth]
Dicaeoma eriophori (Thüm.) Kuntze
See: Puccinia eriophori Thüm.
Dicaeoma extensicola (Plowr.) Kuntze
See: Puccinia dioicae Magnus. s.lat.
Dicaeoma flosculosorum (Alb. & Schwein.) H. Mart.
See: Puccinia hieracii (Röhl.) H. Mart.
Dicaeoma fraxini (Schwein.) Arthur
See: Puccinia sparganioides Ellis & Barthol.
Dicaeoma galiorum (Link) Arthur, nom. ambig.
See: Puccinia punctata Link; P. punctata var. troglodytes (Lindr.) Arthur and P. difformis Kunze
Dicaeoma grossulariae (Schumach.) H.S. Jacks.
See: Puccinia caricina DC., s.lat.
Dicaeoma helianthii (Schwein.) Kuntze
See: Puccinia helianthi Schwein.
Dicaeoma heliopsidis (Schwein.) Kuntze
See: Puccinia helianthi Schwein.
Dicaeoma hibisciatum (Schwein.) Arthur
See: Puccinia schedonnardi Kellerm. & Swingle
Dicaeoma impatientis (Schwein.) Arthur
See: Puccinia impatientis (Schwein.) Arthur
Dicaeoma iridis (DC.) Kuntze
See: Puccinia iridis Wallr.
Dicaeoma kuhniae (Schwein.) Kuntze
See: Puccinia kuhniae Schwein.
Dicaeoma lateripes (Berk. & Ravenel) Kuntze
See: Puccinia lateripes Berk. & Ravenel
Dicaeoma lobeliae (W.R. Gerard) Arthur
See: Puccinia lobeliae W.R. Gerard
Dicaeoma ludibundum (Ellis & Everh.) Kuntze
See: Puccinia dioicae Magnus, s.lat.
Dicaeoma majanthae (Schumach.) Arthur
See: Puccinia sessilis J. Schröt.
Dicaeoma melicae (P. Syd. & Syd.) Arthur
See: Puccinia melicae P. Syd. & Syd.
Dicaeoma menthae (Pers.) Gray
See: Puccinia menthae Pers.
Dicaeoma minutissimum (Arthur) H.S. Jacks.
See: Puccinia minutissima Arthur
Dicaeoma montanense (Ellis) Kuntze
See: Puccinia montanensis Ellis
Dicaeoma obscurum (J. Schröt.) Kuntze
See: Puccinia obscura J. Schröt.
Dicaeoma obtectum (Peck) Kuntze
See: Puccinia obtecta Peck
Dicaeoma orbicula (Peck & Clinton) Kuntze
See: Puccinia orbicula Peck & Clinton
Dicaeoma pammelii (Trel.) Arthur
See: Puccinia emaculata Schwein.
Dicaeoma patruelis (Arthur) H.S. Jacks.
See: Puccinia dioicae Magnus, s.lat.
Dicaeoma peckii (De Toni) Arthur
See: Puccinia dioicae Magnus, s.lat.
Dicaeoma physostegiae (Peck & Clinton) Kuntze
See: Puccinia physostegiae Peck & Clinton
Dicaeoma poculiforme (Jacq.) Kuntze
See: Puccinia graminis Pers.
Dicaeoma podophylli (Schwein.) Kuntze
See: Allodus podophylli (Schwein.) Arthur
Dicaeoma polygoni-amphibii (Pers.) Arthur
See: Puccinia polygoni-amphibii Pers.
Dicaeoma polygoni-convolvuli (Hedw.) Arthur
See: Puccinia polygoni-amphibii Pers.
Dicaeoma prenanthis (Pers.) Kuntze
See: Puccinia orbicula Peck & Clinton
Dicaeoma punctatum (Link) Arthur
See: Puccinia punctata Link and P. punctata var. troglodytes (Lindr.) Arthur
Dicaeoma pustulatum (M.A. Curtis) Arthur
See: Puccinia andropogonis Schwein.
Dicaeoma ranunculi (Seym.) Kuntze
See: Puccinia andina Dietel & Neger
Dicaeoma rhamni (J.F. Gmel.) Kuntze, nom. ambig.
See: Puccinia coronata Corda, s.lat., P. coronata var. avenae f. sp. avenae Urban & Marková and P. coronati-calamagrostidis M. Liu & Hambl.
Dicaeoma ruelleae (Berk. & Broome) Kuntze
See: Puccinia lateripes Berk. & Ravenel
Dicaeoma sambuci (Schwein.) Arthur
See: Puccinia sambuci (Schwein.) Arthur
Dicaeoma saniculae (Grev.) Kuntze
See: Puccinia marylandica Lindr.
Dicaeoma silphii (Schwein.) Kuntze
See: Puccinia silphii Schwein.
Dicaeoma smilacis (Schwein.) Kuntze
See: Puccinia smilacis Schwein.
Dicaeoma sorghi (Schwein.) Kuntze
See: Puccinia sorghi Schwein.
Dicaeoma tenue (Schwein.) Kuntze
See: Puccinia tenuis Burrill
Dicaeoma thalictri (Chevall.) Kuntze
See: Tranzschelia thalictri (Chev.) Dietel
Dicaeoma triticinum (Erikss.) H.S. Jacks.
See: Puccinia persistens subsp. triticina (Erikss.) Z. Urb. & J. Marková
Dicaeoma troglodytes (Lindr.) H.S. Jacks.
See: Puccinia punctata var. troglodytes (Lindr.) Arthur
Dicaeoma urticae (Schumach.) Kuntze
This name as applied by Arthur (1899) is ambiguous and probably refers to Puccinia urticata
See: Puccinia urticata F. Kern
Dicaeoma verbenicola (Ellis & Kellerm.) Arthur [reported as ‘verbenicolum’]
See: Puccinia vilfae Arthur & Holw.
Dicaeoma vernoniae (Schwein.) Kuntze (reported by Jackson 1915, but this combination was never made by Kuntze)
See: Puccinia longipes Lagerh.
Dicaeoma vexans (Farl.) Kuntze
See: Puccinia vexans Farl.
Dicaeoma vilfae (Arthur & Holw.) Arthur
See: Puccinia vilfae Arthur & Holw.
Dicaeoma violae (Schumach.) Kuntze
See: Puccinia violae (Schumach.) DC.
Dicaeoma vulpinodis (Dietel & Holw.) Kuntze
See: Puccinia dioicae Magnus, s.lat.
Dicaeoma windsoriae (Schwein.) Kuntze
See: Puccinia windsoriae Schwein.
Dicaeoma xanthii (Schwein.) Kuntze
See: Puccinia xanthii Schwein.
Diorchidium lateripes Magnus
See: Puccinia lateripes var. strepentis G.F. Laundon
Earlea speciosa (Fr.) Arthur
See: Phragmidium speciosum (Fr.) Bonard
Endocronartium harknessii (J.P. Moore) Y. Hirats.
See: Cronartium harknessii E. Meinecke
Endophyllum tuberculatum (Ellis & Kellerm.) Arthur & Fromme
See: Pucciniosira tuberculata (Ellis & Kellerm.) Buriticá & J.F. Hennen
Frommea obtusa (F. Strauss) Arthur
See: Phragmidium tormentillae Fuckel
Frommeëlla duchesneae (Arthur) Yohem, Cummins & Gilb.
See: Phragmidium mexicanum (Mains) H.Y. Yun, Minnis & Aime
Frommeëlla mexicana var. indicae J.W. McCain & J.F. Hennen
See: Phragmidium mexicanum (Mains) H.Y. Yun, Minnis & Aime
Frommeëlla tormentillae (Fuckel) Cummins & Y. Hirats.
See: Phragmidium tormentillae Fuckel
Gymnoconia interstitialis (Schltdl.) Lagerh.
See: Gymnoconia peckiana (Howe) Trotter
Gymnoconia nitens (Schwein.) Arthur
Rubus allegheniensis cf. Porter
R. allegheniensis Porter (Rosaceae) (Jackson 1920)
R. argutus var. floridus (Tratt.) L.H. Bailey
R. flagellaris cf. Willd.
R. flagellaris Willd.; (=R. villosus Aiton) (Snyder 1898); (=R. procumbens Muhl.) (Jackson 1920)
R. frondosus Bigelow (=R. sativus Brainerd) (Jackson 1920)
R. hispidus L.
R. occidentalis L. (Jackson 1920)
Rubus sp.
Host not reported (Van Hook 1910)
Gymnoconia peckiana (Howe) Trotter
Rubus allegheniensis Porter (Rosaceae) (Jackson 1915; Anderson and Anderson 1919; Emmons et al. 1960)
R. flagellaris Willd. (=R. procumbens Muhl.) (Jackson 1915); (=R. villosus Aiton) (Underwood 1893; Snyder 1896; Arthur 1899; Anderson and Anderson 1919)
R. idaeus L. ssp. strigosus (Michx.) Focke (=Ru. strigosus Michx.) (Jackson 1915)
R. occidentalis L. (Underwood 1893; Arthur 1899; Jackson 1915; Anderson and Anderson 1919)
Rubus spp. (Underwood 1893; Pipal 1915; Ruhl et al. 1980; Ruhl et al. 1981)
Gymnosporangium bethelii Kern
Crataegus cerronis Nels. (Rosaceae) PUR: 10499
C. coccinea L. PUR: 10467
C. pringlei Sarg. PUR: 10510
C. punctata Jacq. PUR: 10511, 10512
Crataegus sp. PUR: 10525, 10521
C. chrysocarpa var. chrysocarpa Ashe (=C. coccinea L.)
C. phaenopyrum (L. f.) Medik. (=C. cordata (Mill.) Ait. nom. rej.) PUR: 10477, 10494, 10501
Sorbus americana Marshall (Rosaceae) PUR: 10531, 10532, 10533 [Doubtful record]
Gymnosporangium biseptatum Ellis
Amelanchier canadensis (L.) Medik. (Rosaceae)
A. ovalis Medik. (= A. vulgaris Moench) Medik.
Amelanchier × intermedia Spach (pro sp.) [arborea × canadensis] Spach.
Gymnosporangium clavariiforme (Jacq.) DC.
Amelanchier canadensis (L.) Medik. (Rosaceae) (McCain et al. 1990)
A. erecta Blanch.
Amelanchier × intermedia Spach (pro sp.) [arborea × canadensis] Spach.
Gymnosporangium clavipes Cooke & Peck
Amelanchier canadensis (L.) Medik. (Rosaceae)
A. erecta Blanch.
A. laevis × humilis (Jackson 1920)
Crataegus crus-galli L. (Rosaceae) PUR: N11564
C. mollis Scheele (Jackson 1915)
C. punctata Jacq.
C. sp. (Ruhl et al. 1983)
C. tomentosa Anon.
Cydonia oblonga Mill. (=C. vulgaris (L.) Pers.) (Rosaceae) (Jackson 1915; Pipal 1915; Ruhl et al. 1980)
Juniperus communis L. (Cupressaceae)
J. virginiana L. (Jackson 1920; Van Hook 1921; McCain and Hennen 1981)
Mespilus germanica L. (Rosaceae) (Walker and Earhart 1962)
Pyrus calleryana Decne. (Rosaceae) (Creswell et al. 2016)
Gymnosporangium confusum Plowr.
Crataegus chrysocarpa var. chrysocarpa Ashe (=C. coccinea L.) (Rosaceae)
Gymnosporangium corniculans Kern
Amelanchier canadensis (L.) Medik. (Rosaceae)
Gymnosporangium cornutum Arthur ex F. Kern
Sorbus americana Marshall (Rosaceae)
Gymnosporangium davisii Kern
Photinia melanocarpa (Michx.) K.R. Robertson & Phipps (=Aronia melanocarpa (Michx.) Ellis) (Rosaceae)
Gymnosporangium ellisii (Berkeley) Ellis
See: Gymnotelium myricatum (Schwein.) Arthur
Gymnosporangium exiguum Kern
Crataegus pringlei Sarg. (Rosaceae)
Gymnosporangium exterum Arthur & Kern
Gillenia stipulata (Muhl. ex Willd.) Baill. (Rosaceae) (Jackson 1920; USDA 1960)
Juniperus virginiana L. (Cupressaceae)
Gymnosporangium floriforme Thaxt.
Crataegus chrysocarpa var. chrysocarpa Ashe (=C. coccinea L.) (Rosaceae)
Gymnosporangium germinale (Schwein.) F. Kern
See: Gymnosporangium clavipes Cooke & Peck
Gymnosporangium globosum Farl.
Crataegus anduennae Sarg. (Rosaceae) (Jackson 1920) [Doubtful host]
C. chrysocarpa var. chrysocarpa Ashe (=C. coccinea L.) (Underwood 1893; Arthur 1899; Jackson 1915; Anderson and Anderson 1919)
C. crus-galli L. (Arthur 1899; Jackson 1915; Anderson and Anderson 1919)
C. erythropoda Ashe
Crataegus × lavallei Hérincq ex Lavallee
C. mollis Scheele (Arthur 1899; Jackson 1915)
C. pringlei Sarg. (Jackson 1915)
C. punctata Jacq. (Underwood 1893; Arthur 1899; Jackson 1915; Anderson and Anderson 1919) PUR: 10771
Crataegus sp. (Ruhl et al. 1980, 1981; Snyder 1896; Anderson and Anderson 1919)
C. subvillosa Schrad. ex Torr. & A. Gray (Snyder 1898)
Juniperus virginiana L. (Cupressaceae) (Underwood 1893; Arthur 1899; Jackson 1915; Cooke 1967)
Malus sylvestris (L.) Mill. (Rosaceae)
M. angustifolia (Aiton) Michx. (USDA 1960)
M. coronaria (L.) Mill. (=M. glaucescens Rehd.)
Pyrus calleryana (Rosaceae) (PURN24059)
Gymnosporangium inconspicuum Kern
Amelanchier erecta Blanch. (Rosaceae)
Gymnosporangium juniperi-virginianae Schwein.
Ben Davis Apple (Rosaceae) (Van Hook 1924)
Crataegus punctata Jacq. (Rosaceae)
C. chrysocarpa var. chrysocarpa Ashe (=C. coccinea L.)
Juniperus virginiana L. (Cupressaceae) (Underwood 1893; Snyder 1896; Arthur 1899; Jackson 1915; Anderson and Anderson 1919; Emmons et al. 1960; Ruhl et al. 1981, 1983)
Malus coronaria (L.) Mill. (Rosaceae) (Arthur 1899; Jackson 1915); (=Pyrus coronaria L.) (Underwood 1893; Snyder 1896)
M. ioensis (Alph. Wood) Britton (Jackson 1915)
Malus sp. [‘CRABAPPLE’] (Ruhl et al. 1980, 1981)
M. sylvestris (L.) Mill. (Ruhl et al. 1980, 1982, 1983); (=M. malus (L.) Britton and Pyrus malus L.) (Arthur 1899; Jackson 1915; Pipal 1915; Anderson and Anderson 1919)
Pyrus communis L. (Rosaceae) (Underwood 1893; Arthur 1899) [Doubtful host, likely a Malus sp.]
Host not reported (Van Hook 1910)
Gymnosporangium kernianum Bethel
Amelanchier canadensis (L.) Medik. (Rosaceae)
Gymnosporangium libocedri (P. Henn.) Kern
See: Gymnotelium blasdaleanum (Dietel & Holw.) Arthur
Gymnosporangium macropus Link
See: Gymnosporangium juniperi-virginianae Schwein.
Gymnosporangium nelsonii Arthur
Amelanchier canadensis (L.) Medik. (Rosaceae)
A. erecta Blanch.
A. ovalis (=vulgaris Moench) Medik.
Malus coronaria (L.) Mill. (Rosaceae)
M. ioensis (Alph. Wood) Britton
Sorbus americana Marshall (Rosaceae)
Gymnosporangium nidus-avis Thaxt.
Malus sylvestris (L.) Mill. (Rosaceae) (USDA 1960) [Doubtful host, likely an Amelanchier sp.]
Gymnosporangium speciosum Peck
See: Gymnotelium speciosum (Peck) Aime & McTaggart
Gymnosporangium trachysorum Kern
Crataegus cerronis Nels. (Rosaceae)
C. punctata Jacq.
Gymnotelium blasdaleanum (Dietel & Holw.) Arthur
Amelanchier ovalis (= A. vulgaris Moench) Medik. (Rosaceae)
Crataegus pringlei Sarg. (Rosaceae)
Gymnotelium myricatum (Schwein.) Arthur
Morella cerifera (L.) Small (=Myrica cerifera L.) (Myricaceae)
Gymnotelium speciosum (Peck) Aime & McTaggart
Philadelphus coronarius L. (Saxifragaceae)
Hyalopsora polypodii (Pers.) Magnus
Cystopteris fragilis (L.) Bernh. (Cystopteridaceae) (Underwood 1893; Jackson 1915; Emmons et al. 1960)
Kuehneola obtusa (F. Strauss) Arthur
See: Phragmidium tormentillae Fuckel
Kuehneola uredinis (Link) Arthur
Rubus allegheniensis Porter (Rosaceae) (Jackson 1915; Anderson and Anderson 1919; Emmons et al. 1960)
R. cuneifolius Pursh (Underwood 1893; Arthur 1899; Jackson 1915)
R. flagellaris Willd. (=R. procumbens Muhl.) (Jackson 1915); (=R. villosus Aiton) (Underwood 1893; Arthur 1899)
R. hispidus L. (Jackson 1915)
Rubus sp. (Pipal 1915)
Melampsora abietis-caprearum Tubeuf
Salix sericea Marshall (Salicaceae) (USDA 1960)
Salix sp. (Cooke 1967)
Melampsora aff. abietis-caprearum
Note: Similar to M. abietis-caprearum but with amphigenous uredinia and telia
Salix interior Rowlee (PURN15351)
Melampsora abietis-canadensis C.A. Ludw. ex Arthur
[Considered as a taxonomic synonym for M. medusae (Cellerino 1999)]
Populus grandidentata Michx. (Salicaceae) (Underwood 1893; Arthur 1899; Jackson 1915)
P. heterophylla L. (Jackson 1915)
P. tremuloides Michx. (Jackson 1915)
P. deltoides W. Bartram ex Marshall(Jackson 1915)
Tsuga canadensis (L.) Carriere (Pinaceae)
Melampsora bigelowii Thüm.
See: Melampsora paradoxa Diet. et Holw.
Melampsora epitea Thüm., sensu Hylander et al. 1953
Salix cordata Michx. (Salicaceae) (Underwood 1893)
S. eriocephala Michx.
S. interior Rowlee; (=S. longifolia Muhl.) (Underwood 1893)
S. nigra Marshall (Underwood 1893)
S. sericea Marshall (reported as Salix sp.) (Snyder 1898)
Salix sp. (Snyder 1898)
Note: Melampsora epitea is a species complex comprised of at least 14 cryptic species in North America; without examination of the original material, it is not possible to determine to which precise species these reports refer (Bennett et al. 2011).
Melampsora euphorbiae (Schwein.) Castagne
Euphorbia commutata Engelm. ex A. Gray (Euphorbiaceae)
E. cyparissias L. (USDA 1960)
Melampsora euphorbiae-gerardianae W. Müll.
Euphorbia commutata Engelm. ex A. Gray (Euphorbiaceae); (=Tithymalus commutatus (Engelm. ex A. Gray) Klotzsch & Garcke) (Jackson 1917)
Euphorbia sp. (USDA 1960)
Melampsora farinosa (Pers.) J. Schröt., sensu Arthur 1899
Note: this is a European species and a synonym of M. caprearum. However, the concept defined by Arthur (1899) includes two different species viz. M. epitea and M. paradoxa and it is most likely these reports refer to one or both of these.
Salix cordata Michx. (Salicaceae) (Arthur 1899)
S. discolor Muhl. (Arthur 1899)
S. fluviatilis Nutt. (Arthur 1899)
S. nigra Marshall (Arthur 1899; Van Hook 1912)
Melampsora ferrinii Toome & Aime
Salix babylonica L. (Salicaceae)
Melampsora lini (Ehrenb.) Lév.
Linum usitatissimum L. (Linaceae)
Melampsora medusae Thüm.
Larix decidua Mill. (Pinaceae) (USDA 1960)
L. laricina (Du Roi) K. Koch
Populus balsamifera L. (Salicaceae) (Underwood 1893; Jackson 1915; Anderson and Anderson 1919; Emmons et al. 1960)
P. deltoides Bartram ex Marshall (Arthur 1899; Jackson 1915; Anderson and Anderson 1919; Newcombe et al. 2000); ssp. monilifera (Aiton) Eckenwalder (=P. monilifera Aiton) (Underwood 1893; Snyder 1896)
P. grandidentata Michx. (Jackson 1915)
P. nigra L. (McCain et al. 1990)
Populus sp. (Anderson and Anderson 1919)
P. tremuloides Michx. (Underwood 1893; Snyder 1898; Arthur 1899; Jackson 1915; Anderson and Anderson 1919)
Pseudotsuga menziesii (Mirb.) Franco (Pinaceae)
Host not reported (Van Hook 1910)
Melampsora paradoxa Dietel & Holw.
Larix decidua Mill. (Pinaceae) (USDA 1960)
Salix amygdaloides Andersson (Salicaceae) (Jackson 1915)
S. bebbiana Sarg. (McCain et al. 1990)
S. caroliniana Michx. (=S. wardii (Bebb) Bebb) (Jackson 1915)
S. cordata Michx. (Jackson 1915; Anderson and Anderson 1919)
S. discolor Muhl. (Underwood 1893; Snyder 1896; Jackson 1915; Anderson and Anderson 1919)
S. interior Rowlee (=S. longifolia Muhl.) (Jackson 1915; Anderson and Anderson 1919)
S. nigra Marshall (Anderson and Anderson 1919; Van Hook 1928)
Salix sp. (Anderson and Anderson 1919; USDA 1960)
Melampsora populina (Jacq.) Lév., sensu Underwood, 1893 [Followed by Arthur (1899)]
See: Melampsora abietis-canadensis C.A. Ludw. ex Arthur and M. medusae Thüm.
Melampsora populina (Jacq.) Lév.
See: Melampsora medusae Thüm.
