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. 2025 Sep 4;262:145–160. doi: 10.3897/phytokeys.262.164617

Petrocosmea pengzhouensis (Gesneriaceae), a new species from Sichuan Province, China

Xinyi Zheng 1, Tianmeng Qu 1, Xinyu Chen 1, Yanru Zhang 1, Jin Wang 2, Li Li 3, Wen Wu 4, Yujie Wang 5, Bo Li 6, Ke Huang 1,✉, Zhixi Fu 1,✉
PMCID: PMC12426627  PMID: 40948883

Abstract

A new species of Gesneriaceae, Petrocosmea pengzhouensis K.Huang & Z.X.Fu, sp. nov., from the Feilaifeng Scenic Area of Longmen Mountain National Geological Park (Pengzhou, Chengdu City, Sichuan Province, China), is described and illustrated. Morphological and phylogenetic analyses provide robust evidence for its recognition as a new species. While morphologically most similar to P. duclouxii Craib, phylogenetic reconstructions reveal its closest affinities with P. duclouxii and P. intraglabra (W. T. Wang) Z. J. Qiu. The new species is morphologically distinguished from P. duclouxii by a suite of diagnostic characters: leaf shape (ovate to oblique-ovate vs. ovate to nearly orbicular), corolla throat (spotless vs. bearing two deep purple spots), and filament pubescence (translucent-puberulent vs. rust-brown puberulent). A distribution map and a comparative morphological table with P. duclouxii, P. intraglabra, and P. hexiensis S. Z. Zhang & Z. Y. Liu, and a preliminary conservation assessment following IUCN criteria are provided.

Key words: Morphological characters, new species, Petrocosmea , taxonomy

Introduction

Petrocosmea Oliv., a genus within the Gesneriaceae family and subfamily Didymocarpoideae (Weber et al. 2013; Qiu et al. 2020; Tang et al. 2022), has become an important indoor ornamental cultivar due to its emerald-green leaves, brightly colored flowers, and unique morphology (Kan et al. 2024). In 1985, Wang conducted a taxonomic revision of this genus, which was then considered to comprise 27 species and four varieties classified into three sections: sect. Anisochilus Hemsl., sect. Deinanthera W. T. Wang, and sect. Petrocosmea Oliv. (Wang 1985). In 2015, based on the first phylogenetic study, the genus was reclassified to include 33 species and three varieties distributed across five sections, with the addition of sect. Minor Zhi J. Qiu and sect. Barbata Zhi J. Qiu (Wang et al. 1990, 1998; Qiu et al. 2015; Tang et al. 2021).

To date, approximately 70 species of Petrocosmea have been reported worldwide (Li et al. 2023a; GRC 2025). Most Petrocosmea species exhibit narrow-range distributions (Wei and Wen 2009; Tang et al. 2021) and are primarily found in southern and southwestern China, which represent two major diversity centers of Gesneriaceae (Xu and Liu 2011; Tang et al. 2021). According to the Catalog of Life China 2025 edition, 11 Petrocosmea species are distributed in Sichuan Province, southwestern China (http://www.sp2000.org.cn/).

During field expeditions in Pengzhou, Chengdu City, Sichuan Province, in April 2025, we discovered living specimens of Petrocosmea (Gesneriaceae) growing on limestone cliffs within the Feilaifeng Scenic Area of Longmen Mountain National Geopark. Flowering specimens were collected for further study. These plants inhabit moist limestone cliff environments. Their rosette growth habit, cymes with solitary flowers, pale purple corollas, and anther dehiscence patterns show remarkable similarity to those of P. duclouxii, suggesting their possible taxonomic placement within sect. Petrocosmea. Based on preliminary morphological comparison and molecular evidence, we confirm that their comprehensive characteristics differ from those of previously reported taxa, thus supporting their recognition as a new species, which we formally describe here.

Material and methods

Sample collection and morphological analysis

In April 2025, living materials and voucher specimens of the new species were collected from limestone cliffs in Pengzhou, Chengdu City, Sichuan Province. The morphology of the new species was comprehensively compared with its closest relatives, P. duclouxii and P. intraglabra. Type specimens have been deposited in the herbarium of Sichuan Normal University (SCNU). The conservation status of the new species was assessed following the IUCN Red List Categories and Criteria (IUCN 2024).

DNA extraction and sequencing

Total genomic DNA was extracted using the modified CTAB method (Doyle and Doyle 1987). Paired-end DNA libraries were prepared following the Illumina DNA Library Construction Protocol (Allen et al. 2006) and sequenced on an Illumina HiSeq X Ten platform (San Diego, CA, USA). High-quality reads were assembled using GetOrganelle v1.7.2 (Jin et al. 2020) with default parameters. After annotation analysis using CPGAVAS2, the results were manually checked and adjusted in Geneious (Kearse et al. 2012; Shi et al. 2019). The gene map of the plastomes was generated using the Organellar Genome Draw (OGDraw) tool (https://chlorobox.mpimp-golm.mpg.de/OGDraw.html, accessed on 6 June 2025) (Greiner et al. 2019). Various plastid features, such as gene length and GC content, were analyzed using CPJSdraw (Li et al. 2023b). The complete chloroplast genome sequence and ITS (internal transcribed spacer) sequence are deposited in the NCBI GenBank database (http://www.ncbi.nlm.nih.gov/) with registration numbers PV750053 and PV819303 (P. pengzhouensis), respectively.