Melampsora salicina (DC.) Lev., sensu Underwood, 1893
Note: This name as applied by Underwood (1893) could refer to either Melampsora epitea Thüm. or M. paradoxa Dietel & Holw.
Salix sp. (Salicaceae) (Underwood 1893)
Melampsora salicis-capreae (Pers.) G. Winter
See: Melampsora paradoxa Dietel & Holw.
Melampsora sp.
Populus sp. (Salicaceae) (Ruhl et al. 1981)
Melampsorella caryophyllacearum (DC.) J. Schröt.
See: Melampsorella elatina (Alb. & Schwein.) Arthur
Melampsorella elatina (Alb. & Schwein.) Arthur
Cerastium arvense subsp. strictum (L.) Ugborogho (Caryophyllaceae)
Melampsoridium betulae Arthur
See: Melampsoridium betulinum (Pers.) Kleb.
Melampsoridium betulinum (Pers.) Kleb.
Betula alleghaniensis Britton (Betulaceae); var. alleghaniensis Britton (=B. lutea Michx. f.) (Jackson 1915; Pipal 1915; McCain et al. 1990)
Micropuccinia andina (Dietel & Neger) Arthur & H.S. Jacks.
See: Puccinia andina Dietel & Neger
Milesia marginalis Faull & W.R. Watson
Dryopteris marginalis (L.) A. Gray (Dryopteridaceae)
Dryopteris sp.
Naohidemyces vaccinii (Jørst.) S. Sato, Katsuya & Y. Hirats. ex Vanderweyen & Fraiture
Tsuga canadensis (L.) Carrière (Pinaceae) (Jackson 1920; USDA 1960)
Nigredo amphidyma (Syd. & P. Syd.) Arthur
See: Uromyces amphidymus Syd. & P. Syd.
Nigredo appendiculata (Pers.) Arthur
See: Uromyces appendiculatus var. appendiculatus (Pers.) Unger and U. vignae Barclay
Nigredo caladii (Schwein.) Arthur
See: Uromyces ari-triphylli (Schwein.) Seeler
Nigredo caryophyllina (Schrank) Arthur
See: Uromyces dianthi (Pers.) Niessl
Nigredo fabae (Pers.) Arthur
See: Uromyces viciae-fabae (Pers.) J. Schröt.
Nigredo fallens (Desm.) Arthur
See: Uromyces trifolii-repentis var. fallens (Desm.) Arthur.
Nigredo hedysari-paniculati (Schwein.) Arthur
See: Uromyces hedysari-paniculati (Schwein.) Farl.
Nigredo howei (Peck) Arthur
See: Uromyces asclepiadis Cooke
Nigredo hyperici-frondosi (Schwein.) Arthur
See: Uromyces triquetrus Cooke
Nigredo lespedezae-procumbentis (Schwein.) Arthur
See: Uromyces lespedezae-procumbentis (Schwein.) M.A. Curtis
Nigredo medicaginis (Pass.) Arthur
See: Uromyces striatus J. Schröt.
Nigredo minuta (Dietel) Arthur
See: Uromyces minutus Dietel
Nigredo perigynia (Halst.) Arthur
See: Uromyces perigynius Halst.
Nigredo plumbaria (Peck) Arthur
See: Uromyces plumbarius Peck
Nigredo polygoni (Pers.) Arthur
See: Uromyces polygoni-avicularis (Pers.) G.H. Otth
Nigredo polemonii (Peck) Arthur
See: Uromyces acuminatus Arthur
Nigredo proëminens (DC.) Arthur
See: Uromyces proëminens (DC.) Lév.
Nigredo rhynchosporae (Ellis) Arthur [as ‘rhyncosporae’]
See: Uromyces rhynchosporae Ellis
Nigredo scirpi (Castagne) Arthur
See: Uromyces lineolatus subsp. nearcticus Savile
Nigredo silphii (Burrill) Arthur
See: Uromyces silphii (Syd. & P. Syd.) Arthur
Nigredo spermacoces (Schwein.) Arthur
See: Uromyces spermacoces (Schwein.) M.A. Curtis
Nigredo trifolii (R. Hedw.) Arthur
Note: As applied by Arthur (N. American Flora 7: 255) this likely refers to U. trifolii-repentis (not U. trifolii).
See: Uromyces trifolii-repentis Liro and U. trifolii-repentis var. fallens (Arthur) Cummins
Nigredo valens (F. Kern) Arthur
See: Uromyces valens F. Kern
Phakopsora crotonis (Burrill) Arthur
Croton capitatus Michx. (Euphorbiaceae) (McCain et al. 1990)
C. monanthogynus Michx. (McCain et al. 1990; Buriticá 1999)
Phakopsora pachyrhizi Syd. & P. Syd.
Glycine max (L.) Merr. (Fabaceae)
Phragmidium americanum (Peck) Dietel
Rosa carolina L. (Rosaceae) (Jackson 1915)
R. rubiginosa L. (Jackson 1915)
R. setigera Michx. (Jackson 1915; Anderson and Anderson 1919)
R. virginiana Mill. (Jackson 1915)
Rosa spp. (Jackson 1915; Pipal 1915)
Phragmidium disciflorum (Tode) J. James
See: Phragmidium mucronatum (Pers.) Schltdl.
Phragmidium fragariae (DC.) Rossmann
See: Phragmidium tormentillae Fuckel
Phragmidium ivesiae Syd. & P. Syd.
Potentilla recta L. (Rosaceae) (Savile 1976)
Phragmidium mexicanum (Mains) H.Y. Yun, Minnis & Aime
Duchesnea indica (Andrews) Focke (Rosaceae) (Ruhl and McCain 1986)
Phragmidium montivagum Arthur
(Rosaceae) (PURN24227)
Phragmidium mucronatum (Pers.) Schltdl.
Rosa carolina L. (Rosaceae) (Underwood 1893; Arthur 1899)
R. humilis Marshall (Arthur 1899)
R. lucida Ehrh. (Underwood 1893)
R. setigera Michx. (Underwood 1893; Arthur 1899)
Rosa spp. (Underwood 1893; Jackson 1915; Pipal 1915; Lee 1929) PUR: 8197
Phragmidium rosae-setigerae Dietel
See: Phragmidium americanum (Peck) Dietel
Phragmidium speciosum (Fr.) Bonard
Rosa arkansana var. suffulta (Greene) Cockerell (=R. suffulta Greene) (Rosaceae)
R. carolina L. (Jackson 1915; Snyder 1896; Arthur 1899)
R. humilis Marshall (Arthur 1899)
R. nitida Willd.
Rosa sp. (Van Hook 1924)
R. virginiana Mill. (Jackson 1915)
Phragmidium subcorticium (Schrank) G. Winter
See: Phragmidium mucronatum (Pers.) Schltdl.
Phragmidium tormentillae Fuckel
Potentilla canadensis L. (Rosaceae) (Underwood 1893; Snyder 1896; Arthur 1899; Jackson 1915; Anderson and Anderson 1919; Jackson 1920; Van Hook 1923, 1929)
P. simplex Michx. (Emmons et al. 1960; Ruhl and McCain 1986)
Phragmidium triarticulatum Berk. & M.A. Curtis
See: Phragmidium tormentillae Fuckel
Pileolaria brevipes Berk. & Ravenel
Toxicodendron pubescens Mill. (=Rhus toxicodendron L.) (Anacardiaceae) (Underwood 1893; Snyder 1896; Anderson and Anderson 1919) [Doubtful host]
T. radicans (L.) Kuntze (=Rhus radicans L.) (Arthur 1899; Jackson 1915); subsp. radicans (=R. radicans L.) (Cooke 1967)
Pileolaria cotini-coggygriae Tai & Cheo
Cotinus coggygria Scop. (Kaishian et al. 2023; PUR058489)
Pileolaria toxicodendri (Berk. & Ravenel) Arthur
See: Pileolaria brevipes Berk. & Ravenel
Polythelis fusca Arthur
See: Tranzschelia pseudofusca M. Scholler & M. Abbasi
Polythelis thalictri (Chevall.) Arthur
See: Tranzschelia thalictri (Chev.) Dietel
Prospodium transformans (Ellis & Everh.) Cummins
Tecoma stans (L.) Juss. ex Kunth (Bignoniaceae)
Puccinia acetosae (Schumach.) Körn.
Rumex acetosella L. (Polygonaceae) (Jackson 1917; USDA 1960)
Puccinia albiperidia Arthur
See: Puccinia caricina DC., s.lat.
Puccinia aletridis Berk. & M.A. Curtis
Aletris farinosa L. (Nartheciaceae) (Jackson 1915; USDA 1960)
Puccinia amphigena Dietel
Ammophila breviligulata Fernald (Poaceae) (USDA 1960)
Calamovilfa longifolia (Hook.) Scribn. (Poaceae) (Cummins and Greene 1961); var. magna Scribn. & Merr. (McCain et al. 1990)
Smilax herbacea L. (Smilacaceae) (Cummins and Greene 1961)
S. tamnoides L. (=hispida Muhl. ex Torr.) (Cummins and Greene 1961)
Puccinia andina Dietel & Neger
Ranunculus hispidus var. nitidus (Chapm.) T. Duncan (=R. septentrionalis Poir.) (Ranunculaceae) (Arthur 1899; Jackson 1915; USDA 1960)
Puccinia andropogi Schwein.
See: Puccinia andropogonis Schwein
Puccinia andropogonis Schwein.
Aesculus glabra Willd. (Hippocastanaceae)
A. hippocastanum L.
Andropogon gerardii Vitman (Poaceae); (=A. furcatus Muhl. ex Willd.) (Snyder 1896; Arthur 1899; Jackson 1915)
Andropogon sp. (Underwood 1893)
Comandra umbellata (L.) Nutt. (Santalaceae) (Underwood 1893; Jackson 1915; Anderson and Anderson 1919)
Orbexilum onobrychis (Nutt.) Rydb. (=Psoralea onobrychis Nutt.) (Fabaceae) (Underwood 1893)
Penstemon digitalis Nutt. ex Sims (Plantaginaceae)
P. hirsutus (L.) Willd. (Jackson 1915; USDA 1960) PUR: 88314
P. laevigatus Aiton; (=P. penstemon (L.) Britton) (Jackson 1920)
P. glaber var. alpinus (Torr.) A. Gray (=P. alpinus Torr.)
P. pubescens Aiton [as ‘Pentstemon pubescens’] (Snyder 1896)
Schizachyrium scoparium (Michx.) Nash (Poaceae) (Jackson 1915); (=Andropogon scoparius Michx., as [‘scoparus’]) (Snyder 1896; Arthur 1899)
Zanthoxylum americanum Mill. (Rutaceae) (USDA 1960; McCain et al. 1990)
Puccinia andropogonis var. pentastemonis (Schwein.) Arthur
See: Puccinia andropogonis Schwein.
Puccinia anemones-virginianae Schwein.
Anemone canadensis L. (Ranunculaceae) (Cooke 1967; McCain and Hennen 1981)
A. cylindrica A. Gray (Snyder 1896; Arthur 1899; Jackson 1915)
Anemone sp.
A. virginiana L. (Jackson 1915; Anderson and Anderson 1919)
Puccinia angelicae (Schumach.) Fuckel
Taenidia integerrima (L.) Drude (Apiaceae) (Jackson 1915; USDA 1960)
Puccinia angustata Peck
Eriophorum virginicum L. (Cyperaceae)
E. viridicarinatum (Engelm.) Fernald
Lycopus americanus Muhl. ex W.P.C. Barton (=L. sinuatus Elliott) (Lamiaceae) (Snyder 1898; Jackson 1915)
Lycopus sp. (Underwood 1893)
L. uniflorus Michx. (Jackson 1915)
Pycnanthemum virginianum (L.) T. Dur. & B.D. Jacks. ex B.L. Rob. & Fernald (Lamiaceae) (USDA 1960); (=P. lanceolatum Pursh); (=Koellia virginiana (L.) MacMill.) (Jackson 1920)
Scirpus atrovirens Willd. (Cyperaceae) (Underwood 1893; Snyder 1896; Arthur 1899; Jackson 1915; Anderson and Anderson 1919; Savile 1972)
S. cyperinus (L.) Kunth (Arthur 1899); (=Eriophorum cyperinum L.) (Underwood 1893)
Scirpus sp.
Puccinia anomala Rostr.
See: Puccinia hordei G.H. Otth
Puccinia antirrhini Dietel & Holw.
Antirrhinum majus L. (Plantaginaceae) (Jackson 1915; Pipal 1915; Anderson and Anderson 1919) PUR: 42777, 42487, 42489,42490, 42492, 42493
Antirrhinum sp. PUR: N10830
Puccinia apocrypta Ellis & Tracy
Asperella hystrix (L.) Humb. in Roem. & Usteri (Poaceae) (Underwood 1893)
Elymus hystrix L. (=Hystrix patula Moench) (Poaceae) (USDA 1960); var. hystrix (=H. hystrix (L.) Millsp.) (Arthur 1899)
Puccinia argentata (Schultz) G. Winter
Impatiens capensis Meerb. (=I. fulva Nutt.) (Balsaminaceae) (Snyder 1896); (=I. biflora Walter) (Arthur 1899; Jackson 1915)
I. flava (Underwood 1893) [Doubtful host, likely refers to I. fulva]
I. capensis Meerb. (reported as I. flava) (Underwood 1893)
I. pallida Nutt. (USDA 1960)
Puccinia aristidae Tracy
See: Puccinia subnitens Dietel
Puccinia arundinariae Schweinitz
Smilax sp. (Smilacaceae) (Cummins 1971) PUR: 45961
Puccinia asparagi DC.
Allium cepa L. (Amaryllidaceae) (Ruhl et al. 1983) [Doubtful host]
Asparagus officinalis L. (Asparagaceae) (Jackson 1915; Anderson and Anderson 1919; Ruhl et al. 1980)
Asparagus sp. (Pipal 1915)
Host not reported (Van Hook 1910)
Puccinia asperifolii (Pers.) Wettst. [as’ asperifolia’ by Pipal (1915)]
See: Puccinia recondita Roberge ex Desm. s.str.
Puccinia asteris Duby
See: Puccinia cnici-oleracei Pers. Ex Desm.
Puccinia asterum (Schwein.) F. Kern
See: Puccinia dioicae Magnus, s.lat.
Puccinia atrofusca (Dudley & C.H. Thomps.) Holw.
Artemisia (=Dracunculoides L.) dracunculus L. (Asteraceae)
Artemisia sp.
Puccinia bardanae (Wallr.) Corda
Arctium lappa L. (Asteraceae) (Jackson 1915; Anderson and Anderson 1919) [Doubtful host, likely refers to A. minus]
A. minus (Hill) Bernh. (Savile 1970); PUR: N2751, 38010, 38119, 38120, 38124, 38125, 38126, 38141, 38143, 38154, 49274, 61540, 88329, 89269, N1943, N4508, N4509, N6319
Puccinia bolleyana Saccardo
See: Puccinia sambuci (Schwein.) Arthur
Puccinia brachypodii G.H. Otth
See: Puccinia poae-nemoralis G.H. Otth
Puccinia brachypodii var. arrhenatheri (Kleb.) Cummins & H.C. Greene
See: Puccinia magelhaenica Peyr.
Puccinia brachypodii var. poae-nemoralis (G.H. Otth) Cummins & H.C. Greene
See: Puccinia poae-nemoralis G.H. Otth
Puccinia bromina Erikss., s.lat.
Bromus pubescens Muhl. ex Willd. (Poaceae) PUR: 25031, 25039, 25041
B. secalinus L. PUR: 25083
Puccinia calcitrapae var. bardanae (Wallr.)
See: Puccinia bardanae (Wallr.) Corda
Puccinia calcitrapae var. centaureae (DC.) Cummins
See: Puccinia laschii var. laschii Lagerh.
Puccinia calthae Link
Caltha palustris L. (Ranunculaceae) (Jackson 1915)
Puccinia canaliculata (Schwein.) Lagerh.
Ambrosia trifida L. (Asteraceae) (1, Jackson 1915; Kern 1919)
Cyperus esculentus L. (Cyperaceae) (Jackson 1915; Kern 1919; Van Hook 1929)
C. filiculmis Vahl. (Jackson 1915)
C. odoratus L.; (=Cyperus engelmannii Steud.) (Cyperaceae) (Jackson 1915); (=C. speciosus Vahl) (Jackson 1920)
Cyperus sp.
C. strigosus L. (Snyder 1896; Jackson 1915; Kern 1919)
C. schweinitzii Torr. (Jackson 1915)
Xanthium strumarium L. var. canadense (Mill.) Torr. & A. Gray (=X. commune Britton) (Asteraceae) (Jackson 1915); var. glabratum (DC.) Cronquist (=X. americanum Walter) (Jackson 1915)
Xanthium sp. (Kern 1919)
Puccinia caricis (Schumach.) Rebent.
See: Puccinia caricina DC.
Puccinia caricina DC., s.lat.
Carex blanda Dewey (Cyperaceae)
C. bullata Schkuhr ex Willd. (Cyperaceae) (Underwood 1893)
C. cephalophora Muhl. ex Willd. (Jackson 1920)
C. conoidea Schkuhr ex Willd. (Jackson 1920)
C. crinita Lam. (Jackson 1915)
C. davisii Schwein & Torr. (McCain and Hennen 1981)
C. digitalis Willd. (Jackson 1915)
C. emoryi Dewey
C. frankii Kunth (Snyder 1898)
C. gracilescens Steud. (McCain and Hennen 1981)
C. hirtifolia Mack. (Jackson 1915) PUR: 27716; (=C. pubescens Muhl. ex Schkuhr) (Anderson and Anderson 1919; Jackson 1920)
C. hitchcockiana Dewey (Jackson 1915)
C. laxiflora Lam. (Jackson 1915)
C. lupuliformis Sartwell ex Dewey (Jackson 1920)
C. lurida Wahlenb. (Underwood 1893)
C. rosea Schkuhr ex Willd.
C. siccata Dewey (=Carex foenea Willd.) (Underwood 1893)
Carex sp. (Snyder 1896)
C. sparganioides Muhl. ex Willd. (Jackson 1920)
C. squarrosa L. (Jackson 1915)
C. stricta Lam. (Jackson 1915)
C. tetanica Schkuhr (Jackson 1915)
C. trichocarpa Muhl. ex Willd.
Ribes americanum Mill. [as’americana’] (Grossulariaceae) (Jackson 1920)
R. aureum Pursh
R. cynosbati L. (Underwood 1893; Snyder 1898; Anderson and Anderson 1919; McCain and Hennen 198; PUR: 27120 – Greenhouse)
R. cynosbati (=Grossularia cynosbati (L.) Mill.) (Jackson 1915)
R. floridum L’Herit (Anderson and Anderson 1919) [Doubtful host]
R. hirtellum Michx.; (=G. hirtella (Michx.) Spach) (Jackson 1920)
R. missouriense Nutt. (=G. missouriensis (Nutt.) Coville & Britton) (Jackson 1915)
R. oxyacanthoides L. ssp. oxyacanthoides (=G. oxyacanthoides (L.) Mill.) (Jackson 1915; Anderson and Anderson 1919); ssp. setosum (Lindl.) Sinnott (=G. setosa (Lindl.) Coville & Britton) (Jackson 1915)
R. rotundifolium Michx. (Underwood 1893); (=G. rotundifolia (Michx.) Coville & Britton) (Jackson 1915)
R. rubrum L. (=R. vulgare Lam.) (Jackson 1920)
Ribes sp. (Van Hook 1929)
R. uva-crispa L.
Puccinia caricina var. limosae (Magnus) Jørst.
Lysimachia thyrsiflora L. (=Naumburgia thyrsiflora (L.) Duby) (Primulaceae) (Jackson 1917)
Puccinia caricis-erigerontis Arthur
See: Puccinia dioicae Magnus, s.lat.
Puccinia caulicola Tracy & Gall.
Salvia lanceolata Lam. (Lamiaceae)
Puccinia chloridis Speg.
Asclepias incarnata L. (Apocynaceae)
A. syriaca L.
Puccinia chrysanthemi Roze
See: Puccinia tanaceti var. tanaceti DC.
Puccinia circaeae Pers.
Circaea lutetiana L. (Onagraceae) (Underwood 1893; Snyder 1896; Arthur 1899; Jackson 1915; Anderson and Anderson 1919); subsp. canadensis (L.) Asch. & Magnus (=C. quadrisulcata (Maxim.) Franch. & Savigny) (Emmons et al. 1960)
Puccinia cirsii Lasch
See: Puccinia laschii var. laschii Lagerh.
Puccinia clematidis (DC.) Lagerh., nom. ambig.
Anemone virginiana L. (Ranunculaceae) (Jackson 1920)
Pascopyrum smithii (Rydb.) Á. Löve (=Agropyron smithii Rydb.) (Poaceae) (Jackson 1920)
Puccinia cnici var. cnici H. Mart.
Cirsium lanceolatum Hill (Asteraceae) (Jackson 1915; Anderson and Anderson 1919)
C. vulgare (Savi) Ten. (Savile 1970)
Puccinia cnici-oleracei Pers. ex Desm.
Aster sp. (Asteraceae) (Underwood 1893; Snyder 1898; Anderson and Anderson 1919)
Calendula officinalis L. (Asteraceae) PUR: 42243 - Greenhouse
Dimorphotheca cuneata (Thunb.) Less. (Asteraceae)*
Symphyotrichum cordifolium (L.) G.L. Nesom (=Aster cordifolius L.) (Asteraceae) (Underwood 1893; Arthur 1899; Jackson 1915; Anderson and Anderson 1919); (=A. sagittifolius Wedemeyer ex Willd.) (Arthur 1899; Jackson 1915)
S. lanceolatum (=A. paniculatus Lam.) (Arthur 1899); subsp. lanceolatum var. lanceolatum (Willd.) G.L. Nesom (=A. paniculatus Lam.) (Underwood 1893; Jackson 1915; Anderson and Anderson 1919); subsp. lanceolatum var. lanceolatum (=A. diffusus) (Snyder 1896)
S. lateriflorum var. lateriflorum (L.) Á. Löve & D. Löve (=A. lateriflorus (L.) Britton) (Arthur 1899; Jackson 1920)
S. novae-angliae (L.) G.L. Nesom (=A. novae-angliae L.) (Jackson 1915)
S. novi-belgii (L.) G.L. Nesom var. novi-belgii (=A. longifolius Lam.) (Jackson 1915)
S. oolentangiense var. oolentangiense (Riddell) G.L. Nesom (=A. azureus Lindl.) (Jackson 1915)
S. praealtum var. praealtum (Poir.) G.L. Nesom (=A. salicifolius Aiton) (Arthur 1899; Jackson 1920)
S. puniceum var. puniceum (L.) Á. Löve & D. Löve (=A. puniceus L.) (Jackson 1915)
S. shortii (Lindl.) G.L. Nesom (=A shortii Lindl.) (Arthur 1899)
Puccinia cockerelliana Bethel ex Arthur
Thalictrum dioicum L. (Ranunculaceae)
Puccinia conoclinii var. conoclinii Seym. ex. Burrill
Conoclinium coelestinum (L.) DC. (=Eupatorium coelestinum (L.) DC) (Asteraceae) (Jackson 1915)
Eupatorium sp. (Asteraceae) (USDA 1960)
Fleischmannia incarnata (Walter) R.M. King & H. Rob. (=E. incarnatum Walter) (Asteraceae) (Jackson 1920; Cummins et al. 1969)
Puccinia convolvuli (Pers.) Castagne
Calystegia sepium (L.) R. Br. (Convolvulaceae) (Jackson 1915); (=Convolvulus sepium L.) (Underwood 1893; Snyder 1896; Arthur 1899; Anderson and Anderson 1919); [reported as Polygonum dumetorum L.] (Snyder 1896)
Convolvulus arvensis L. (Convolvulaceae) (USDA 1960)
Convolvulus sp.
Puccinia coronata Corda, s.lat.
Alopecurus aequalis Sobol. (Poaceae) (USDA 1960); (= A. aristulatus Michx.) (Jackson 1920)
Cinna arundinacea L. (Poaceae) (Jackson 1915; USDA 1960)
Frangula caroliniana (Walter) A. Gray (=Rhamnus caroliniana Walter) (Rhamnaceae) (Jackson 1915; USDA 1960)
F. purshiana (DC.) A. Gray
Hordeum jubatum L. (Poaceae)
Rhamnus alnifolia L’Hér. (Rhamnaceae) (Jackson 1920)
R. cathartica L. (Van Hook 1925)
R. lanceolata Pursh (Arthur 1899; USDA 1960)
Host not reported (Van Hook 1910)
Also see: Puccinia coronata var. avenae f. sp. avenae Urban & Marková, P. coronati-brevispora M. Liu & Hambl.
Puccinia coronata var. avenae f. sp. avenae Urban & Marková
Avena sativa L. (Poaceae) (Snyder 1896, 1898; Arthur 1899; Jackson 1915; Pipal 1915; Anderson and Anderson 1919; PUR: N1190 – Greenhouse)
Lolium perenne L. (Poaceae)
Lolium sp. (Poaceae)
Poa pratensis L. (Poaceae)
Puccinia coronatae-brevispora M. Liu & Hambl.
Bromus sp. (Poaceae) PUR: N11047
Puccinia coronata var. avenae Fraser & Ledinham
See: Puccinia coronata var. avenae f. sp. avenae Urban & Marková
Puccinia coronatae-calamagrostidis M. Liu & Hambl. [as ‘ coronati-calamagrostidis ’]
Calamagrostis canadensis (Michx.) P. Beauv. (Poaceae) (Underwood 1893; Arthur 1899; Jackson 1915)
Puccinia crandallii Pammel & H.H. Hume
Symphoricarpos albus (L.) S.F. Blake (Caprifoliaceae)
Puccinia cyani var. cyani Pass.
Centaurea cyanus L. (Asteraceae) (USDA 1960; Savile 1970)
Puccinia cyperi Arthur
Conyza canadensis (L.) Cronquist (Asteraceae)
Cyperus echinatus (L.) Alph. Wood (=C. ovularis (Michx.) Torr.) (Cyperaceae) (McCain et al. 1990)
C. lupulinus subsp. lupulinus (Spreng.) Marcks (=C. filiculmis Vahl) (Kern 1919; Jackson 1920); subsp. macilentus (Fernald) Marcks (=C. filiculmis var. macilentus Fernald) (McCain et al. 1990)
C. schweinitzii Torr. (Kern 1919; Jackson 1920)
Cyperus sp. (Underwood 1893)
C. strigosus L. (Underwood 1893; Snyder 1896; Arthur 1899; Anderson and Anderson 1919; Kern 1919; Lopez-Franco and Hennen 1990)
Erigeron annuus (L.) Pers. (Asteraceae)
E. strigosus Muhl. ex Willd.
Puccinia cypripedii Arthur
Calopogon tuberosus (L.) Britton, Sterns & Poggenb. (Orchidaceae); var. tuberosus (=Limodorum tuberosum L.) (Jackson 1915)
Calopogon sp. (McCain et al. 1990)
Puccinia dayi Clinton
Lysimachia ciliata L. (Primulaceae); (=Steironema ciliatum (L.) Baudo) (Primulaceae) (Underwood 1893; Arthur 1899; Jackson 1915; Anderson and Anderson 1919)
Puccinia difformis Kunze
Galium aparine L. (Rubiaceae) (Arthur 1899; Jackson 1915)
Puccinia dioicae Magnus, s.lat.
Agoseris glauca (Pursh) Raf. (Asteraceae)
Brachychaeta sphacelata (Raf.) Britton (Asteraceae) (Jackson 1920)
Calylophus serrulatus (Nutt.) P.H. Raven (=Oenothera serrulata Nutt.) (Onagraceae)
Carex albicans Willd. ex Spreng. var. albicans (=C. varia Muhl. ex Willd.) (Cyperaceae) (Jackson 1920)
C. annectens (E.P. Bicknell) E.P. Bicknell (McCain and Hennen 1981; Schoknecht 1981)
C. brevior (Dewey) Mack.
C. cephaloidea (Dewey) Dewey (Jackson 1915)
C. cephalophora Muhl. ex Willd. (Jackson 1915)
Carex cf. frankii Kunth
C. communis L.H. Bailey (McCain and Hennen 1981)
C. conoidea Schkuhr ex Willd. (Jackson 1915)
C. festucacea Schkuhr ex Willd. (Jackson 1915)
C. jamesii Schwein.
C. laeviculmis Meinsh. (Cyperaceae)
C. laeviconica Dewey (Jackson 1920)
C. lanuginosa Michx. (Jackson 1915)
C. lupuliformis Sartwell ex Dewey (Jackson 1920)
C. muskingumensis Schwein. (Jackson 1920)
C. nebrascensis Dewey (Jackson 1915)
C. normalis Mack. (Jackson 1920)
C. oligocarpa Schkuhr ex Willd. (Jackson 1915)
C. pellita Muhl. ex Willd. (=C. lanuginosa auct. non Michx.)
C. pensylvanica Lam. (Jackson 1915; USDA– inoculation using aeciospores from Dirca palustris)
C. rosea Schkuhr (McCain and Hennen 1981)
C. siccata Dewey (=C. foenea Willd.) (Jackson 1915)
Carex sp. (Underwood 1893; Jackson 1915; Cooke 1967)
C. sparganioides Muhl. ex Willd. (Snyder 1896; Arthur 1899; Arthur 1909; Jackson 1915)
C. stenolepis Less. ex Steud. (Underwood 1893)
C. stipata Muhl. ex Willd. (Arthur 1909; Jackson 1915; Jackson 1920)
C. straminea Willd. ex Schkuhr (Underwood 1893; Jackson 1915)
C. tetanica Schkuhr (Jackson 1915)
C. tribuloides Wahlenb. (Jackson 1920)
C. trichocarpa Muhl. ex Willd. (Jackson 1915)
C. umbellata Schkuhr ex Willd. (=C. abdita E.P. Bicknell) (Jackson 1920)
C. virescens Muhl. ex Willd. (Underwood 1893)
C. vulpinoidea Michx. (Cyperaceae) (Underwood 1893; Snyder 1896; Arthur 1899; Arthur 1909; Jackson 1915)
C. willdenowii Schkuhr ex Willd. (Jackson 1920)
Conyza canadensis var. canadensis (L.) Cronquist (=Erigeron canadensis L.) (Asteraceae); (=Leptilon canadense (L.) Britton) (Jackson 1915)
Dirca palustris L. (Thymelaeaceae) (Underwood 1893; Jackson 1917; Anderson and Anderson 1919; Van Hook 1927) PUR: 24 specimens
Dulichium arundinaceum (L.) Britton (Cyperaceae) (Jackson 1915)
D. spathaceum Rich. (Underwood 1893)
Erigeron annuus (L.) Pers. (Asteraceae) (Jackson 1915; Anderson and Anderson 1919) PUR: 28800
E. pulchellus Michx. (Jackson 1920)
E. strigosus var. strigosus Muhl. ex Willd. (=E. ramosus (Walter) Britton, Sterns & Poggenb.) (Jackson 1915)
Euthamia graminifolia (L.) Nutt. (Asteraceae)
Lactuca canadensis L. (Asteraceae) (Jackson 1915; Anderson and Anderson 1919; Jackson 1920)
L. floridana (L.) Gaertn. (Jackson 1915) [Doubtful host, likely is L. canadensis]
L. sativa L. (Jackson 1915; USDA 1960)
L. virosa L. (Jackson 1915)
Oenothera biennis L. (Onagraceae) (Underwood 1893; Snyder 1896; Jackson 1915; Anderson and Anderson 1919)
Phryma leptostachya L. (Phrymaceae)
Sericocarpus sp. (Asteraceae) (McCain et al. 1990)
Solidago altissima L. (Asteraceae) (Jackson 1920)
S. arguta Aiton (Anderson and Anderson 1919)
S. caesia L. (Underwood 1893; Jackson 1915; Anderson and Anderson 1919)
S. canadensis L. (Underwood 1893; Jackson 1915; Savile 1972)
S. flexicaulis L. (Jackson 1915); (=S. latifolia L.) (Underwood 1893)
S. gigantea Aiton (=S. serotina Aiton) (Jackson 1915)
S. juncea Aiton
S. patula Muhl. ex Willd. (Jackson 1915) PUR: 88315
S. rugosa Mill.
Solidago sp. (Underwood 1893; McCain and Hennen 1981)
S. ulmifolia Muhl. ex Willd. (Jackson 1915)
S. missouriensis var. fasciculata Holz. (=S. glaberrima M. Martens)
Symphyotrichum cordifolium (L.) G.L. Nesom (=Aster cordifolius L.) (Asteraceae) (Underwood 1893; Jackson 1915; Anderson and Anderson 1919); (=A. sagittifolius Wedemeyer ex Willd.) (Underwood 1893; Jackson 1915; Anderson and Anderson 1919)
S. drummondii var. drummondii (Lindl.) G.L. Nesom (=A. drummondii Lindl.) (Jackson 1915)
S. ericoides var. ericoides (L.) G.L. Nesom (=A. ericoides L.)
S. foliaceum var. canbyi (A. Gray) G.L. Nesom (=A. tweedyi Rydb.)
S. lanceolatum subsp. lanceolatum var. lanceolatum (Willd.) G.L. Nesom (=A. paniculatus Lam.) (Jackson 1915)
S. praealtum var. praealtum (Poir.) G.L. Nesom (=A. salicifolius Aiton, non Lam.)
S. puniceum var. puniceum (L.) Á. Löve & D. Löve (=A. puniceus L.) (Jackson 1920)
S. shortii (Lindl.) G.L. Nesom (=A. shortii Lindl.) (Anderson and Anderson 1919)
Puccinia dispersa Erikss. & Henning
See: Puccinia recondita Roberge ex Desm. s.str.
Puccinia distichlidis Ellis & Everh.
Lysimachia ciliata L. (Primulaceae)
Puccinia dochmia Berk. & M.A. Curtis
Note: Doubtful record, likely refers to Puccinia schedonnardi
Muhlenbergia diffusa (Muhl.) Farw. (Poaceae) (Underwood 1893; Arthur 1899)
M. sylvatica (Torr.) Torr. ex A. Gray (Arthur 1899)
Puccinia dulichii P. Syd. & Syd.
See: Puccinia dioicae Magnus, s.lat.
Puccinia eatoniae Arthur
Myosotis verna Nutt. (=M. virginica (L.) Britton, Sterns & Poggenb.) (Boraginaceae) (USDA 1960)
Ranunculus abortivus L. (Ranunculaceae) (Underwood 1893; Jackson 1915; Anderson and Anderson 1919)
Sphenopholis intermedia (Rydb.) Rydb. (Poaceae)
S. obtusata (Michx.) Scribn.
Sphenopholis sp.
Sphenopholis × pallens (Biehler) Scribn. (pro sp.) [obtusata × pensylvanica] (Jackson 1915)
Puccinia eleocharidis Arthur
Ageratina altissima (L.) R.M. King & H. Rob. (Asteraceae)
Eupatorium maculatum (L.) E.E. Lamont (Asteraceae) (Jackson 1915; Kern 1919)
E. perfoliatum L. (Jackson 1915; Kern 1919)
E. purpureum (L.) E.E. Lamont (Snyder 1897; Jackson 1915; Kern 1919) PUR: 26729
Eleocharis obtusa (Willd.) Schult. (Cyperaceae) (Savile 1972; Jackson 1920; Kern 1919)
Eleocharis palustris (L.) Roem. & Schult. (Underwood 1893; Snyder 1896; Arthur 1899; Arthur 1909; Jackson 1915; Anderson and Anderson 1919; Kern 1919)
Puccinia ellisiana Thüm.
Schizachyrium scoparium (Michx.) Nash (Poaceae) (Jackson 1915)
Viola nuttallii Pursh (Violaceae)
V. palmata L. var. cucullata A. Gray (Underwood 1893)
V. sororia Willd.; (=Viola papilionacea Pursh) (Jackson 1915)
Puccinia emaculata Schwein.
Euphorbia corollata L. (Euphorbiaceae); (=Tithymalopsis corollata (L.) Klotzsch) (Jackson 1915)
E. marginata Pursh
Panicum capillare L. (Poaceae) (Underwood 1893; Snyder 1896; Arthur 1899; Jackson 1915; Anderson and Anderson 1919; Ramachar and Cummins 1965; Demers et al. 2017)
P. flexile (Gattinger) Scribn.
P. miliaceum L. (Jackson 1915; USDA 1960)
Panicum sp. (USDA 1960)
P. virgatum L. (Jackson 1915)
Puccinia epiphylla (L.) Wettst.
See: Puccinia poarum Nielsen
Puccinia eriophori Thüm.
Eriophorum angustifolium Honck. (Cyperaceae) (Jackson 1915)
E. virginicum L. (Jackson 1915)
Puccinia eumacrospora Cummins
Smilax tamnoides L. (=S. hispida Muhl. ex Torr.) (Smilacaceae)
Puccinia extensicola Plowr.
See: Puccinia dioicae Magnus, s.lat.
Puccinia extensicola var. asteris (Thüm.) Arthur
See: Puccinia dioicae Magnus, s.lat.
Puccinia extensicola var. hieraciata (Schwein.) Arthur
See: Puccinia dioicae Magnus, s.lat.
Puccinia extensicola var. solidaginis Arthur
See: Puccinia dioicae Magnus, s.lat.
Puccinia flaveriae H.S. Jacks.
See: Puccinia melampodii Dietel & Holw.
Puccinia flosculosorum Röhl.
See: Puccinia hieracii (Röhl.) H. Mart.
Puccinia fraxinata (Link) Arthur
See: Puccinia sparganioides Ellis & Barthol.
Puccinia galii (Pers.) Schwein.
See: Puccinia punctata Link and P. punctata var. troglodytes (Lindr.) Arthur
Puccinia gentianae (F. Strauss) Link
Gentiana andrewsii Griseb. (Gentianaceae)
Puccinia glechomatis DC.
Agastache nepetoides (L.) Kuntze (Lamiaceae) (Jackson 1915; Anderson and Anderson 1919)
Glechoma hederacea L. (Lamiaceae) (Scholler 2000; Böllmann and Scholler 2006)
Puccinia globosipes Peck
Lycium barbarum L. (=L. halimifolium Mill.) (Solanaceae) (Jackson 1915); (=halimifolium Mill.; vulgare Dunal) (Cash 1952)
Puccinia globosum (Farl.) Kuntze
See: Gymnosporangium globosum Farl.
Puccinia goldsbroughii M. Abbasi & Aime
Melica nitens (Scribn.) Nutt. ex Piper (Poaceae) (Abbasi and Aime 2016)
Melica sp. (Poaceae) (Abbasi and Aime 2016)
Puccinia graminicola (Burrill) Demers & Castl.
Panicum virgatum L. (Poaceae) (Arthur 1899; Jackson 1920; Kenaley et al. 2018) PUR: 11737
Unknown grasses (Poaceae) (Underwood 1893)
Puccinia graminis Pers.
Agrostis alba L. (Poaceae) (Jackson 1915; Anderson and Anderson 1919)
A. gigantea Roth
Agrostis sp. (Underwood 1893; Arthur 1899)
Alopecurus aequalis Sobol. (Poaceae) (USDA 1960); var. aequalis Sobol. (=A. aristulatus Michx.) (Jackson 1920)
A. pratensis L.
Arrhenatherum elatius (L.) P. Beauv. ex J. Presl & C. Presl (Poaceae) (Jackson 1915; USDA 1960)
Avena sativa L. (Poaceae) (Underwood 1893; Snyder 1896; Arthur 1899; Jackson 1915; Anderson and Anderson 1919)
Berberis canadensis Mill. (Berberidaceae)
B. vulgaris L. (Underwood 1893; Arthur 1899; Jackson 1915; Jackson 1920)
Bromus carinatus Hook. & Arn. (Poaceae)
B. ciliatus L.
B. kalmii A. Gray (=Bromus purgans L.) (Jackson 1920; USDA 1960)
B. marginatus Nees ex Steud. PUR: N5237 - Greenhouse
B. secalinus L. (Jackson 1915)
B. tomentellus Boiss. PUR: N5238 - Greenhouse
Cinna arundinacea L. (Poaceae) (Jackson 1915; USDA 1960)
Dactylis glomerata L. (Poaceae) (Snyder 1896; Arthur 1899; Jackson 1915; USDA 1960; Cooke 1986)
Elymus alaskanus subsp. latiglumis (Scribn. & J.G. Sm.) Á. Löve (Poaceae)
E. canadensis L. (Jackson 1920)
E. elymoides (Raf.) Swezey
E. repens (L.) Gould; (=Agropyron repens (L.) P. Beauv.) (Jackson 1920)
E. trachycaulus (Link) Gould ex Shinners
E. virginicus L. (Jackson 1920)
Festuca idahoensis Elmer (Poaceae)
Hordeum jubatum L. (Poaceae) (Snyder 1896; Arthur 1899; Jackson 1915) PUR: N1436
H. vulgare L. (Jackson 1915)
Hordeum sp. (Pipal 1915)
Koeleria macrantha (Ledeb.) Schult. (Poaceae)
Oryzopsis asperifolia Michx. (Poaceae) PUR: 62066 - Greenhouse
Pascopyrum smithii (Rydb.) Á. Löve (Poaceae)
Phalaris arundinacea L. (Poaceae)
Phleum pratense L. (Poaceae) (Jackson 1915; Pipal 1915; Anderson and Anderson 1919)
Poa alpina L. (Poaceae) PUR: 62060, 62062, N5161, N5262 - Greenhouse
P. annua L. PUR: N5263 - Greenhouse
P. compressa L. (Underwood 1893; Arthur 1899)
P. cusickii Vasey PUR: N5264 - Greenhouse
P. iridifolia Hauman PUR: N5265 - Greenhouse
P. pratensis L. (Underwood 1893; Arthur 1899; Abbasi et al. 2005)
P. secunda J. Presl PUR: 61061, 62067, 62069, N5272 N5280, N5281, N5282, N5283 - Greenhouse
P. palustris L. (USDA 1960)
Pseudoroegneria spicata (Pursh) Á. Löve (Poaceae)
Schedonorus pratensis (Huds.) P. Beauv. (Poaceae); (=Festuca elatior L.) (Jackson 1920)
Secale cereale L. (Poaceae) (Jackson 1915; Pipal 1915)
Trisetum spicatum (L.) K. Richt. (Poaceae) PUR: N5275 - Greenhouse
Triticum aestivum L. (Poaceae) (Ruhl et al. 1982; Abbasi et al. 2005); (=Triticum vulgare Vill.) (Underwood 1893; Snyder 1898; Arthur 1899; Jackson 1915; Pipal 1915; Anderson and Anderson 1919)
Host not reported (Van Hook 1910)
Puccinia graminis subsp. graminicola Z. Urb.
Poa pratensis L. (Poaceae)
Schedonorus arundinaceus (Schreb.) Dumort., nom. cons. (=S. phoenix (Scop.) Holub) (Poaceae)
Puccinia granulispora Ellis & Galloway
Allium cernuum Roth (Amaryllidaceae)
Puccinia grindeliae Peck
Gutierrezia sarothrae (Pursh) Britton & Rusby (Asteraceae)
Lygodesmia juncea (Pursh) D. Don ex Hook. (Asteraceae)
Solidago canadensis L. (Asteraceae)
Puccinia grossulariae Lagerh.
See: Puccinia caricina DC., s.lat.
Puccinia helianthi Schwein.
Helianthus annuus L. (Asteraceae) (Underwood 1893; Arthur 1899; Jackson 1915; Anderson and Anderson 1919)
H. cf. hirsutus
H. decapetalus L. (Jackson 1920); (=H. tracheliifolius Mill.) (Underwood 1893; Arthur 1899; Jackson 1915; Anderson and Anderson 1919)
H. divaricatus L. (Underwood 1893; Arthur 1899; Jackson 1915; Anderson and Anderson 1919; Emmons et al. 1960)
H. giganteus L. (Jackson 1915)
H. grosseserratus M. Martens (Underwood 1893; Snyder 1896; Arthur 1899; Jackson 1915; Anderson and Anderson 1919)
H. hirsutus Raf. (Jackson 1915; Anderson and Anderson 1919)
Helianthus × laetiflorus Pers. (pro sp.) [pauciflorus × tuberosus] (Jackson 1920; Emmons et al. 1960)
Helianthus × laetiflorus (=H. scaberrimus Elliott) (Jackson 1920)
H. maximiliani Schrad. (McCain et al. 1990)
H. microcephalus Torr. & A. Gray (McCain and Hennen 1981)
H. mollis Lam. (Jackson 1915)
H. occidentalis Riddell (Jackson 1915)
H. parviflorus Hornem. (Anderson and Anderson 1919)
H. pauciflorus Nuttall
H. petiolaris Nutt. (Jackson 1915)
Helianthus sp. (Underwood 1893)
H. s strumosus L. (Underwood 1893; Arthur 1899; Jackson 1915; Anderson and Anderson 1919)
H. tomentosus Michx.
H. tuberosus L. (Jackson 1915)
Helianthus × glaucus Small (pro sp.) [divaricatus × microcephalus]
Heliopsis helianthoides (L.) Sweet (Asteraceae) (Jackson 1915); var. scabra (Dunal) Fernald (=Heliopsis scabra Dunal) (Underwood 1893; Arthur 1899; USDA 1960)
Host not reported (Van Hook 1910)
Puccinia heliopsidis Schwein.
See: Puccinia helianthi Schwein.
Puccinia hemerocallidis Thüm.
Hemerocallis sp. (Liliaceae) PUR: N1322 (Hernández et al. 2002)
H. cv. ‘Strawberry Candy’ PUR: 11602
Puccinia heterospora Berk. & M.A. Curtis
Sida spinosa L. (Malvaceae)
Puccinia heucherae (Schwein.) Dietel
Saxifraga pensylvanica L. (=Micranthes pensylvanica (L.) Haw.) (Saxifragaceae) (Jackson 1915)
Mitella diphylla L. (Saxifragaceae)
Puccinia hibisciata (Schwein.) Kellerm.
See: Puccinia schedonnardi Kellerm. & Swingle
Puccinia hieracii (Röhl.) H. Mart.
Chrysanthemum sp. (Asteraceae)
Cnicus lanceolatus (L.) Willd. (=Carduus lanceolatus L.) (Asteraceae) (Underwood 1893; Arthur 1899) [Doubtful host]
Hieracium canadense Michx. (Asteraceae) (Jackson 1920)
H. gronovii L.
H. scabrum Michx. (Jackson 1915; Anderson and Anderson 1919)
Taraxacum dens-leonis Desf. (Asteraceae) (Snyder 1898) [Doubtful host]
T. officinale F.H. Wigg. (Underwood 1893; Snyder 1896; Anderson and Anderson 1919; Parmelee and Savile 1981); ssp. officinale (=Leontodon taraxacum L.) (Jackson 1915)
T. laevigatum (Willd.) DC. (=T. erythrospermum (Willd.) DC.) (Jackson 1915)
T. taraxacum H. Karst. (Arthur 1899)
Puccinia hieraciata (Schwein.) H.S. Jacks.
See: Puccinia dioicae Magnus, s.lat.
Puccinia hordei G.H. Otth
Hordeum vulgare L. (Poaceae) (Jackson 1920)
Ornithogalum umbellatum L. (Asparagaceae)
Puccinia hydnoidea (Berk. & M.A. Curtis) Arthur
See: Puccinia dioicae Magnus, s.lat.
Puccinia hydrophylli Peck & Clinton
Hydrophyllum virginianum L. (Hydrophyllaceae)
Puccinia hyssopi Schwein.
Agastache nepetoides (L.) Kuntze (Lamiaceae)
Puccinia impatientis (Schwein.) Arthur
Agrostis perennans (Walter) Tuck. (Poaceae) (Jackson 1915; Emmons et al. 1960)
Elymus canadensis L. (Poaceae) (Jackson 1915)
E. virginicus L. (Underwood 1893; Snyder 1896; Arthur 1899; Jackson 1915; Mains 1933); (=E. striatus Willd.) (Jackson 1915; Anderson and Anderson 1919)
E. hystrix L. (=Hystrix patula Moench) (USDA 1960); (=H. hystrix (L.) Millsp) (Jackson 1915)
Impatiens capensis Meerb. (Balsaminaceae); (=I. fulva Nutt.) (Underwood 1893)
I. pallida Nutt. (Snyder 1896; Jackson 1915; Van Hook 1928, 1929)
Impatiens sp.
I. capensis Meerb. (=I. biflora Walter) (Jackson 1915; Anderson and Anderson 1919)
Puccinia indusiata Dietel & Holw., ined.
See: Puccinia cyperi Arthur
Puccinia interstitialis Tranzsch.
See: Gymnoconia peckiana (Howe) Trotter
Puccinia interveniens (Peck) Bethel
Achnatherum nelsonii subsp. dorei (Barkworth & Maze) Barkworth (Poaceae)
Sidalcea candida A. Gray (Malvaceae)
Puccinia iridis Wallr.
Iris fulva Ker Gawl. (Iridaceae) (USDA 1960)
Iris sp. (USDA 1960)
I. versicolor L. (Jackson 1915; Anderson and Anderson 1919)
Iris × filifolia-xiphium Anon. (USDA 1960)
I. xiphioides Ehrh. (USDA 1960)
I. xiphium L. (USDA 1960)
Puccinia juniperi-virginianae (Schwein.) Arthur
See: Gymnosporangium juniperi-virginianae Schwein.
Puccinia kansensis Ellis & Barthol.
Buchloe dactyloides (Nutt.) Engelm. (Poaceae)
Physalis heterophylla Nees (Solanaceae) (Jackson 1917)
P. longifolia var. subglabrata (Mack. & Bush) Cronquist
Puccinia kuhniae Schwein.
Brickellia eupatorioides var. eupatorioides (L.) Shinners (=Kuhnia eupatorioides L.) (Asteraceae) (Underwood 1893; Snyder 1896; Arthur 1899; Jackson 1915; Jackson 1920; Cummins et al. 1969)
Kuhnia sp. (Asteraceae) (Cummins 1978)
Puccinia laschii var. laschii Lagerh.
Arctium minus (Hill) Bernh. (Asteraceae) [Doubtful host]
Cirsium sp. (Asteraceae)
C. hillii (Canby) Fernald (=C. pumilum subsp. hillii (Canby) J.W. Moore & Frankton (Savile 1970)
C. undulatum (Nutt.) Spreng. [as ‘undulatus’] (Jackson 1920)
Puccinia lateripes Berk. & Ravenel
Ruellia caroliniensis subsp. ciliosa var. cinerascens (Fernald) Kartesz & Gandhi (=R. ciliosa Pursh) (Acanthaceae) (Jackson 1915; Anderson and Anderson 1919; Laundon 1963)
R. strepens L. (Underwood 1893; Arthur 1899; Jackson 1915; Anderson and Anderson 1919; Laundon 1963)
Puccinia lateripes var. lateripes Berk. & Ravenel
Ruellia caroliniensis subsp. ciliosa var. cinerascens (Fernald) Kartesz & Gandhi (=R. ciliosa Pursh) (Acanthaceae) (Laundon 1963)
Puccinia lateripes var. strepentis G.F. Laundon
Ruellia strepens L. (Acanthaceae) (Snyder 1896; Laundon 1963)
Puccinia leptochloae Arthur & Fromme
Talinum paniculatum (Jacq.) Gaertn. (Portulacaceae)
Puccinia liatridis Bethel.
Liatris pycnostachya Michx. (Asteraceae) (USDA 1960)
Puccinia liliacearum Duby
Ornithogalum umbellatum L. (Asparagaceae) (Abbasi et al. 2016b)
Puccinia lithospermi Ellis & Kellerm.
Evolvulus nuttalliana R & S. (Convolvulaceae)
Puccinia lobeliae W.R. Gerard
Lobelia siphilitica L. (Campanulaceae) (Underwood 1893; Snyder 1896; Arthur 1899; Jackson 1915; Anderson and Anderson 1919)
Puccinia longipes var. brevipes (Dietel) Z. Urb.
Vernonia altissima (Asteraceae) (Arthur 1934)
Puccinia longipes var. longipes Lagerh.
Vernonia fasciculata Michx. (Asteraceae) (Underwood 1893; Arthur 1899; Jackson 1915; Cash 1952)
V. gigantea (Walter) Trel. (Urban 1971)
Vernonia sp. (Urban 1971)
Puccinia ludibunda Ellis & Everh.
See: Puccinia dioicae Magnus, s.lat.
Puccinia ludovicianae Fahrend.
Artemisia ludoviciana Nutt. (Asteraceae)
Puccinia lysimachiata (Link) Kern
See: Puccinia caricina var. limosae (Magnus) Jørst.
Puccinia magelhaenica Peyr.
Berberis fendleri A. Gray (Berberidaceae)
Koeleria macrantha (Ledeb.) Schult. (Poaceae)
Mahonia sp. (Berberidaceae)
Puccinia magnusiana Körn.
Anemone canadensis L. (Ranunculaceae)
Puccinia majanthae Arthur & Holw.
See: Puccinia sessilis J. Schröt.
Puccinia malvacearum Bertero ex Mont.
Alcea rosea L. (Malvaceae) (Jackson 1915); [as ‘Althaea’] (Ruhl et al. 1981, 1982, 1983; Pipal 1915; Anderson and Anderson 1919)
Malva neglecta cf. Wallr.
M. neglecta Wallr. (Malvaceae) (McCain and Hennen 1981)
M. rotundifolia L. (Jackson 1915; Lee 1929)
Host not reported (Van Hook 1910)
Puccinia mariae-wilsoniae G.W. Clinton [as ‘mariae-wilsoni’]
Claytonia virginica L. (Portulacaceae) (Jackson 1915; Anderson and Anderson 1919; Arthur 1934)
Host not reported (Van Hook 1934)
Puccinia marylandica Lindr.
Sanicula canadensis L. (Apiaceae) (Underwood 1893; Arthur 1899; Jackson 1915; Anderson and Anderson 1919; Jackson 1920; Emmons et al. 1960)
S. marilandica L. (USDA 1960)
Sanicula sp. (Cooke 1967)
Puccinia maydis Carr.
See: Puccinia sorghi Schwein.
Puccinia melampodii Dietel & Holw.
Calendula officinalis L. (Asteraceae) (USDA 1960)
Puccinia melicae P. Syd. & Syd.
Melica mutica Walter (Poaceae) (Jackson 1915)
Puccinia menthae Pers.
Blephilia ciliata (L.) Benth. (Lamiaceae) (Jackson 1915; USDA 1960)
B. hirsuta (Pursh) Benth. (Underwood 1893; Snyder 1896; Arthur 1899; Jackson 1915; Anderson and Anderson 1919; USDA 1960)
Cunila origanoides (L.) Britton (Lamiaceae) (Jackson 1915; Arthur 1899; Emmons et al. 1960); (=C. mariana L.) (Underwood 1893)
Hedeoma pulegioides (L.) Pers. (Lamiaceae)
Koellia flexuosa MacMill. (Lamiaceae) (Jackson 1920)
Mentha arvensis L. (=M. canadensis L.) (Lamiaceae) (Underwood 1893; Arthur 1899; Jackson 1915; Anderson and Anderson 1919)
Mentha × piperita L. (pro sp.) [aquatica × spicata]
M. spicata L. (Jackson 1915)
Monarda clinopodia L. (Lamiaceae)
M. didyma L.
M. fistulosa L. (Underwood 1893; Snyder 1896; Arthur 1899; Jackson 1915; Anderson and Anderson 1919; Emmons et al. 1960; Cooke 1967)
M. punctata L. (Jackson 1920)
Monarda sp. (Underwood 1893)
Pycnanthemum flexuosum (Walter) Britton, Sterns & Poggenb. (Lamiaceae)
P. lanceolatum Pursh (Underwood 1893)
P. muticum (Michx.) Pers. (Underwood 1893)
Pycnanthemum sp. (Snyder 1896)
P. virginianum (L.) T. Dur. & B.D. Jacks. ex B.L. Rob. & Fernald; (=Koellia virginiana (L.) MacMill.) (Arthur 1899; Jackson 1915)
P. verticillatum var. pilosum (Nutt.) Cooperr. (=P. pilosum Nutt.); (=Koellia pilosa (Nutt.) Britton) (Arthur 1899; Jackson 1915)
Puccinia minutissima Arthur
Decodon verticillatus (L.) Elliott (Lythraceae) (Jackson 1915; USDA 1960)
Carex lasiocarpa Ehrh. (Cyperaceae) (Jackson 1915)
Puccinia modiolae P. Syd & Syd.
Althaea sp. (Malvaceae) (Aime and Abbasi 2018)
Puccinia monoica Arthur
Arabis sp. (Brassicaceae)
Koeleria macrantha (Ledeb.) Schult. (Poaceae)
Trisetum spicatum (L.) K. Richt. (Poaceae)
Puccinia montanensis Ellis
Elymus canadensis L. (Poaceae) (Jackson 1915; USDA 1960; Cummins and Greene 1966)
E. hystrix L.
Elymus sp.
Puccinia muhlenbergiae Arthur & Holw.
See: Puccinia schedonnardi Kellerm. & Swingle
Puccinia nigrovelata Ellis & Tracy
See: Puccinia canaliculata (Schwein.) Lagerh.
Puccinia nolitangeris Corda [reported as ‘nolitangere’]
See: Puccinia argentata (Schultz) G. Winter
Puccinia novopanici Demers, Miao Liu & Hambl.
Panicum virgatum (Poaceae) PUR: N22686
Puccinia obliqua Berk. & M.A. Curtis
Gonolobus suberosus var. granulatus (Scheele) Krings & Q.Y. Xiang (Apocynaceae)
Puccinia obscura J. Schröt.
Luzula bulbosa (Alph. Wood) Smyth & Smyth (Juncaceae) (McCain et al. 1990)
L. campestris (L.) DC. (=Juncoides campestre (L.) Kuntze) (Jackson 1915; Anderson and Anderson 1919)
L. multiflora subsp. frigida (Buchenau) Krecz. (=L. intermedia (Thuill.) A. Nelson)
L. multiflora (Ehrh.) Lej.
Puccinia obtecta Peck
Bidens connata Muhl. ex Willd. (Asteraceae)
B. frondosa L. (Jackson 1920; USDA 1960; Savile 1972)
B. coronata (L.) Britton (=B. trichosperma (Michx.) Britton) (Jackson 1920)
Schoenoplectus lacustris (L.) Palla (=Scirpus lacustris L.) (Cyperaceae) (Arthur 1899)
S. pungens (Vahl) Palla
S. tabernaemontani (C.C. Gmel.) Palla (=Scirpus validus Vahl) (Jackson 1915; Anderson and Anderson 1919; Savile 1972)
S. americanus (Pers.) Volkart ex Schinz & R. Keller (=Scirpus americanus Pers.) (Jackson 1915; Savile 1972)
Puccinia orbicula Peck & Clinton
Prenanthes alba L. (Asteraceae) (Underwood 1893; Snyder 1896); (=Nabalus albus (L.) Hook.) (Arthur 1899; Jackson 1915)
Prenanthes sp. (Asteraceae)
Puccinia pammelii (Trel.) Arthur
Euphorbia corollata (Kenaley et al. 2018)
Puccinia panici Dietel
See: Puccinia emaculata Schwein.
Puccinia patruelis Arthur
See: Puccinia dioicae Magnus, s.lat.
Puccinia pattersoniae P. Syd. & Syd.
Tripsacum dactyloides (L.) L. (Poaceae)
Puccinia peckii (De Toni) Kellerm.
See: Puccinia dioicae Magnus, s.lat.
Puccinia peckiana Howe
See: Gymnoconia peckiana (Howe) Trotter
Puccinia persistens Plowr., s.lat.
Actaea racemosa L. (Ranunculaceae)
Anemone virginiana L. (Ranunculaceae)
Anemonella thalictroides (L.) Eames & B. Boivin (Ranunculaceae) (USDA 1960)
Aquilegia canadensis L. (Ranunculaceae)
Aquilegia sp. (Underwood 1893)
Puccinia persistens subsp. agropyri (Ellis & Everh.) J. Marková & Z. Urb.
Bromus kalmii A. Gray (=purgans auct. non L.) (Poaceae) (USDA 1960)
Clematis drummondii Torr. & A. Gray (Ranunculaceae)
C. hirsutissima var. scottii (Porter) Erickson
C. ligusticifolia Nutt.
C. virginiana L.
Elymus hystrix L. (=Hystrix patula Moench) (Poaceae) (USDA 1960)
Puccinia persistens subsp. triticina (Erikss.) Z. Urb. & J. Marková
Triticum aestivum L. (Poaceae) (Ruhl et al. 1981, 1982); (=T. vulgare Vill.) Poaceae (Jackson 1915; Pipal 1915; Anderson and Anderson 1919)
Puccinia phragmitis var. phragmitis (Schumach.) Körn.
Rumex crispus L. (Polygonaceae) (Arthur 1902)
R. obtusifolius L. (Arthur 1902)
Puccinia physostegiae Peck & Clinton
Physostegia parviflora Nutt. ex A. Gray (Lamiaceae)
P. virginiana (L.) Benth. (Arthur 1899; Jackson 1915; USDA 1960; Cooke 1967); [reported as ‘virginica’] (Snyder 1896)
Puccinia pimpinellae (F. Strauss) Link
Chaerophyllum procumbens (L.) Crantz (Apiaceae) (McCain et al. 1990)
Osmorhiza claytonii (Michx.) C.B. Clarke (Apiaceae)
O. longistylis (Torr.) DC. (Jackson 1920)
Puccinia poae-nemoralis G.H. Otth
Alopecurus aequalis Sobol. (Poaceae) (USDA 1960)
A. geniculatus L. (Jackson 1915; USDA 1960) PUR: 23412
Catabrosa aquatica (L.) P. Beauv. (Poaceae) (USDA 1960)
Poa pratensis L. (Poaceae) (McCain et al. 1990)
Poa sp.
Puccinia poarum Nielsen
Festuca sp. (Poaceae)
Hymenoxys hoopesii (A. Gray) Bierner (Asteraceae)
Koeleria macrantha (Ledeb.) Schult. (=nitida Nutt., nom. utique rej.) (Poaceae) (McCain et al. 1990)
Liatris punctata Hook. (Asteraceae)
L. scariosa (L.) Willd.
L. spheroidea Michx.
L. spicata (L.) Willd.
Poa pratensis L. (Poaceae) (Snyder 1898; Arthur 1899; Jackson 1915; Pipal 1915; Anderson and Anderson 1919)
Puccinia poculiformis (Jacq.) Wettst.
See: Puccinia graminis Pers.
Puccinia podophylli Schwein.
See: Allodus podophylli (Schwein.) Arthur
Puccinia polemonii Dietel & Holw.
Polemonium reptans L. (Polemoniaceae) (Jackson 1920; USDA 1960)
Puccinia polygoni-amphibii Pers.
Geranium maculatum L. (Geraniaceae) (Jackson 1915)
Polygonum amphibium L. (Polygonaceae); var. emersum Michx. (=Persicaria amphibia (L.) Gray & P. emersum (Michx.) Britton) (Arthur 1899; Jackson 1915); var. emersum (=Persicaria muehlenbergii (S. Watson) Small & P. muehlenbergii S. Watson) (Underwood 1893; Jackson 1915; Anderson and Anderson 1919)
P. convolvulus L. (Arthur 1899; Anderson and Anderson 1919); (=Tiniaria convolvulus (L.) Webb & Moq.) (Jackson 1915)
P. erectum L. (Snyder 1896)
P. hydropiperoides Michx. (Snyder 1898; Arthur 1899); (=Persicaria hydropiperoides (Michx.) Small) (Jackson 1915)
P. lapathifolium L.; (=Persicaria lapathifolia (L.) Gray) (Jackson 1915)
P. pensylvanicum L. (=pensylvanicum var. laevigatum Fernald) (Arthur 1899; McCain et al. 1990); (=Persicaria pensylvanica (L.) Small) (Jackson 1915)
P. punctatum Elliott (Arthur 1899); var. punctatum (=P. acre Kunth) (Underwood 1893; Anderson and Anderson 1919); (=Persicaria punctata (Elliott) Small) (Jackson 1915)
P. scandens L. (Arthur 1899; Cooke 1967); (=Tiniaria scandens (L.) Small) (Jackson 1915)
Polygonum sp.
Polygonum virginianum L. (Cooke 1967)
P. amphibium var. stipulaceum Coleman (=P. natans L.)
Puccinia polysora Underw.
Zea mays L. (Poaceae) (Ullstrup and Lavoilette 1959; Hanlin et al. 1978; Schall et al. 1983)
Puccinia prenanthis (Pers.) Fuckel
See: Puccinia orbicula Peck & Clinton
Puccinia pringsheimiana Kleb.
See: Puccinia caricina DC., s.lat.
Puccinia procera Dietel & Holway
Phacelia distans Benth. (Hydrophyllaceae)
Puccinia proserpinacae Farl.
Proserpinaca palustris L. (Haloragaceae)
Puccinia pruni-spinosae Pers., sensu Anderson and Anderson (1919)
See: Tranzschelia arthurii Tranzschel & M.A. Litv.
Puccinia pseudostriiformis M. Abbasi, Hedjar. & M. Scholler
Poa compressa L. (Poaceae)
P. nemoralis L.
P. pratensis L.
P. secunda J. Presl
Puccinia punctata Link
Galium asprellum Michx. (Rubiaceae) (Underwood 1893; Arthur 1899; Jackson 1915)
G. concinnum Torr. & A. Gray (Underwood 1893; Arthur 1899; Jackson 1915; Anderson and Anderson 1919; Cooke 1967)
Galium sp.
Puccinia punctata var. troglodytes (Lindr.) Arthur
Galium triflorum Michx. (Rubiaceae) (Underwood 1893; Arthur 1899; Jackson 1915; Anderson and Anderson 1919; Lee 1929; Emmons et al. 1960)
Puccinia punctiformis (F. Strauss) Röhl.
Cirsium arvense (L.) Scop. (Asteraceae) (McCain et al. 1990)
Puccinia purpurea Cooke
Sorghum bicolor (L.) Moench (Poaceae) (Ullstrup and Lavoilette 1959); subsp. bicolor (L.) Moench (=S. vulgare Pers.) (Ullstrup and Lavoilette 1959); subsp. drummondii (Nees ex Steud.) de Wet & Harlan (=S. sudanense (Piper) Stapf) (Ullstrup and Lavoilette 1959)
S. halepense (L.) Pers. (Ullstrup and Lavoilette 1959)
Puccinia pustulata Arthur
See: Puccinia andropogonis Schwein.
Puccinia pygmaea Erikss.
Calamagrostis × acutiflora (Schrad.) Rchb. (Poaceae)
Puccinia ranunculi Seym., Nom. illegit.
See: Tranzschelia pseudofusca M. Scholler & M. Abbasi
Puccinia recondita Roberge ex Desm. s.str.
Anchusa arvensis (L.) M. Bieb. (Boraginaceae)
A. capensis Thunb. PUR: 23972, 65619, 6580 – Greenhouse & Hort. Garden
A. officinalis L. (USDA 1960)
Anchusa sp. PUR: 65620 – Greenhouse
Hordeum jubatum L. (Poaceae) N1436
Secale cereale L. (Poaceae) (Snyder 1896; Arthur 1899; Arthur 1909; Jackson 1915; Anderson and Anderson 1919; Pipal 1915; USDA 1960)
Puccinia recondita Roberge ex Desm., sensu Cummins 1971, nom. ambig.
See: Puccinia persistens Plowr., s.lat.; P. impatientis (Schwein.) Arthur; P. bromina Erikss. s.lat.
Avena sativa L. (Poaceae) (USDA 1960) [Doubtful host]
Bromus japonicus Thunb. (Poaceae) (USDA 1960)
B. kalmii A. Gray (=purgans auct. non L.) (USDA 1960; Cooke 1967)
Notes: records on Bromus species probably refer to Puccinia bromina
Elymus canadensis L. (Poaceae)
E. hystrix L. (=Hystrix patula Moench) (Emmons et al. 1960; USDA 1960)
E. repens (L.) Gould
E. trachycaulus (Link) Gould ex Shinners
E. villosus Muhl. ex Willd.
E. virginicus L.
Glyceria grandis S. Watson (Poaceae) (USDA 1960) PUR: 25351
Note: probably refers to Puccinia persistens subsp. agropyri
Pascopyrum smithii (Rydb.) A. Löve (Poaceae)
Ranunculus cymbalaria Pursh (Ranunculaceae)
Note: Probably refers to Puccinia eatoniae
Thalictrum alpinum L. (Ranunculaceae)
T. delavayi Franch.
T. dioicum L.
T. flavum L.
T. glaucum Sieber ex Lecoy.
T. pubescens Pursh
Thalictrum sp.
T. delavayi var. delavayi Franch. (=T. dipterocarpum Franch.)
Note: records on Thalictrum spp. probably refer to Puccinia cockerelliana or P. persistens.
Puccinia rhamni (Pers. ex J.F. Gmel.) Wettst.
See: Puccinia coronata Corda, s.lat. and P. coronata var. avenae f. sp. avenae Urban & Marková
Puccinia rubella (Pers. ex J.F. Gmel.) Arthur
See: Puccinia phragmitis var. phragmitis (Schumach.) Tul.
Puccinia rubigo-vera (DC.) G. Winter, nom. ambig.
This is an ambiguous name used by Underwood (1893) and Snyder (1897)
See: Puccinia impatientis (Schwein.) Arthur, P. recondita Roberge ex Desm. s.str. and P. schedonnardi Kellerm. & Swingle
Avena sativa L. (Poaceae) (Underwood 1893) [Doubtful host]
Puccinia rubigo-vera var. agropyrina (Erikss.) Arthur
See: Puccinia persistens subsp. agropyri (Ellis & Everh.) J. Marková & Z. Urb.
Puccinia rubigo-vera var. apocrypta (Ellis & Tracy) Arthur
See: Puccinia apocrypta Ellis & Tracy
Puccinia rubigo-vera f. sp. arthuri Mains
See: Puccinia persistens subsp. agropyri (Ellis & Everh.) J. Marková & Z. Urb.
Puccinia rubigo-vera f. sp. impatiensis Mains
See: Puccinia impatientis (Schwein.) Arthur
Puccinia rubigo-vera f. sp. indianensis Mains
See: Puccinia impatientis (Schwein.) Arthur
Puccinia rubigo-vera f. secalis (Erikss. & Henn.) Mains
See: Puccinia recondita Roberge ex Desm. s.str.
Puccinia ruelliae (Berk. & Broome) Lagerh.
See: Puccinia lateripes Berk. & Ravenel
Puccinia sambuci (Schwein.) Arthur
Carex aureolensis Steud. (=C. frankii Kunth) (Cyperaceae) (Jackson 1915; Anderson and Anderson 1919; Schoknecht 1981) PUR: 30297
C. lupuliformis Sartwell ex Dewey (Jackson 1920)
C. lupulina Muhl. ex Willd. (Jackson 1915)
C. lurida Wahlenb. (Arthur 1909; Jackson 1915)
Carex sp. (Underwood 1893; Snyder 1896; Arthur 1899)
C. trichocarpa Muhl. ex Willd. (Jackson 1915; Anderson and Anderson 1919)
Sambucus racemosa L. (Caprifoliaceae)
S. nigra subsp. canadensis (L.) R. Bolli (=S. canadensis L.) (Underwood 1893; Jackson 1915; Anderson and Anderson 1919; Van Hook 1928)
S. racemosa var. racemosa L. (=S. pubens Michx.) (Jackson 1920; USDA 1960)
Puccinia saniculae Grev.
See: Puccinia marylandica Lindr.
Puccinia schedonnardi Kellerm. & Swingle
Callirhoe involucrata (Torr. & A. Gray) A. Gray (Malvaceae)
Melica mutica Walter (Poaceae) (Jackson 1920)
Melica sp. L. (USDA 1960)
Muhlenbergia mexicana (L.) Trin. (Poaceae) (Jackson 1915)
M. racemosa (Michx.) Britton, Sterns & Poggenb. (Jackson 1920)
M. schreberi J.F. Gmel. (Jackson 1915; Emmons et al. 1960; Cummins and Greene 1961); (as M. diffusa) (Underwood 1893) PUR: 21416
M. sobolifera (Muhl. ex Willd.) Trin. (Jackson 1920)
M. sylvatica (Torr.) Torr. ex A. Gray (=M. umbrosa Scribn.) (Snyder 1896; Jackson 1915)
M. tenuiflora (Willd.) Britton, Sterns & Poggenb. (Jackson 1915; Anderson and Anderson 1919)
Napaea dioica L. (Malvaceae) (Jackson 1915)
Sphaeralcea angustifolia (Cav.) G. Don (Malvaceae)
S. coccinea (Nutt.) Rydb.
S. incana Torr. ex A. Gray
Triodia seslerioides Benth. ex Vasey (Poaceae) (Snyder 1896)
Puccinia schroeteriana Plowr. & Magnus
See: Puccinia dioicae Magnus, s.lat.
Puccinia sessilis J. Schröt.
Iris versicolor L. (Iridaceae) (Jackson 1920; USDA 1960)
Phalaris arundinacea L. (Poaceae) (Jackson 1915)
Puccinia seymeriae Burrill
Dasistoma macrophylla (Nutt.) Raf. (=Afzelia macrophylla (Nutt.) Kuntze) (Orobanchaceae) (Jackson 1915)
Puccinia seymouriana Arthur
Apocynum cannabinum L. (Apocynaceae) (Jackson 1920; USDA 1960)
Asclepias syriaca L. (Apocynaceae)
Cephalanthus occidentalis L. (Rubiaceae) (Jackson 1915)
Spartina pectinata Bosc ex Link (Poaceae); (=S. michauxiana Hitchc.) (Jackson 1915)
S. cynosuroides (L.) Roth (Arthur 1909)
Puccinia silphii Schwein.
Silphium integrifolium Michx. (Asteraceae) (Jackson 1915)
S. perfoliatum L. (Jackson 1915; Anderson and Anderson 1919; Van Hook 1923)
Silphium sp. (Underwood 1893; Arthur 1899; Jackson 1915)
S. terebinthinaceum Jacq.
Puccinia smilacis Schwein.
Apocynum cannabinum L. (Apocynaceae)
Smilax bona-nox L. (Smilacaceae) (Jackson 1920)
S. glauca Walter (Jackson 1915)
S. rotundifolia L.
S. tamnoides L. (=hispida Muhl. ex Torr.) (Emmons et al. 1960)
Puccinia sorghi Schwein.
Oxalis corniculata L. (Oxalidaceae) (Anderson and Anderson 1919; USDA 1960)
O. stricta L. (USDA 1960); (=Xanthoxalis cymosa (Small) Small) (Jackson 1915)
Zea mays L. (Poaceae) (Underwood 1893; Snyder 1898; Arthur 1899; Jackson 1915; Pipal 1915; Anderson and Anderson 1919; Ullstrup and Lavoilette 1959; Hanlin et al. 1978; Ruhl et al. 1980, 1981, 1982)
Zea mexicana (Schrad.) Kuntze (Poaceae)
Host not reported (Van Hook 1910)
Puccinia sparganioidis Ellis & Barthol.
Fraxinus americana L. (Oleaceae) (McCain and Hennen 1981)
F. pennsylvanica Marshall (=F. lanceolata Borkh.) (Arthur 1902)
F. profunda (Bush) Bush (Jackson 1920)
Fraxinus sp. (Ruhl et al. 1981)
F. viridis F. Michx. (Van Hook 1927)
Spartina pectinata Bosc ex Link (Poaceae); (=S. michauxiana Hitchc.) (Jackson 1915)
Puccinia splendens Vize
Hymenoclea monogyra Torr. & A. Gray (Asteraceae)
Puccinia sporoboli Arthur
Allium cernuum Roth (Amaryllidaceae)
A. drummondii Regel
Lilium philadelphicum L. (Liliaceae)
Sporobolus cryptandrus (Torr.) A. Gray (Poaceae) (Snyder 1896)
Puccinia sporoboli var. robusta Cummins & H.C. Greene
Calamovilfa longifolia (Hook.) Scribn. (Poaceae) (Cummins and Greene 1961)
Maianthemum racemosum (L.) Link (Asparagaceae)
M. stellatum (L.) Link (=Smilacina stellata (L.) Desf.) (Cummins and Greene 1961)
Yucca glauca Nutt. (Agavaceae)
Puccinia stipae Arthur
Grindelia squarrosa (Pursh) Dunal (Asteraceae)
Gutierrezia sarothrae (Pursh) Britton & Rusby (Asteraceae)
Hesperostipa spartea (Trin.) Barkworth (Poaceae)
Senecio spartioides Torr. & Gray (Asteraceae)
Symphyotrichum ericoides (L.) G.L. Nesom (Asteraceae)
S. novae-angliae (L.) G.L. Nesom
S. porteri (A. Gray) G.L. Nesom
Puccinia striiformis Westend.
Triticum aestivum L. (Poaceae) (McCain and Hennen 1981; Ruhl et al. 1983; Abbasi et al. 2005)
Also see: ‘Puccinia pseudostriiformis M. Abbasi, Hedjar. & M. Scholler’ for reports on Poa spp.
Puccinia subnitens Dietel
Abronia fragrans Nutt. ex Hook. (Nyctaginaceae)
Atriplex prostrata Bouchér ex DC. (=A. hastata L.) (Chenopodiaceae)
Capsella bursa-pastoris (L.) Medik. (Brassicaceae)
Chenopodium album L. (Chenopodiaceae) (USDA 1960)
Cleome hassleriana Chod. (=C. spinosa auct. non Jacq.) (Capparidaceae) (USDA 1960)
C. serrulata Pursh (USDA 1960)
Descurainia pinnata var. brachycarpa (Richardson) Detling (Brassicaceae)
Erysimum capitatum (Dougl. ex Hook.) Greene (Brassicaceae)
Lepidium densiflorum Schrad. (Brassicaceae)
Lepidium sp. (USDA 1960)
L. virginicum L.
Polygonum aviculare L. (Polygonaceae)
Sarcobatus vermiculatus (Hook.) Torr. (Chenopodiaceae)
Puccinia substerilis Ellis & Everh.
Nassella viridula (Trin.) Barkworth (Poaceae)
Puccinia substriata Ellis & Barthol.
Paspalum setaceum Michx. (=P. setaceum var. muhlenbergia (Nash) D.J. Banks) (Poaceae)
Solanum carolinense L. (Solanaceae)
Puccinia substriata var. imposita (Arthur) Ramachar & Cummins
Solanum carolinense L. (Solanaceae)
S. melongena L.
Puccinia tanaceti DC., sensu Underwood (1893) [also followed by Snyder (1897)]
See: Puccinia helianthi Schwein.
Puccinia tanaceti var. tanaceti DC.
Chrysanthemum × morifolium Ramat. (pro sp.) (Asteraceae)
C. morifolium (Ramat.) Hemsl. var. sinense (Sabine) Makino (=C. sinense Sabine) (Jackson 1915)
Chrysanthemum sp. (Pipal 1915)
Puccinia tanaceti var. dracunculina (Fahrend.) Cummins
Artemisia (=Dracunculoides L.) dracunculus L. (Asteraceae)
Puccinia taraxaci Plowr.
See: Puccinia hieracii (Röhl.) H. Mart.
Puccinia tenuis (Schwein.) Burrill
Ageratina altissima var. altissima (L.) R.M. King & H. Rob. (=Eupatorium rugosum Houtt.) (Asteraceae) (Emmons et al. 1960); var. roanensis (Small) Clewell & Woot. (=E. urticifolium Reichard) (Jackson 1915)
Ageratum altissimum (=E. ageratoides L.f.) (Asteraceae) (Underwood 1893; Snyder 1896; Arthur 1899)
Puccinia thalictri Chevall.
See: Tranzschelia thalictri (Chev.) Dietel
Puccinia thelypodii Cummins
Guillenia lasiophylla (Hook. & Arn.) Greene (=Thelypodium lasiophyllum (Hook. & Arn.) Greene) (Brassicaceae)
Puccinia thompsonii Hume
See: Puccinia sambuci (Schwein.) Arthur
Puccinia tomipara Trel.
Bromus ciliatus L. (Poaceae) (Anderson and Anderson 1919)
Thalictrum thalictroides (L.) Eames & B. Boivin (=Anemonella thalictroides (L.) Spach) (Ranunculaceae) (Anderson and Anderson 1919)
Puccinia triodiae Ellis & Barthol.
See: Puccinia schedonnardi Kellerm. & Swingle
Puccinia tripsaci Dietel & Holw.
Ceanothus americanus L. (Rhamnaceae)
Tripsacum sp. (Poaceae) (USDA 1960)
Puccinia tritici auct. non Oerst.
See: Puccinia persistens subsp. triticina (Erikss.) Z. Urb. & J. Marková
Puccinia triticina Erikss.
See: Puccinia persistens subsp. triticina (Erikss.) Z. Urb. & J. Marková
Puccinia troglodytes Lindr. [as ‘triglodytes’ by Lee (1929)]
See: Puccinia punctata var. troglodytes (Lindr.) Arthur
Puccinia tumidipes Peck
Lycium halimifolium Mill. (= L. barbarum L.) (Solanaceae) (Otálora and Berndt 2018)
Puccinia urticae Lagerh.
See: Puccinia urticata F. Kern.
Puccinia urticata F. Kern
Carex bullata Schkuhr ex Willd. (Cyperaceae) (Arthur 1899)
C. emoryi Dewey (Jackson 1920)
C. frankii Kunth (Arthur 1899)
C. lacustris Willd. (Jackson 1915)
C. lurida Wahlenb. (Arthur 1899)
C. pensylvanica Lam. (Arthur 1899)
C. siccata Dewey (=C. foenea Willd.) (Arthur 1899)
C. stipata Muhl. ex Willd.
C. straminea Willd. ex Schkuhr (Arthur 1899)
C. stricta Lam. (Jackson 1915) PUR: 66442
C. trichocarpa Muhl. ex Willd. (Jackson 1920)
C. virescens Muhl. ex Willd. (Arthur 1899)
Dulichium arundinaceum (L.) Britton (Arthur 1899)
Urtica dioica L. ssp. gracilis (Aiton) Seland. (=U. gracilis Aiton) (Urticaceae) (Arthur 1899; Jackson 1915)
Puccinia verbenicola Arthur
See: Puccinia vilfae Arthur & Holw.
Puccinia verbesinae Schwein.
Verbesina occidentalis (L.) Walter (Asteraceae) (USDA 1960)
Puccinia vernoniae Schwein.
See: Puccinia longipes var. brevipes (Dietel) Z. Urb.
Puccinia vernoniae var. caulicola Ellis & Everh.
See: Puccinia longipes var. longipes Lagerh.
Puccinia veronicarum DC., s.lat.
Veronicastrum virginicum (L.) Farw. (Plantaginaceae) PUR: N6188
Puccinia verrucosa Link
See: Puccinia glechomatis DC.
Puccinia vexans Farl.
Bouteloua curtipendula (Michx.) Torr. (Poaceae); (=Atheropogon curtipendulus (Michx.) Fourn.) (Jackson 1915; Jackson 1920)
Fouquieria splendens Engelm. (Fouquieriaceae) PUR: N5514 - Greenhouse
Puccinia vilfae Arthur & Holw.
Sporobolus compositus (Poir.) Merr. (Poaceae); (=S. asper (P. Beauv.) Kunth) (Arthur 1899; Jackson 1915; Cummins and Greene 1961)
Verbena hastata L. (Verbenaceae) (Jackson 1920)
V. stricta Vent. (Snyder 1896; Jackson 1915)
V. urticifolia L.
Verbena × illicita Moldenke [stricta × urticifolia] (McCain et al. 1990)
Puccinia violae (Schumach.) DC.
Viola blanda Willd. (Violaceae) (Anderson and Anderson 1919); var. palustriformis A. Gray (=V. incognita var. forbesii Brainerd)
V. cucullata Aiton (=V. obliqua Hill) (Arthur 1899; Arthur and Holway 1901; Anderson and Anderson 1919)
V. eriocarpa Schwein. (Jackson 1915)
V. missouriensis Greene (Jackson 1920)
V. palmata var. cucullata A. Gray (Underwood 1893)
V. pubescens Aiton (Arthur and Holway 1901; Jackson 1915; Anderson and Anderson 1919)
V. sororia Willd.; (=V. papilionacea Pursh p.p.) (Jackson 1915; Anderson and Anderson 1919)
Viola sp. (Underwood 1893; Cooke 1967; Ruhl et al. 1981; Schoknecht 1981)
V. striata Aiton (Underwood 1893; Arthur 1899; Arthur and Holway 1901; Jackson 1915; Anderson and Anderson 1919)
Host not reported (Van Hook 1910) [reported as P. violaceae]
Puccinia virgata Ellis & Everh.
Sorghum bicolor subsp. bicolor (L.) Moench (=S. vulgare Pers.) (Poaceae) PUR: N7456
Puccinia vulpinoideae Dietel & Holw. [as’vulpinoidis’]
See: Puccinia dioicae Magnus, s.lat.
Puccinia windsoriae Burrill
See: Puccinia schedonnardi Kellerm. & Swingle
Puccinia windsoriae Schwein.
Ptelea perfoliatum (Rutaceae) (Snyder 1896) [Doubtful host]
P. trifoliata L. (Underwood 1893; Jackson 1915; Anderson and Anderson 1919)
Sieglingia seslerioides Scribn. (Poaceae) (Arthur 1899)
Tridens flavus (L.) Hitchc. (=Triodia flava (L.) Hitchc.) (Poaceae) (Anderson and Anderson 1919; Cooke 1967, 1975); [as ‘flava’] (Jackson 1915)
Puccinia xanthii Schwein.
Ambrosia sp. (Asteraceae)
A. trifida L. (Asteraceae) (Underwood 1893; Snyder 1896; Arthur 1899; Jackson 1915)
Xanthium sp. (Asteraceae)
X. spinosum L. (Jackson 1915; USDA 1960; McCain et al. 1990)
X. strumarium L. (Underwood 1893; Arthur 1899); var. canadense (Mill.) Torr. & A. Gray (McCain et al. 1990); var. canadense (=X. commune Britton [as ‘communis’]) (Jackson 1915); var. canadense (=X. canadense Mill.) (Underwood 1893; Snyder 1896; Arthur 1899; Anderson and Anderson 1919); var. canadense (=X. pensylvanicum Wallr.) (Jackson 1915); var. canadense (Mill.) Torr. & A. Gray (=X. italicum Moretti); var. glabratum (DC.) Cronquist (McCain et al. 1990); var. glabratum (DC.) Cronquist (=X. americanum Walter) (Jackson 1915)
Host not reported (Van Hook 1910)
Puccinia xanthii f. sp. ambrosiae-trifidae S.W.T. Batra [as ‘ ambrosia-trifidae ’]
Ambrosia trifida L. (Asteraceae) (Batra 1981)
Puccinia sp.
Poa pratensis L. (Poaceae) (Ruhl et al. 1981)
Pucciniastrum agrimoniae (Dietel) Tranzschel
Agrimonia eupatoria L. (Rosaceae) (Underwood 1893; Snyder 1896)
A. gryposepala Wallr. (Anderson and Anderson 1919; Emmons et al. 1960)
A. hirsuta Bickn. (Arthur 1899; Jackson 1915)
A. microcarpa Wallr. (Hiratsuka 1958)
A. parviflora Aiton (Underwood 1893; Arthur 1899; Jackson 1915; Hiratsuka 1958)
A. pubescens Wallr. (Cooke 1975); (=A. mollis (Torr. & A. Gray) Britton) (Jackson 1915)
A. rostellata Wallr. (Jackson 1920; Hiratsuka 1958)
Agrimonia sp.
Host not reported (Cooke 1967)
Pucciniastrum americanum (Farl.) Arthur
See: Thekopsora americana (Farl.) Aime & McTaggart
Pucciniastrum goeppertianum (Kuehn) Kleb.
See: Calyptospora columnaris (Alb. & Schwein.) J.G. Kühn
Pucciniastrum hydrangeae (Magnus) Arthur
Hydrangea arborescens L. (Hydrangeaceae) (Underwood 1893;; Snyder 1896; Arthur 1899; Jackson 1915; Anderson and Anderson 1919; Cooke 1967, 1975; Schoknecht 1981)
Hydrangea sp.
H. cinerea Small (=H. arborescens subsp. discolor (Ser. ex DC.) McClint.) (McCain et al. 1990)
H. paniculata Siebold (USDA 1960)
Tsuga canadensis (L.) Carriere (Pinaceae) (USDA 1960)
Pucciniastrum myrtilli Arthur
See: Naohidemyces vaccinii (Jørst.) S. Sato, Katsuya & Y. Hirats. ex Vanderweyen & Fraiture
Pucciniastrum vaccinii (G. Winter) Jørst. 1952
See: Naohidemyces vaccinii (Jørst.) S. Sato, Katsuya & Y. Hirats. ex Vanderweyen & Fraiture
Pucciniosira tuberculata (Ellis & Kellerm.) Buriticá & J.F. Hennen
Callirhoe involucrata (Torr. & A. Gray) A. Gray (Malvaceae) (USDA 1960)
Ravenelia epiphylla (Schwein) Dietel
Tephrosia virginiana (L.) Pers. (=T. virginiana var. holosericea (Nutt.) Torr. & A. Gray) (Fabaceae); (=Cracca virginiana L.) (Jackson 1915)
Roestelia lacerata (Sowerby) Fr.
See: Gymnosporangium globosum Farl.
Roestelia pyrate Thaxt.
See: Gymnosporangium juniperi-virginianae Schwein.
Teleutospora rudbeckiae (Arthur & Holw.) Arthur & Bisby
See: Uromyces rudbeckiae Arthur & Holw.
Thekopsora americana (Farl.) Aime & McTaggart
Rubus idaeus L. (Rosaceae)
Tranzschelia arthurii Tranzschel & M.A. Litv.
Hepatica nobilis var. acuta (Pursh) Steyerm. (Ranunculaceae); var. acuta (=H. acutiloba DC.) (Underwood 1893; Jackson 1915; Anderson and Anderson 1919; Van Hook 1934); var. obtusa (Pursh) Steyerm. (=H. triloba Chaix) (Anderson and Anderson 1919)
Prunus serotina Ehrh. (Rosaceae) (Jackson 1920; Lopez-Franco and Hennen 1990); var. serotina Ehrh. (Lopez-Franco and Hennen 1990)
P. virginiana L. (Rosaceae) (Lopez-Franco and Hennen 1990)
Tranzschelia fusca (Pers.) Dietel
See: Tranzschelia pseudofusca M. Scholler & M. Abbasi
Tranzschelia pruni-spinosae (Pers.) Dietel
Anemone quinquefolia L. (Ranunculaceae) (Jackson 1920; Lopez-Franco and Hennen 1990)
Thalictrum dioicum L. (Ranunculaceae) (USDA 1960)
Tranzschelia pruni-spinosae var. americana López-Franco & Hennen
Prunus americana Marshall (Rosaceae)
Tranzschelia pseudofusca M. Scholler & M. Abbasi
Anemone quinquefolia L. (Ranunculaceae) (Underwood 1893; Arthur 1899; Jackson 1915; Jackson 1920; Lopez-Franco and Hennen 1990)
Ranunculus repens L. (Ranunculaceae) (Underwood 1893)
Tranzschelia punctata Arthur
Note: This name as applied could refer to Tranzschelia arthurii Tranzschel & M.A. Litv., T. pruni-spinosae (Pers.) Dietel or Aecidium dakotense Griffiths
Tranzschelia thalictri (Chev.) Dietel
Thalictrum dioicum L. (Ranunculaceae) (Underwood 1893; Arthur 1899; Jackson 1915; Anderson and Anderson 1919; Lopez-Franco and Hennen 1990)
Triphragmium ulmariae (DC.) Link
Filipendula rubra (Hill.) B.L. Rob. (=Spiraea rubra (Hill) Britton) (Rosaceae) (Jackson 1915; Jackson 1920; Arthur 1934; Monoson 1975)
F. ulmaria (L.) Maxim. (USDA 1960)
Ulmaria sp. (Rosaceae) (Monoson 1975)
Uredinopsis americana Syd. & P. Syd.
Onoclea sensibilis L. (Onocleaceae) (Jackson 1917; Hiratsuka 1958)
Uredinopsis arthurii Faull
Woodwardia virginica (L.) Sm. (Blechnaceae) (USDA 1960)
Uredinopsis atkinsonii Magnus
Cystopteris bulbifera (L.) Bernh. (=Filix bulbifera (L.) Underw.) (Cystopteridaceae) (Jackson 1920)
Thelypteris palustris var. pubescens (G. Lawson) Fernald (=Dryopteris thelypteris var. pubescens (G. Lawson) A.R. Prince ex Weath.) (Thelypteridaceae) (Hiratsuka 1958); (=D. thelypteris (L.) A. Gray) (Jackson 1917)
Host not reported (Van Hook 1934)
Uredinopsis ceratophora Faull
Cystopteris bulbifera (L.) Bernh. (=Filix bulbifera (L.) Underw.) (Cystopteridaceae) (USDA 1960)
Uredinopsis mirabilis (Peck) Magnus
See: Uredinopsis americana Syd. & P. Syd.
Uredinopsis osmundae Magnus
Abies balsamea (L.) Mill. (Pinaceae)
Osmunda claytoniana L. (Osmundaceae)
Uredinopsis pteridis Dietel & Holw.
Pteridium aquilinum (Dennstaedtiaceae) [as Pteris aquilina L. (Pteridaceae)] (Lee 1929)
Uredinopsis struthiopteridis F.C.M. Störmer
Athyrium filix-foemina var. cyclosorum Rupr. (Dryopteridaceae)
Cystopteris bulbifera (L.) Bernh. (=Filix bulbifera (L.) Underw.) (Cystopteridaceae)
Thelypteris palustris Schott (Thelypteridaceae)
Woodwardia virginica (L.) Sm. (Blechnaceae)
W. virginica (=Anchistea virginica (L.) C. Presl) (Jackson 1920)
Uredo bigelowii (Thüm.) Arthur
See: Melampsora paradoxa Dietel & Holw.
Uredo hydrangeae Berk. & M.A. Curtis
See: Pucciniastrum hydrangeae (Magnus) Arthur
Uredo polypodii (Pers.) DC.
See: Hyalopsora polypodii (Pers.) Magnus
Uredo similis Ellis
See: Puccinia globosipes Peck
Uromyces acuminatus Arthur
Collomia linearis Nutt. (Polemoniaceae)
Lysimachia ciliata L. (Primulaceae)
Maianthemum stellatum (L.) Link (=Smilacina stellata (L.) Desf.) (Asparagaceae) (McCain et al. 1990)
Phlox divaricata L. (Polemoniaceae)
P. maculata L. (Jackson 1920)
Polygonatum biflorum (Walter) Elliott (Asparagaceae) (Jackson 1917)
Polemonium reptans L. (Polemoniaceae) (Jackson 1915; USDA 1960)
Smilacina sp. (Liliaceae)
Spartina pectinata Bosc ex Link (Poaceae); (=S. michauxiana Hitchc.) (Jackson 1915)
S. pectinata Link
Spartina sp.
Uromyces acuminatus var. polemonii (Arthur) Davis
See: Uromyces acuminatus Arthur
Uromyces americanus Speg.
Schoenoplectus tabernaemontani (C.C. Gmelin) Palla (=Scirpus validus Vahl) (Cyperaceae) (USDA 1960)
Uromyces amphidymus Syd. & P. Syd.
Glyceria septentrionalis Hitchc. (Poaceae) (USDA 1960); (=Panicularia septentrionalis (Hitchc.) E.P. Bicknell) (Jackson 1915)
Uromyces andropogonis Tracy
Andropogon gyrans Ashe var. gyrans (=elliottii Chapm.) (Poaceae) (Hennen 1965)
A. virginicus L. (Hennen 1965)
Viola cucullata Aiton (=V. obliqua Hill) (Violaceae) (Arthur and Holway 1901)
V. affinis Le Conte
Uromyces appendiculatus var. appendiculatus (Pers.) Unger
Phaseolus diversifolius Pers. (Fabaceae) (Snyder 1896)
Phaseolus sp. (Underwood 1893)
P. vulgaris L. (Jackson 1915; Pipal 1915; Anderson and Anderson 1919; Ruhl et al. 1981, 1982)
Strophostyles angulosa Elliott (Fabaceae) (Underwood 1893)
S. helvola (L.) Elliott (Arthur 1899; Jackson 1915; Anderson and Anderson 1919)
S. leiosperma (Torr. & A. Gray) Piper (=S. pauciflora (Benth.) S. Watson) (Jackson 1915)
S. umbellata (Muhl. ex Willd.) Britton (Jackson 1915)
Vigna angularis (Willd.) Ohwi & Ohashi (=P. angularis (Willd.) W. Wight) (Fabaceae) (USDA 1960) [Doubtful host]
Host not reported (Van Hook 1910)
Uromyces ari-triphylli (Schwein.) Seeler
Arisaema dracontium (L.) Schott (Araceae) (Underwood 1893; Snyder 1896; Arthur 1899; Jackson 1915; Anderson and Anderson 1919)
A. triphyllum (L.) Schott (Underwood 1893; Snyder 1896, 1898; Arthur 1899; Jackson 1915; Anderson and Anderson 1919; Cooke 1967, 1975)
Peltandra virginica (L.) Schott (Araceae) (Jackson 1915; USDA 1960)
Host not reported (Van Hook 1910)
Uromyces asclepiadis Cooke
Asclepias cornuti Decne. (Apocynaceae) (Underwood 1893; Snyder 1896, 1898)
A. incarnata L. (Jackson 1915, 51; Snyder 1896; Arthur 1899; Anderson and Anderson 1919)
A. purpurascens L. (Underwood 1893; Arthur 1899; Jackson 1915; Anderson and Anderson 1919)
A. syriaca L. (Arthur 1899; Jackson 1915; Anderson and Anderson 1919)
A. tuberosa L.
Matelea obliqua (Jacq.) Woodson (Asclepiadaceae); (=Vincetoxicum shortii (A. Gray) Britton) (Jackson 1915)
Host not reported (Van Hook 1910)
Uromyces caladii Farl.
See: Uromyces ari-triphylli (Schwein.) Seeler
Uromyces caryophyllinus (Schrank) J. Schröt.
See: Uromyces dianthi (Pers.) Niessl
Uromyces coloradensis Ellis & Everh.
Vicia americana Muhl. ex Willd. (Fabaceae) (Jackson 1920)
Uromyces dactylidis G.H. Otth
Poa pratensis L. (Poaceae) (Jackson 1915; USDA 1960)
Uromyces dianthi (Pers.) Niessl
Dianthus caryophyllus L. (Caryophyllaceae) (Arthur 1899; Jackson 1915; Pipal 1915; Anderson and Anderson 1919)
Dianthus spp. (Underwood 1893; Van Hook 1912)
Uromyces dictyospermae Ellis & Everh. ex Tranzschel
Euphorbia spathulata Lam. (Euphorbiaceae)
Uromyces elegans (Berk. & M.A. Curtis) Lagerh.
Trifolium carolinianum Michx. (Fabaceae)
Uromyces eragrostidis Tracy
Echeandia flavescens (Schult. & Schult. f.) Cruden (=Anthericum torreyi Baker) (Asparagaceae)
Uromyces euphorbiae Cooke & Peck
See: Uromyces proëminens (DC.) Lév.
Uromyces fabae (Pers.) de Bary
See: Uromyces viciae-fabae (Pers.) J. Schröt.
Uromyces fallens (Desm.) Barthol.
See: Uromyces trifolii-repentis var. fallens (Desm.) Arthur
Uromyces gaurinus (Peck) Snyder [as ‘gaurina’]
See: Uromyces plumbarius Peck
Uromyces geranii (DC.) Lév.
Geranium maculatum L. (Geraniaceae) (Underwood 1893; Snyder 1896, 1898)
Uromyces graminicola Burrill
See: Puccinia graminicola (Burrill) Demers & Castl.
Uromyces halstedii De Toni
Leersia oryzoides (L.) Sw. (Poaceae) (USDA 1960)
Trillium sp. (Melanthiaceae) (Snyder 1896; Jackson 1917)
Uromyces hedysari-paniculati (Schwein.) Farl.
Desmodium canadense (L.) DC. (=Meibomia canadensis (L.) Kuntze) (Fabaceae) (Snyder 1896; Arthur 1899)
D. canescens (L.) DC. (Underwood 1893; Anderson and Anderson 1919); (=Meibomia canescens (L.) Kuntze) (Arthur 1899; Jackson 1915)
D. ciliare (Muhl. ex Willd.) DC. (McCain et al. 1990)
D. cuspidatum (Muhl. ex Willd.) DC. ex D. Don var. cuspidatum (=M. bracteosa (Michx.) Kuntze) (Jackson 1915); var. longifolium (Torr. & A. Gray) B.G. Schub. (McCain et al. 1990)
D. glabellum (Michx.) DC. (=D. dillenii Darl.) (Anderson and Anderson 1919)
D. laevigatum (Nutt.) DC. (=M. laevigata (Nutt.) Kuntze) (Underwood 1893; Arthur 1899; Jackson 1915; Anderson and Anderson 1919)
D. paniculatum (L.) DC. (Underwood 1893; Anderson and Anderson 1919); var. paniculatum (=M. paniculata (L.) Kuntze) (Arthur 1899; Jackson 1915)
D. perplexum B.G. Schub.; (=M. dillenii (Darl.) Kuntze) (Snyder 1896; Arthur 1899; Jackson 1915); (=D. dillenii Darl.) (Underwood 1893)
D. rotundifolium DC. (Emmons et al. 1960); (=M. michauxii Vail) (Jackson 1920)
D. sessilifolium (Torr.) Torr. & A. Gray (McCain et al. 1990)
Desmodium sp. (Underwood 1893; Cooke 1967, 1975)
D. viridiflorum (L.) DC. (Underwood 1893)
D. viridiflorum (L.) DC. (=M. viridiflora (L.) Kuntze) (Arthur 1899; Jackson 1915)
Uromyces hordei Tracy
See: Uromyces hordeinus (Arthur) Barthol.
Uromyces hordeinus (Arthur) Barthol.
Hordeum pusillum Nutt. (Poaceae) (Jackson 1920)
Nothoscordum bivalve (L.) Britton (Amaryllidaceae)
Uromyces houstoniatus (Schwein.) J. Sheld.
Houstonia caerulea L. (Rubiaceae) (Jackson 1917)
H. longifolia Gaertn.
Sisyrinchium angustifolium Mill. (=S. gramineum Curtis) (Iridaceae)
Uromyces howei Peck
See: Uromyces asclepiadis Cooke
Uromyces hyperici M.A. Curtis
See: Uromyces triquetrus Cooke
Uromyces hyperici-frondosi (Schwein.) Arthur
See: Uromyces triquetrus Cooke
Uromyces junci (Desm.) Tul. & C. Tul.
Ambrosia artemisiifolia Bess. (Asteraceae)
Ambrosia psilostachya DC. (Asteraceae)
Cirsium flodmanii Arthur (Asteraceae)
Juncus tenuis Willd. (Juncaceae) (Snyder 1896, 1898; Arthur 1899)
Uromyces junci - effusi P. Syd. & Syd.
Juncus effusus L. (Juncaceae) (McCain et al. 1990)
Uromyces lespedezae Peck
See: Uromyces lespedezae-procumbentis (Schwein.) M.A. Curtis
Uromyces lespedezae-procumbentis (Schwein.) M.A. Curtis
Lespedeza capitata Michx. (Fabaceae) (Jackson 1915)
L. frutescens (L.) Hornem. (Arthur 1899; Jackson 1915; Anderson and Anderson 1919); (=L. intermedia (S. Watson) Britton, nom. inq.) (McCain et al. 1990)
L. hirta (L.) Hornem. (Jackson 1915)
L. procumbens Michx. (Underwood 1893; Arthur 1899; Jackson 1915; Anderson and Anderson 1919)
L. repens (L.) W.P.C. Barton (Snyder 1896; Arthur 1899; Jackson 1915; McCain et al. 1990)
L. reticulata Pers. (Underwood 1893)
Lespedeza sp. (Underwood 1893; Cooke 1975, 1967)
L. stuevei Nutt. (Jackson 1915)
L. violacea (L.) Pers. (Coulter 1876; McCain et al. 1990)
L. virginica (L.) Britton (Jackson 1915); f. deamii M. Hopkins (McCain et al. 1990)
Lespedeza × brittonii E.P. Bicknell (pro sp.) [procumbens × virginica] (McCain et al. 1990)
Lespedeza × nuttallii Darl. (pro sp.) [hirta × violacea] (McCain et al. 1990)
Uromyces lineolatus (Desm.) J. Schröt.
See: Uromyces lineolatus subsp. nearcticus Savile
Uromyces lineolatus subsp. nearcticus Savile
Cicuta maculata L. (Apiaceae) (Jackson 1915)
Sium suave Walter (=Sium cicutifolium Schrank) (Apiaceae)
Schoenoplectus americanus (Pers.) Volkart ex Schinz & R. Keller (=Scirpus americanus Pers.) (Cyperaceae) (Jackson 1915; Anderson and Anderson 1919)
S. fluviatilis (Torr.) M.T. Strong (=Bolboschoenus fluviatilis (Torr.) Soják); (=Scirpus fluviatilis (Torr.) A. Gray) (Jackson 1920; Savile 1972)
S. tabernaemontani (C.C. Gmel.) Palla (=Scirpus validus Vahl) (Jackson 1915; Cummins 1935)
Uromyces magnatus Arthur
See: Uromyces acuminatus Arthur
Uromyces medicaginis Pass.
See: Uromyces striatus J. Schröt.
Uromyces minutus Dietel
Carex lanuginosa Michx. (Cyperaceae) (Jackson 1915)
C. virescens Muhl. ex Willd. (Jackson 1915)
Uromyces orobi (Pers.) G. Winter
See: Uromyces viciae-fabae (Pers.) J. Schröt.
Uromyces peckianus Farl.
Atriplex prostrata Bouchér ex DC. (=Atriplex hastata L.) (Chenopodiaceae)
Uromyces perigynius Halst.
Carex albicans var. albicans Willd. ex Spreng. (=C. varia Muhl. ex Wild) (Cyperaceae) (Jackson 1915) PUR: 12781; (=C. artitecta Mack.)
C. grayi Carey; (=C. asa-grayi L.H. Bailey) (Jackson 1920)
C. intumescens Rudge
C. pubescens Poir. (Underwood 1893; Arthur 1899)
Carex sp.
C. tribuloides Wahlenb. (Jackson 1915)
Gaura biennis L. (Onagraceae)
Oenothera biennis L. (Onagraceae)
Rudbeckia laciniata L. (Asteraceae) (Jackson 1920)
Solidago caesia L. (Asteraceae)
S. canadensis L. PUR: 12708
S. flexicaulis L.
S. rugosa Mill.
S. serotina Aiton
Symphyotrichum ericoides var. ericoides (L.) G.L. Nesom (=Aster ericoides L.) (Asteraceae)
Symphyotrichum foliaceum var. canbyi (A. Gray) G.L. Nesom (=A. tweedyi Rydb.)
Symphyotrichum lanceolatum subsp. lanceolatum var. lanceolatum (Willd.) G.L. Nesom (=A. paniculatus Lam. p.p., non)
Uromyces phaseoli G. Winter
See: Uromyces appendiculatus (Pers.) Unger
Uromyces phaseoli var. strophostylis Arthur
See: Uromyces appendiculatus var. appendiculatus (Pers.) Unger
Uromyces phaseoli var. typica Arthur, nom. inval.
See: Uromyces appendiculatus var. appendiculatus (Pers.) Unger
Uromyces plumbarius Peck
Gaura biennis L. (Onagraceae) (Jackson 1915; Snyder 1896; Arthur 1899)
Oenothera biennis L. (Onagraceae) (Jackson 1915)
Uromyces polemonii (Peck) Barthol.
See: Uromyces acuminatus Arthur
Uromyces polygoni (Pers.) Fuckel
See: Uromyces polygoni-avicularis (Pers.) G.H. Otth
Uromyces polygoni-avicularis (Pers.) G.H. Otth
Polygonum aviculare L. (Polygonaceae) (Underwood 1893; Snyder 1896; Arthur 1899; Jackson 1915; Anderson and Anderson 1919)
P. erectum L. (Underwood 1893; Arthur 1899; Jackson 1915)
P. punctatum Elliott var. punctatum (=P. acre Kunth) (Underwood 1893)
Uromyces porosus (Peck) H.S. Jacks.
See: Uromyces coloradensis Ellis & Everh.
Uromyces proëminens (DC.) Lév.
Chamaesyce humistrata (Engelm. ex A. Gray) Small (Euphorbiaceae) (Jackson 1915); (=Euphorbia humistrata Engelm. ex A. Gray) (Anderson and Anderson 1919)
C. hypericifolia (L.) Millsp. (=E. hypericifolia L.) (Underwood 1893; Snyder 1896, 1898)
C. hyssopifolia (L.) Small
C. maculata (L.) Small (=E. maculata L.) (Underwood 1893; Snyder 1896; Jackson 1915; Anderson and Anderson 1919)
C. nutans (Lag.) Small (=C. preslii (Guss.) Arthur) (Jackson 1915); (=E. nutans Lag.) (Arthur 1899)
Euphorbia corollata L. (Euphorbiaceae) (Underwood 1893); (=E. preslii Guss.) (Underwood 1893; Anderson and Anderson 1919; Van Hook 1934)
E. dentata Michx. (Underwood 1893; Snyder 1896; Anderson and Anderson 1919); var. dentata (=Poinsettia dentata (Michx.) Klotzsch & Garcke) (Snyder 1896; Arthur 1899; Jackson 1915)
E. glyptosperma (Engelm.) Small
E. heterophylla L.; (=P. heterophylla (L.) Klotzsch & Garcke) (Jackson 1915)
E. maculata L.
Euphorbia sp. (Underwood 1893)
Host not reported (Van Hook 1910)
Uromyces punctatus J. Schröt.
Astragalus canadensis L. (Fabaceae)
Uromyces rhynchosporae Ellis
Rhynchospora alba (L.) Vahl (Cyperaceae) (Jackson 1915)
R. capitellata (Michx.) Vahl
R. glomerata (L.) Vahl (Jackson 1920)
Uromyces rudbeckiae Arthur & Holw.
Rudbeckia laciniata L. (Asteraceae) (Snyder 1898; Arthur 1899; Jackson 1915; Anderson and Anderson 1919)
Uromyces scirpi Burrill
See: Uromyces lineolatus subsp. nearcticus Savile
Uromyces seditiosus F. Kern
Aristida longespica Poir. (Poaceae) (Cummins and Husain 1966)
A. oligantha Michx. (=A. ramosissima Engelm.) (Jackson 1917; Jackson 1920; Cummins and Husain 1966)
A. adscensionis L. (McCain et al. 1990)
Plantago aristata Michx. (Plantaginaceae)
P. lanceolata L.
P. rugelii Decne.
Uromyces silphii (Syd. & P. Syd.) Arthur
Heliopsis helianthoides (L.) Sweet (Asteraceae)
Juncus anthelatus (Wiegand) R.E. Brooks (=J. tenuis var. anthelatus Wiegand) (Juncaceae)
J. dudleyi Wiegand (Jackson 1915)
Juncus sp.
J. tenuis Willd. (Jackson 1915)
Silphium integrifolium Michx. (Asteraceae)
S. perfoliatum L. (Jackson 1915)
Uromyces solidaginis-caricis Arthur
See: Uromyces perigynius Halst.
Uromyces sparganii Cooke & Peck
Acorus calamus L. (Acoraceae) (Parmelee and Savile 1954)
Uromyces spermacoces (Schwein.) M.A. Curtis
Diodia teres Walter (Rubiaceae) (Jackson 1915; Emmons et al. 1960)
Uromyces sporoboli Ellis & Everh.
Allium canadense L. (Amaryllidaceae)
Allium sp.
Sporobolus compositus var. compositus (Poir.) Merr. (=S. asper (P. Beauv.) Kunth) (Poaceae) (Cummins and Greene 1961)
S. vaginiflorus (Torr. ex A. Gray) Alph. Wood
Uromyces striatus J. Schröt.
Medicago lupulina L. (Fabaceae) (Jackson 1915; McCain et al. 1990)
M. sativa L. (Jackson 1915; Pipal 1915; Ruhl et al. 1982)
Uromyces terebinthi (DC.) G. Winter
See: Pileolaria brevipes Berk. & Ravenel
Uromyces trifolii (Alb. & Schwein.) G. Winter, sensu Underwood (1893)
See: Uromyces trifolii-repentis var. trifolii-repentis Liro and U. trifolii-repentis var. fallens (Arthur) Cumm.
Uromyces trifolii (Alb. & Schwein.) G. Winter
See: Uromyces trifolii-repentis var. fallens (Arthur) Cumm.
Host not reported (Van Hook 1910)
Uromyces trifolii (R. Hedw.) Lév.
See: Uromyces trifolii-repentis var. trifolii-repentis
Uromyces trifolii var. fallens (Desm.) Arthur
See: Uromyces trifolii-repentis var. fallens (Desm.) Arthur
Uromyces trifolii-repentis var. trifolii-repentis Liro
Trifolium hybridum L. (Fabaceae) (Underwood 1893; Arthur 1899; Jackson 1915)
T. medium L. (Jackson 1915)
T. repens L. (Underwood 1893; Arthur 1899; Jackson 1915)
T. campestre Schreb. (=T. procumbens L.) (Anderson and Anderson 1919) [Doubtful host]
Trifolium sp. (Pipal 1915)
Uromyces trifolii-repentis var. fallens (Desm.) Cumm.
Trifolium medium L. (Fabaceae) (Underwood 1893; USDA 1960)
T. pratense L. (Underwood 1893; Snyder 1896, 1898; Arthur 1899; Jackson 1915; Anderson and Anderson 1919)
T. repens L. (Jackson 1915) PUR: 15471
Trifolium sp. (Pipal 1915)
Uromyces triquetrus Cooke
Elodes campanulata Pursh (Clusiaceae) (Underwood 1893)
Hypericum canadense L. (Clusiaceae) (Underwood 1893; Arthur 1899; Jackson 1915)
H. adpressum W.P.C. Barton (McCain et al. 1990)
H. cistifolium Lam.
H. ellipticum Hook.
H. kalmianum L. (McCain and Hennen 1981)
H. mutilum L. (Underwood 1893; Arthur 1899; Jackson 1915)
H. prolificum L. (Jackson 1920; Emmons et al. 1960)
Hypericum sp. (Emmons et al. 1960)
Triadenum virginicum (L.) Raf. (Clusiaceae) (Arthur 1899; Jackson 1915)
Uromyces valens F. Kern
Carex lupulina Muhl. ex Willd. (Cyperaceae) (Jackson 1915); (=C. lupulina Muhl. ex Willd. var. pedunculata A. Gray) (Underwood 1893)
Carex sp. (USDA 1960)
C. rostrata Stokes (Jackson 1915)
C. utriculata Boott (Kern 1910)
Uromyces verruculosus J. Schröt.
Silene latifolia subsp. alba (Mill.) Greuter & Burdet (=Lychnis alba Mill.) (Caryophyllaceae) (USDA 1960)
Uromyces viciae-fabae (Pers.) J. Schröt.
Lathyrus palustris L. (Fabaceae) (Anderson and Anderson 1919)
Lathyrus sp.
L. venosus Muhl. ex Willd. (Jackson 1915); (reported as Vicia americana Muhl. ex Willd.) (Snyder 1896; Arthur 1899)
Uromyces vignae Barcl.
Vigna unguiculata (L.) Walp. (Fabaceae); (=Vigna sinensis (L.) Savi ex Hassk.) (Fabaceae) (Jackson 1915)
Uropyxis amorphae (M.A. Curtis) J. Schröt.
Amorpha canescens Pursh (Fabaceae) (Underwood 1893; Arthur 1899; Jackson 1915)
Acknowledgments
We are thankful to the Arthur Fungarium (PUR) and the numerous individuals who have enriched this collection.
Citation
Torres-Cruz TJ, Abbasi M, Simmons DR, Aime MC (2025) Checklist of Rust Fungi (Basidiomycota, Pucciniales) and their hosts in Indiana, United States of America. MycoKeys 121: 167–235. https://doi.org/10.3897/mycokeys.121.148853
Contributor Information
Terry J. Torres-Cruz, Email: terryjari@gmail.com.
M. Catherine Aime, Email: maime@purdue.edu.
Additional information
Conflict of interest
The authors have declared that no competing interests exist.
Ethical statement
No ethical statement was reported.
Funding
This work was funded by NSF DBI 1502887 and 1458290; NSF DEB 2127290; USDA NIFA Hatch project 1010662; USDA APHIS AP24PPQS & T00C027 and AP20PPQS & T00C077; and the Indiana Academy of Sciences IP 14033562.
Author contributions
Conceptualization: MA, MCA. Data curation: TJJTC, MCA, MA. Formal analysis: MA, TJJTC. Funding acquisition: MCA. Investigation: MA. Methodology: MCA, MA. Project administration: MCA. Resources: DRS. Supervision: MCA. Validation: MA, MCA. Visualization: TJJTC. Writing - original draft: TJJTC. Writing - review and editing: TJJTC, MCA, DRS, MA.
Author ORCIDs
Terry J. Torres-Cruz https://orcid.org/0000-0002-9193-3249
D. Rabern Simmons https://orcid.org/0000-0002-7033-225X
M. Catherine Aime https://orcid.org/0000-0001-8742-6685
Data availability
All of the data that support the findings of this study are available in the main text or Supplementary Information.
Supplementary materials
Number of rust specimens deposited in the Arthur Fungarium by number of specimens per Indiana county ranked from highest to lowest
This dataset is made available under the Open Database License (http://opendatacommons.org/licenses/odbl/1.0/). The Open Database License (ODbL) is a license agreement intended to allow users to freely share, modify, and use this Dataset while maintaining this same freedom for others, provided that the original source and author(s) are credited.
Terry J. Torres-Cruz
Data type
xls
Metadata of specimens evaluated in the study
This dataset is made available under the Open Database License (http://opendatacommons.org/licenses/odbl/1.0/). The Open Database License (ODbL) is a license agreement intended to allow users to freely share, modify, and use this Dataset while maintaining this same freedom for others, provided that the original source and author(s) are credited.
Terry J. Torres-Cruz
Data type
xls
Results of searching all taxa reported for Indiana in MycoBank and IndexFungorum databases
This dataset is made available under the Open Database License (http://opendatacommons.org/licenses/odbl/1.0/). The Open Database License (ODbL) is a license agreement intended to allow users to freely share, modify, and use this Dataset while maintaining this same freedom for others, provided that the original source and author(s) are credited.
Terry J. Torres-Cruz
Data type
xlsx
References
- Abbasi M, Aime MC. (2016) Two new Puccinia species on Melica (Poaceae) from USA. Mycotaxon 131(1): 247–253. 10.5248/131.247 [DOI] [Google Scholar]
- Abbasi M, Goodwin SB, Scholler M. (2005) Taxonomy, phylogeny, and distribution of Puccinia graminis, the black stem rust: New insights based on rDNA sequence data. Mycoscience 46(4): 241–247. 10.47371/mycosci.MYC46241 [DOI] [Google Scholar]
- Abbasi M, Aime MC, Creswell TC, Ruhl GE. (2016a) First report of rust disease caused by Coleosporium apocynaceum on Amsonia ‘Blue Ice’ in Indiana. Plant Disease 100(8): 1786. 10.1094/PDIS-01-16-0019-PDN [DOI] [Google Scholar]
- Abbasi M, Klimek JF, Aime MC. (2016b) First report of rust disease caused by Puccinia liliacearum on Ornithogalum umbellatum from Indiana and Maryland with notes on the spread of the rust fungus in the United States. Plant Disease 100(10): 2169. 10.1094/PDIS-04-16-0509-PDN [DOI] [Google Scholar]
- Aime MC, Abbasi M. (2018) Puccinia modiolae in North America: Distribution and natural host range. MycoKeys 11(39): 63–73. 10.3897/mycokeys.39.27378 [DOI] [PMC free article] [PubMed] [Google Scholar]
- Aime MC, McTaggart AR. (2021) A higher-rank classification for rust fungi, with notes on genera. Fungal Systematics and Evolution 7(1): 21–47. 10.3114/fuse.2021.07.02 [DOI] [PMC free article] [PubMed] [Google Scholar]
- Aime MC, Bell CD, Wilson AW. (2018) Deconstructing the evolutionary complexity between rust fungi (Pucciniales) and their plant hosts. Studies in Mycology 89: 143–152. 10.1016/j.simyco.2018.02.002 [DOI] [PMC free article] [PubMed] [Google Scholar]
- Anderson HW, Anderson PJ. (1919) The parasitic Fungi of Montgomery County, I. Proceedings of the Indiana Academy of Sciences 29: 175–222. https://journals.indianapolis.iu.edu/index.php/ias/article/view/13500 [Google Scholar]
- Arthur JC. (1899) Indiana plant rusts, listed in accordance with latest nomenclature. Proceedings of the Indiana Academy of Sciences 8: 174–186. https://journals.indianapolis.iu.edu/index.php/ias/article/view/13487 [Google Scholar]
- Arthur J. (1902) The Uredineæ Occurring upon Phragmites, Spartina, and Arundinaria in America. Botanical Gazette (Chicago, Ill. ) 34(1): 1–20. 10.1086/328256 [DOI] [Google Scholar]
- Arthur JC. (1909) Cultures of Uredineae in 1908. Mycologia 1(6): 225–256. 10.1080/00275514.1909.12020595 [DOI] [Google Scholar]
- Arthur JC. (1918) New Species of Uredineae-X. Bulletin of the Torrey Botanical Club 45(4): 141–156. 10.2307/2479631 [DOI] [Google Scholar]
- Arthur JC. (1934) Manual of the rusts in United States and Canada. Purdue Research Foundation, Lafayette, Indiana, 438 pp. [Google Scholar]
- Arthur JC, Holway E. (1901) Violet rusts in North America. Minnesota Botanical Studies 2: 631–641. [Google Scholar]
- Bates ST, Bailey EM, Tobey NT. (2019) Checklist of Indiana Fungi II: Microfungi. Proceedings of the Indiana Academy of Sciences 128(2): 107–130. [Google Scholar]
- Batra SWT. (1981) Puccinia xanthii forma specialis ambrosiatrifidae, a microcyclic rust for the biological control of giant ragweed, Ambrosia trifida (Compositae). Mycopathologia 73(2): 61–64. 10.1007/BF00562590 [DOI] [Google Scholar]
- Bean DW, Gladem K, Rosen K, Blake A, Clark RE, Henderson C, Kaltenbach J, Price J, Smallwood EL, Berner DK, Young SL, Schaeffer RN. (2024) Scaling use of the rust fungus Puccinia punctiformis for biological control of Canada thistle (Cirsium arvense (L.) Scop.): First report on a U.S. statewide effort. Biological Control 192: 105481. 10.1016/j.biocontrol.2024.105481 [DOI]
- Bennett C, Aime MC, Newcombe G. (2011) Molecular and pathogenic variation within Melampsora on Salix in western North America reveals numerous cryptic species. Mycologia 103(5): 1004–1018. 10.3852/10-289 [DOI] [PubMed] [Google Scholar]
- Böllmann J, Scholler M. (2006) Life cycle and life strategy features of Puccinia glechomatis (Uredinales) favorable for extending the natural range of distribution. Mycoscience 47(3): 152–158. 10.47371/mycosci.MYC47152 [DOI] [Google Scholar]
- Buriticá P. (1998) La Familia Phakopsoraceae en el Neotrópico - II. Generos: Arthuria, Nothoravenelia, Uredopeltis, Kweilingia, Aplopsora y Pucciniostele. Revista de la Academia Colombiana de Ciencias Exactas, Físicas y Naturales 22(80): 325–334. 10.18257/raccefyn.22(84).1998.2916 [DOI] [Google Scholar]
- Buriticá P. (1999) La familia Phakopsoraceae en el Neotrópico III, géneros: Batistopsora y Phakopsora. Revista de la Academia Colombiana de Ciencias Exactas, Físicas y Naturales 23(87): 271–305. 10.18257/raccefyn.23(87).1999.2906 [DOI]
- Cantonwine E, Nienow J, Blackmore M, Griffin B, Bergstrom B. (2019) Results of a fall and spring BioBlitz at Grassy Pond Recreational Area, Lowndes County, Georgia. Georgia Journal of Science 77: 17.
- Cash EK. (1952) A record of the fungi named by J.B. Ellis., Part I. Division of Mycology and Disease Survey, Bureau of Plant Industry, Soils, and Agricultural Engineering, Agricultural Research Administration, USDA. 10.5962/bhl.title.149755 [DOI]
- Caubel J, Launay M, Ripoche D, Gouache D, Buis S, Huard F, Huber L, Brun F, Bancal MO. (2017) Climate change effects on leaf rust of wheat: Implementing a couple crop-disease model in a French regional application. European Journal of Agronomy 90: 53–66. 10.1016/j.eja.2017.07.004 [DOI] [Google Scholar]
- Cellerino GP. (1999) Review of fungal diseases in Poplar. Grugliasco, Turin, 53 pp. https://openknowledge.fao.org/server/api/core/bitstreams/a247e7ff-5ddf-4e78-9bfc-2f8326c0e8d1/content [Google Scholar]
- Çelekli A, Zariç ÖE. (2023) Utilization of herbaria in ecological studies: biodiversity and landscape monitoring. Herbarium Turcicum. 10.26650/HT.2023.1345916 [DOI]
- Cooke WB. (1967) The 1961 Indiana Foray. Mycologia 59(2): 375–381. https://www.jstor.org/stable/3756813 [Google Scholar]
- Cooke WB. (1975) The 1970 Indiana Foray. Mycologia 67(5): 1065–1071. 10.1080/00275514.1975.12019846 [DOI] [Google Scholar]
- Cooke WB. (1986) The 1981 Indiana Foray. Mycologia 78(2): 321–323. 10.1080/00275514.1986.12025252 [DOI] [Google Scholar]
- Coulter JM. (1876) Uromyces lespedezae. Botanical Gazette (Chicago, Ill. ) 1(5): 20. [Google Scholar]
- Creswell TC, Ruhl GE, Aime MC, Beckerman JL, Abbasi M. (2016) First report of rust disease of Bradford pear caused by Gymnosporangium clavipes. Plant Disease 100(4): 860. 10.1094/PDIS-10-15-1125-PDN [DOI]
- Cummins GB. (1935) Notes on some species of the Uredinales. Mycologia 27(6): 605–614. 10.1080/00275514.1935.12017104 [DOI] [Google Scholar]
- Cummins GB. (1962) Supplement to Arthur’s Manual of the rusts in United States and Canada. Hafner, New York, 24 pp. [Google Scholar]
- Cummins GB. (1971) The Rust Fungi of cereals, grasses and bamboos. Springer-Verlag, New York, 570 pp. 10.1007/978-3-642-88451-1 [DOI] [Google Scholar]
- Cummins GB. (1978) Rust fungi on legumes and composites in North America. University of Arizona Press, Tucson, AZ.
- Cummins G, Greene H. (1961) The rust fungi of Muhlenbergia, Sporobolus, and related genera. Brittonia 13(3): 271–285. 10.2307/2805343 [DOI] [Google Scholar]
- Cummins GB, Greene HC. (1966) A review of the grass rust fungi that have uredial paraphyses and aecia on Berberis-Mahonia. Mycologia 58(5): 702–721. 10.1080/00275514.1966.12018364 [DOI] [Google Scholar]
- Cummins GB, Husain SM. (1966) The rust fungi on the genus Aristida. Bulletin of the Torrey Botanical Club 93(1): 56–67. 10.2307/2483886 [DOI]
- Cummins G, Britton M, Baxter J. (1969) The autoecious species of Puccinia on North American Eupatoriae. Mycologia 61(5): 924–944. 10.1080/00275514.1969.12018816 [DOI] [Google Scholar]
- Demers JE, Liu M, Hambleton S, Castlebury LA. (2017) Rust fungi on Panicum. Mycologia 109(1): 1–17. 10.1080/00275514.2016.1262656 [DOI] [PubMed]
- Dudney J, Willing CE, Das AJ, Latimer AM, Nesmith JCB, Battles JJ. (2021) Nonlinear shifts in infections rust disease due to climate change. Nature Communications 12(1): 5102. 10.1038/s41467-021-25182-6 [DOI] [PMC free article] [PubMed] [Google Scholar]
- Durkin L, Jansson T, Sanchez M, Khomich M, Ryberg M, Kristiansson E, Nilsson RH. (2020) When mycologists describe new species, not all relevant information is provided (clearly enough). MycoKeys 72: 109–128. 10.3897/mycokeys.72.56691 [DOI] [PMC free article] [PubMed] [Google Scholar]
- Emmons CW, Cummins GB, Cooke WB. (1960) The 1958 Foray of the Mycological Society of America. Mycologia 52(5): 808–817. https://www.jstor.org/stable/3755883 [Google Scholar]
- Farr DF, Rossman AY. (2018) Fungal Databases, U.S. National Fungus Collections, ARS, USDA. https://nt.ars-grin.gov/fungaldatabases/ [Accessed October 2024]
- Furnas A. (1874) Rust in blackberries. Transactions of the Indiana Horticultural Society 14: 74.
- Greve M, Lykke AM, Fagg CW, Gereau RE, Lewis GP, Marchant R, Marshall AR, Ndayishimiye J, Bogaert J, Svenning J-C. (2016) Realising the potential of herbarium records for conservation biology. South African Journal of Botany 105: 317–323. 10.1016/j.sajb.2016.03.017 [DOI] [Google Scholar]
- Hanlin RT, Foudin LL, Berisford Y, Glover SU, Jones JP, Huang LH. (1978) Plant disease index for maize in the US part 1: Host index. Agricultural Experimental Stations, University of Georgia. Research Reports (Montgomery) 277: 1–62. [Google Scholar]
- Heberling JM, Prather LA, Tonsor SJ. (2019) The changing uses of herbarium data in an era of global change: An overview using automated content analysis. Bioscience 69(10): 812–822. 10.1093/biosci/biz094 [DOI] [Google Scholar]
- Hennen JF. (1965) The species of Uromyces parasitic on the grass tribe Andropogoneae. Mycologia 57(1): 104–113. 10.1080/00275514.1965.12018196 [DOI] [Google Scholar]
- Hernández JR, Palm ME, Castlebury LA. (2002) Puccinia hemerocallidis, cause of daylily rust, a newly introduced disease in the Americas. Plant Disease 86(11): 1194–1198. 10.1094/PDIS.2002.86.11.1194 [DOI] [PubMed] [Google Scholar]
- Hiratsuka N. (1958) Revision of Taxonomy of the Pucciniastreae: With Special Reference to Species of the Japanese Archipelago. KASAI Publishing and Printing Co., Tokyo, 167 pp. [Google Scholar]
- IPBES (2016) The assessment report of the intergovernmental science-policy platform on biodiversity and ecosystem services on pollinators, pollination and food production. In: Potts SG, Imperatriz-Fonseca VL, Ngo HT (Eds) Secretariat of the Intergovernmental Science-Policy Platform on Biodiversity and Ecosystem Services. IPBES Secretariat, Bonn, Germany.
- IUCN (2020) The IUCN red list of threatened species. Version 2020–1.
- Jackson HS. (1915) The Uredinales of Indiana. Proceedings of the Indiana Academy of Sciences 25: 429–476. https://journals.indianapolis.iu.edu/index.php/ias/article/view/14504 [Google Scholar]
- Jackson HS. (1917) The Uredinales of Indiana II. Proceedings of the Indiana Academy of Sciences 27: 133–138. https://journals.indianapolis.iu.edu/index.php/ias/article/view/13289 [Google Scholar]
- Jackson HS. (1920) The Uredinales of Indiana III. Proceedings of the Indiana Academy of Sciences 30: 165–182. https://journals.indianapolis.iu.edu/index.php/ias/article/view/13580 [Google Scholar]
- Jones CA, Daehler CC. (2018) Herbarium specimens can reveal impacts of climate change on plant phenology; a review of methods and applications. PeerJ 6: e4576. 10.7717/peerj.4576 [DOI] [PMC free article] [PubMed]
- Kaishian P, Layug CRK, Anderson M, Berg DR, Aime MC. (2024) Rust HUBB: DNA barcode-based identification of Pucciniales. IMA Fungus 15(1): 3. 10.1186/s43008-023-00132-7 [DOI] [PMC free article] [PubMed] [Google Scholar]
- Kaishian P, Creswell T, Bonkowski J, Aime MC. (2023) First report of smoketree rust caused by Pileolaria cotini-coggygriae in the Midwestern United States. Plant Health Progress 24(2): 226–227. 10.1094/PHP-08-22-0080-BR [DOI] [Google Scholar]
- Kenaley SC, Quan M, Aime MC, Bergstrom GC. (2018) New insight into the species diversity and life cycles of rust fungi (Pucciniales) affecting bioenergy switchgrass (Panicum virgatum) in the Eastern and Central United States. Mycological Progress 17(11): 1251–1267. 10.1007/s11557-018-1434-1 [DOI] [Google Scholar]
- Kern FD. (1910) Two New Species of Uromyces on Carex. Rhodora 12(138): 124–127. https://biostor.org/reference/179036 [Google Scholar]
- Kern FD. (1919) North American rusts on Cyperus and Eleocharis. Mycologia 11(3): 134–147. 10.1080/00275514.1919.12016788 [DOI]
- Lacher Jr TE, Mallon D, Kennerley RJ, Relton C, Young RP. (2022) Tools and metrics for species prioritization for conservation planning and action: Case studies for antelopes and small mammals. Diversity 14(9): 704. 10.3390/d14090704 [DOI] [Google Scholar]
- Lang PLM, Willems FM, Scheepens JF, Burbano HA, Bossdorf O. (2019) Using herbaria to study global environmental change. The New Phytologist 221(1): 110–122. 10.1111/nph.15401 [DOI] [PMC free article] [PubMed] [Google Scholar]
- Laundon GF. (1963) Rust fungi 1: On Acanthaceae. Mycological Papers 89: 1–89. [Google Scholar]
- Lee HJ. (1929) New parasitic Fungi of Montgomery County. Proceedings of the Indiana Academy of Sciences 39: 135–136. https://journals.indianapolis.iu.edu/index.php/ias/article/view/5519 [Google Scholar]
- Lopez-Franco RM, Hennen JF. (1990) The genus Tranzschelia (Uredinales) in the Americas. Systematic Botany 15(4): 560–591. 10.2307/2419155 [DOI] [Google Scholar]
- Lowman M, Ruppert N, Mohd Nor SA. (2018) Further advancing the expert bioblitz for the rainforest conservation toolkit. Conservation Science and Practice 1(1): e2. 10.1111/csp2.2 [DOI]
- Maharani N, Kusrini MD, Hamidy A. (2022) Increasing herpetofauna data through citizen science in Indonesia. IOP Conference Series. Earth and Environmental Science 950(1): 012063. 10.1088/1755-1315/950/1/012063 [DOI] [Google Scholar]
- Mains EB. (1933) Host specialization in the leaf rust of grasses Puccinia rubigo-vera. Papers of the Michigan Academy of Science, Arts and Letters 17: 289–394.
- McCain JW. (1984) A preliminary review and multiple-entry key to the rust fungi on Cyperaceae and Juncaceae in Indiana. Proceedings of the Indiana Academy of Sciences 94: 447–454. https://journals.indianapolis.iu.edu/index.php/ias/article/view/8380 [Google Scholar]
- McCain JW, Hennen JF. (1981) Notes on biogeography and new records of rust Fungi in the Great Lakes Region. Proceedings of the Indiana Academy of Sciences 91: 504–514. https://journals.indianapolis.iu.edu/index.php/ias/article/view/7773 [Google Scholar]
- McCain JW, Hennen JF. (1982) The Arthur Herbarium centennial: 100 years of uredinology in Indiana and the Great Lakes Region. Proceedings of the Indiana Academy of Sciences 92: 389–395. https://journals.indianapolis.iu.edu/index.php/ias/article/view/8115 [Google Scholar]
- McCain JW, Hennen JF, Ono Y. (1990) New host species and state distribution records for North American rust fungi (Uredinales). Mycotaxon 39: 281–300. [Google Scholar]
- McTaggart AR, Aime MC. (2018) The species of Coleosporium (Pucciniales) on Solidago in North America. Fungal Biology 122(8): 800–809. 10.1016/j.funbio.2018.04.007 [DOI] [PubMed] [Google Scholar]
- Menchetti M, Cianferoni F, Mazza G, Dal Cin M, Barbato D, Benocci A, Cervo R, Dapporto L, Picchi MS, Vanni L, Cabrini R, Mori E. (2021) Checklist of macro-invertebrates of the special conservation area “Poggi di Prata” (Grosseto, Central Italy) through a citizen-science and expert-based approach. Redia (Firenze) 104: 63–68. 10.19263/REDIA-104.21.07 [DOI] [Google Scholar]
- Microsoft Corporation (2024) Microsoft Excel.
- Miller IF, Jiranek J, Brownell M, Coffrey S, Gray B, Stahl M, Metcalf JE. (2022) Predicting the effects of climate change on the cross-scale epidemiological dynamics of a fungal plant pathogen. Scientific Reports 12(1): 14823. 10.1038/s41598-022-18851-z [DOI] [PMC free article] [PubMed] [Google Scholar]
- Molano-Flores B, Johnson SA, Marcum PB, Feist MA. (2023) Utilizing herbarium specimens to assist with the listing of rare plants. Frontiers in Conservation Science 4: 1–15. 10.3389/fcosc.2023.1144593 [DOI] [Google Scholar]
- Monoson HL. (1975) The species of Triphragmium, Nyssopsora, and Triphragmiopsis. Mycopathologia 52(2): 115–131. 10.1007/BF02128054 [DOI] [Google Scholar]
- MyCoPortal (2025) MyCoPortal. http://www.mycoportal.org/portal/index.php [Accessed on April 2025]
- Newcombe G, Striling B, McDonals S, Bradshaw HD. (2000) Melampsora × columbiana, a natural hybrid of M. medusae and M. occidentalis. Mycological Research 104(3): 261–274. 10.1017/S0953756299001665 [DOI]
- Nicolai A, Guernion M, Guillocheau S, Hoeffner K, Le Gouar P, Ménard N, Piscart C, Vallet D, Hervé ME, Benezeth E, Chedanne H, Blémus J, Vernon P, Cylly D, Hotte H, Loïs G, Mai B, Perez G, Ouisse T, Monard C, Wiegand C, Caudal J-P, Butet A, Dahirel M, Barbe L, Balbi M, Briand V, Bormans M, Charrier M, Bouger G, Jung V, Lann CL, Pannard A, Petillon J, Rantier Y, Marguerie D, Tougeron K, Devogel P, Dugravot S, Dubos T, Garrin M, Carnet M, Gouraud C, Chambet A, Esnault J, Poupelin M, Welk E, Bütof A, Dubois GF, Humbert G, Marie-Réau O, Norvez O, Richard G, Froger B, Rochais C, Potthoff M, Ayati K, Bellido A, Rissel A, Santonja M, Farcy J-O, Collias E, Sene L, Cluzeau D, Supper R. (2020) Transdisciplinary Bioblitz: Rapid biotic and abiotic inventory allows studying environmental changes over 60 years at the biological field station of Paimpont (Brittany, France) and opens new interdisciplinary research opportunities. Biodiversity Data Journal 8: e50451. 10.3897/BDJ.8.e50451 [DOI] [PMC free article] [PubMed]
- Otálora MAG, Berndt R. (2018) A taxonomic revision of the genus Puccinia on Lycieae, a tribe of Solanaceae. Mycologia 110(4): 692–709. 10.1080/00275514.2018.1478538 [DOI] [PubMed] [Google Scholar]
- Park DA, Lyra GM, Ellison AM, Barbosa Maruyama RK, Reis Torquato D, Asprino RC, Cook BI, Davis CC. (2022) Herbarium records provide reliable phenology estimates in the understudied tropics. Journal of Ecology 111(2): 327–337. 10.1111/1365-2745.14047 [DOI] [Google Scholar]
- Parker SS, Pauly GB, Moore J, Fraga NS, Knapp JJ, Principe Z, Brown BV, Randall JM, Cohen BS, Wake TA. (2018) Adapting the bioblitz to meet conservation needs. Conservation Biology 32(5): 1007–1019. 10.1111/cobi.13103 [DOI] [PubMed] [Google Scholar]
- Parmelee JA, Savile DBO. (1954) Life history and relationship of the rusts of Sparganium and Acorus. Mycologia 46(6): 823–836. 10.1080/00275514.1954.12024419 [DOI]
- Parmelee JA, Savile DBO. (1981) Autoecious species of Puccinia on Cichorieae in North America. Canadian Journal of Botany 59(6): 1078–1101. 10.1139/b81-147 [DOI] [Google Scholar]
- Pearman PB, Broennimann O, Aavik T, Albayrak T, Alves PC, Aravanopoulos FA, Bertola LD, Biedrzycka A, Buzan E, Cubric-Curik V, Djan M, Fedorca A, Fuentes-Pardo AP, Fussi B, Godoy JA, Gugerli F, Hoban S, Holderegger R, Hvilsom C, Lacolina L, Kalamujic B, Klinga P, Konopiński MK, Kopatz A, Laikre L, Lopes-Fernandes M, McMahon BJ, Mergeay J, Neophytou C, Pálsson S, Paz-Vinas I, Posledovich D, Primmer CR, Raymaekers JAM, Rinkevich B, Rolečková B, Runġis D, Schuerz L, Segelbacher G, Kavčič K, Stefanovic M, Thurfjell H, Träger S, Tsvetkov IN, Velickovic N, Vergeer P, Vernesi C, Vilà C, Westergren M, Zachos FE, Guisan A, Bruford M. (2024) Monitoring of species’ genetic diversity in Europe varies greatly and overlooks potential climate change impacts. Nature Ecology & Evolution 8(2): 267–281. 10.1038/s41559-023-02260-0 [DOI] [PMC free article] [PubMed] [Google Scholar]
- Pierson T, Stratmann T, White EC, Clause A, Carter C, Herr M, Jenkins A, Vogel H, Knoerr M, Folt B. (2014) New county records of amphibians and reptiles resulting from a bioblitz competition in North-Central Georgia, USA. Herpetological Review 45: 296–297. [Google Scholar]
- Pipal FJ. (1915) A list of plant diseases of economic importance in Indiana with bibliography. Proceedings of the Indiana Academy of Sciences 25: 379–414. https://journals.indianapolis.iu.edu/index.php/ias/article/view/14498 [Google Scholar]
- Ramachar P, Cummins GB. (1965) The species of Puccinia on the Paniceae. Mycopathologia 25(1–2): 7–60. 10.1007/BF02049611 [DOI] [Google Scholar]
- Reyserhove L, Desmet P, Oldoni D, Adriaens T, Strubbe D, Davis AJS, Vanderhoeven S, Verloove F, Groom Q. (2020) A checklist recipe: Making species data open and FAIR. Database (Oxford) 2020: baaa084. 10.1093/database/baaa084 [DOI] [PMC free article] [PubMed]
- Ruhl GE, McCain JW. (1986) First report of rust caused by Frommeella duchesneae on false strawberry in Indiana. Plant Disease 70(5): 475. 10.1094/PD-70-475c [DOI] [Google Scholar]
- Ruhl GE, Scott DH, Pecknold PC. (1980) A compilation of plant diseases and disorders in Indiana. Proceedings of the Indiana Academy of Sciences 90: 107–124. https://journals.indianapolis.iu.edu/index.php/ias/article/view/5835 [Google Scholar]
- Ruhl GE, Latin RX, Pecknold PC, Scott DH. (1981) A compilation of plant diseases and disorders in Indiana. Proceedings of the Indiana Academy of Sciences 91: 120–139. https://journals.indianapolis.iu.edu/index.php/ias/article/view/7435 [Google Scholar]
- Ruhl GE, Latin RX, Pecknold PC, Scott DH. (1982) Compilation of plant diseases and disorders in Indiana. Proceedings of the Indiana Academy of Sciences 92: 97–118. https://journals.indianapolis.iu.edu/index.php/ias/article/view/8035 [Google Scholar]
- Ruhl GE, Latin RX, Pecknold PC, Scott DH. (1983) A compilation of plant diseases and disorders in Indiana. Proceedings of the Indiana Academy of Sciences 93: 103–120. https://journals.indianapolis.iu.edu/index.php/ias/article/view/7515 [Google Scholar]
- Savile DBO. (1970) Autoecious Puccinia species attacking Cardueae in North America. Canadian Journal of Botany 48(9): 1553–1566. 10.1139/b70-232 [DOI] [Google Scholar]
- Savile DBO. (1971) Coevolution of the Rust Fungi and Their Hosts. The Quarterly Review of Biology 46(3): 211–218. 10.1086/406895 [DOI] [Google Scholar]
- Savile DBO. (1972) Some rusts of Scirpus and allied genera. Canadian Journal of Botany 50(12): 2579–2596. 10.1139/b72-331 [DOI] [Google Scholar]
- Savile DBO. (1976) Phragmidium ivesiae and its allies in North America. Canadian Journal of Botany 54(14): 1690–1696. 10.1139/b76-182 [DOI] [Google Scholar]
- Schall RA, McCain JW, Hennen JF. (1983) Distribution of Puccinia polysora in Indiana and the absence of cool weather form as determined by comparison with P. sorghi. Plant Disease 67(7): 767–770. 10.1094/PD-67-767 [DOI] [Google Scholar]
- Schoknecht JD. (1981) Contribution to the Flora of Indiana: Fungi of west-central Indiana on the occasion of the sixth annual A. H. Smith Great Lakes foray. Proceedings of the Indiana Academy of Sciences 91: 140–143. https://journals.indianapolis.iu.edu/index.php/ias/article/view/7436 [Google Scholar]
- Scholler M. (2000) Rust on ground-ivy found for the first time in North America. Plant Disease 84(3): 371. 10.1094/PDIS.2000.84.3.371D [DOI] [PubMed] [Google Scholar]
- Seebens H, Blackburn TM, Dyer EE, Genovesi P, Hulme PE, Jeschke JM, Pagad S, Pyšek P, Winter M, Arianoutsou M, Bacher S, Blasius B, Brundu G, Capinha C, Celesti-Grapow L, Dawson W, Dullinger S, Fuentes N, Jäger H, Kartesz J, Kenis M, Kreft H, Kühn I, Lenzner B, Liebhold A, Mosena A, Moser D, Nishino M, Pearman D, Pergl J, Rabitsch W, Rojas-Sandoval J, Roques A, Rorke S, Rossinelli S, Roy HE, Scalera R, Schindler S, Štajerová K, Tokarska-Guzik B, van Kleunen M, Walker K, Weigelt P, Yamanaka T, Essl F. (2017) No saturation in the accumulation of alien species worldwide. Nature Communications 8(1): 14435. 10.1038/ncomms14435 [DOI] [PMC free article] [PubMed] [Google Scholar]
- Snyder L. (1896) The Uredineae of Tippecanoe County, Ind. Proceedings of the Indiana Academy of Sciences 6: 216–224. https://journals.indianapolis.iu.edu/index.php/ias/article/view/13130 [Google Scholar]
- Snyder L. (1898) The Uredineae of Madison and Noble Counties with additional specimens from Tippecanoe County. Proceedings of the Indiana Academy of Sciences 8: 186–189. https://journals.indianapolis.iu.edu/index.php/ias/article/view/13488 [Google Scholar]
- Tanner RA, Pollard KM, Varia S, Evans HC, Ellison CA. (2015) First release of a fungal classical biocontrol agent against an invasive alien weed in Europe: Biology of the rust, Puccinia komarovii var. glanduliferae. Plant Pathology 64(5): 1130–1139. 10.1111/ppa.12352 [DOI]
- Ullstrup AJ, Lavoilette FA. (1959) Diseases of corn and of sorghum species in Indiana in 1958. The Plant Disease Reporter 43: 334–336. [Google Scholar]
- Underwood LM. (1893) List of cryptogams at present known to inhabit the state of Indiana. Proceedings of the Indiana Academy of Sciences 3: 30–67. https://journals.indianapolis.iu.edu/index.php/ias/article/view/12643 [Google Scholar]
- Underwood LM. (1896) Additions to the published lists of Indiana cryptogams. Proceedings of the Indiana Academy of Sciences 6: 171–172. https://journals.indianapolis.iu.edu/index.php/ias/article/view/13120 [Google Scholar]
- United States Department of Agriculture (1960) Index of Plant Diseases in the United States. US Government Printing Office, Washington, DC. https://www.govinfo.gov/content/pkg/GOVPUB-A-PURL-gpo20004/pdf/GOVPUB-A-PURL-gpo20004.pdf
- Urban Z. (1971) The autoecious species of Puccinia on Vernonieae in North America. Acta Universitatis Carolinae. Biologica: 1–84.
- Van Hook JM. (1910) Indiana Fungi. Proceedings of the Indiana Academy of Sciences 20: 205–212. https://journals.indianapolis.iu.edu/index.php/ias/article/view/14883 [Google Scholar]
- Van Hook JM. (1912) Indiana Fungi— III. Proceedings of the Indiana Academy of Sciences 22: 99–101. https://journals.indianapolis.iu.edu/index.php/ias/article/view/14099 [Google Scholar]
- Van Hook JM. (1921) Indiana Fungi— VI. Proceedings of the Indiana Academy of Sciences 31: 143–148. https://journals.indianapolis.iu.edu/index.php/ias/article/view/13884 [Google Scholar]
- Van Hook JM. (1923) Indiana Fungi— VII. Proceedings of the Indiana Academy of Sciences 33: 233–238. https://journals.indianapolis.iu.edu/index.php/ias/article/view/13999 [Google Scholar]
- Van Hook JM. (1924) Indiana Fungi— VIII. Proceedings of the Indiana Academy of Sciences 34: 317–320. https://journals.indianapolis.iu.edu/index.php/ias/article/view/3901 [Google Scholar]
- Van Hook JM. (1925) Indiana Fungi— IX. Proceedings of the Indiana Academy of Sciences 35: 233–236. https://journals.indianapolis.iu.edu/index.php/ias/article/view/4196 [Google Scholar]
- Van Hook JM. (1927) Indiana Fungi— X. Proceedings of the Indiana Academy of Sciences 37: 365–372. https://journals.indianapolis.iu.edu/index.php/ias/article/view/4572 [Google Scholar]
- Van Hook JM. (1928) Indiana Fungi— XI. Proceedings of the Indiana Academy of Sciences 38: 127–130. https://journals.indianapolis.iu.edu/index.php/ias/article/view/4976 [Google Scholar]
- Van Hook JM. (1929) Indiana Fungi, XII. Proceedings of the Indiana Academy of Sciences 39: 75–84. https://journals.indianapolis.iu.edu/index.php/ias/article/view/5182 [Google Scholar]
- Van Hook JM. (1934) Indiana Fungi, XIII. Proceedings of the Indiana Academy of Sciences 44: 55–64. https://journals.indianapolis.iu.edu/index.php/ias/article/view/4654 [Google Scholar]
- Vargas-Gaete R, Doussoulin H, Smith-Ramírez C, Bravo S, Salas-Eljatib C, Andrade N, Trávníček B. (2019) Evaluation of rust pathogenicity (Phragmidium violaceum) as a biological control agent for the invasive plant Rubus ulmifolius on Robinson Crusoe Island, Chile. Australasian Plant Pathology 48(3): 201–208. 10.1007/s13313-019-0615-y [DOI] [Google Scholar]
- Walker JT, Earhart EF. (1962) Medlar is host for Gymnosporangium clavipes. The Plant Disease Reporter 46: 293.
- Williams TM, Schlichting CD, Holsinger KE. (2021) Herbarium records demonstrate changes in flowering phenology associated with climate change over the past century within the Cape Floristic Region, South Africa. Climate Change Ecology 1: 100006. 10.1016/j.ecochg.2021.100006 [DOI]
- Wilson A, Beckerman J, Aime M. (2014) First report of the White Pine Blister Rust Fungus, Cronartium ribicola, on Ribes odoratum in Indiana. Plant Disease 98(2): 277. 10.1094/PDIS-04-13-0442-PDN [DOI] [PubMed] [Google Scholar]
Associated Data
This section collects any data citations, data availability statements, or supplementary materials included in this article.
Supplementary Materials
Number of rust specimens deposited in the Arthur Fungarium by number of specimens per Indiana county ranked from highest to lowest
This dataset is made available under the Open Database License (http://opendatacommons.org/licenses/odbl/1.0/). The Open Database License (ODbL) is a license agreement intended to allow users to freely share, modify, and use this Dataset while maintaining this same freedom for others, provided that the original source and author(s) are credited.
Terry J. Torres-Cruz
Data type
xls
Metadata of specimens evaluated in the study
This dataset is made available under the Open Database License (http://opendatacommons.org/licenses/odbl/1.0/). The Open Database License (ODbL) is a license agreement intended to allow users to freely share, modify, and use this Dataset while maintaining this same freedom for others, provided that the original source and author(s) are credited.
Terry J. Torres-Cruz
Data type
xls
Results of searching all taxa reported for Indiana in MycoBank and IndexFungorum databases
This dataset is made available under the Open Database License (http://opendatacommons.org/licenses/odbl/1.0/). The Open Database License (ODbL) is a license agreement intended to allow users to freely share, modify, and use this Dataset while maintaining this same freedom for others, provided that the original source and author(s) are credited.
Terry J. Torres-Cruz
Data type
xlsx
Data Availability Statement
All of the data that support the findings of this study are available in the main text or Supplementary Information.