Phylogenetic analysis

Based on previous studies (Qiu et al. 2015), we obtained chloroplast DNA regions (rps16, trnT-L, etc.) and ITS sequences of 37 Petrocosmea species (including the newly discovered species) and two outgroup species (Raphiocarpus begoniifolius and R. petelotii) (Table 1). After comparing nodal support values and the topological structure of phylogenetic trees constructed separately or in combination with different chloroplast DNA fragments and ITS sequences, rps16, trnT-L, and ITS sequences were selected as the optimal molecular combinations in this study. Sequence alignments for each of the three selected markers were performed using MAFFT v7.520 (Katoh and Standley 2013) with default parameters. The aligned sequences were then trimmed with the “trimAL Wrapper” in TBtools v2.322 to remove poorly aligned regions. The processed sequences were concatenated using SequenceMatrix 1.10 to generate a combined dataset for phylogenetic reconstruction.

Table 1.

Species and GenBank accession number for taxa included for phylogenetic reconstruction in this study.

Taxon rps16 trnT-L ITS
Ingroups Petrocosmea pengzhouensis PV750053 PV750053 PV819303
P. barbata Craib KR006491 KR006424 KR006475
P. begoniifolia C. Y. Wu & H. W. Li KR006494 KR006431 KR006482
P. cavaleriei H. Lévl. KR006487 KR006420 KR006476
P. coerulea C. Y. Wu & W. T. Wang KR006492 KR006427 KR006483
P. duclouxii Craib KR006498 KR006433 KR006478
P. flaccida Craib KR006517 KR006414 KR006471
P. forrestii Craib KR006520 KR006416 KR006464
P. glabristoma Z. J. Qiu & Y. Z. Wang. KR006516 KR006417 KR006468
P. grandiflora Hemsl. KR006504 KR006437 KR006467
P. grandifolia W. T. Wang KR006507 JN092505 JN092439
P. hexiensis S. Z. Zhang & Z. Y. Liu KR006497 KR006415 KR006469
P. huangjiangensis Yan Liu & W. B. Xu KR006503 KR006435 KR006484
P. iodioides Hemsl. KR006513 JN092506 JN092440
P. kerrii Craib KR006509 JN092507 JN092441
P. kerrii Craib var. crinita W. T. Wang KR006505 KR006425 KR006462
P. longianthera Z. J. Qiu & Y. Z. Wang KR006514 JN092481 JN092398
P. longipedicellata W.T.Wang KR006490 KR006422 KR006474
P. mairei H. Lévl. KR006519 KR006418 KR006465
P. intraglabra (W. T. Wang) Z. J. Qiu KR006495 KR006430 KR006479
P. martinii H. Lévl. KR006501 JN092508 JN092442
P. martinii var. leiandra W. T. Wang KR006512 JN092509 JN092443
P. melanophthalma Huan C. Wang, Z. R. He & Li Bing Zhang KR006493 KR006436 KR006481
P. menglianensis H. W. Li KR006506 JN092510 JN092444
P. minor Hemsl. Hook. KR006515 JN092511 JN092445
P. nanchuanensis sp. nov. KR006489 KR006423 KR006472
P. nervosa Craib KR006523 JN092512 JN092446
P. oblata Craib KR006518 JN092513 GU350661
P. parryorum C. E. C. Fisch. KR006508 KR006426 KR006463
P. qinlingensis W. T. Wang KR006521 KR006419 KR006470
P. rosettifolia C. Y. Wu & H. W. Li KR006510 KR006428 KR006473
P. sericea C. Y. Wu & H. W. Li KR006500 JN092500 JN092434
P. shilinensis Y. M. Shui & H. T. Zhao KR006511 KR006434 KR006486
P. sinensis Oliver KR006522 JN092514 GU350660
P. xanthomaculata G. Q. Gou & X. Y. Wang KR006488 KR006421 KR006477
P. xingyiensis Y. G. Wei & F. Wen KR006502 KR006432 KR006485
P. yanshanensis Z. J. Qiu & Y. Z. Wang KR006499 JN092495 JN092429
P. barbata Craib KR006491 KR006424 KR006475
Outgroups Raphiocarpus begoniifolius (Lévl) Burtt KR006524 JN092515 GU350648
R. petelotii (Pellegr) Burtt KR006525 JN092516 JN092447
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We performed comprehensive phylogenetic analyses using both maximum likelihood (ML) and Bayesian inference (BI) approaches. For the ML analysis, model selection was conducted using ModelFinder (Kalyaanamoorthy et al. 2017), which identified K3P+G4 as the optimal substitution model based on the Bayesian Information Criterion (BIC). This model incorporated unequal base frequencies (+F), the K3P substitution matrix, and among-site rate variation (+G4). Phylogenetic reconstruction was performed in IQ-TREE v2.3.6 (Minh et al. 2020), with branch support assessed via 1000 ultrafast bootstrap replicates (UFBoot ≥95% considered strong support; Hoang et al. 2018). For the BI analysis, the trimmed ITS sequence alignment was analyzed using MrBayes v3.2.7 (Ronquist and Huelsenbeck 2003). The analysis consisted of four independent MCMC runs of 10 million generations each, sampling every 1000 generations. After discarding the first 25% of samples as burn-in, posterior probabilities were calculated from the remaining trees to generate a majority-rule consensus tree. All phylogenetic trees were visualized and annotated using FigTree v1.4.4.

Taxonomic treatment

. Petrocosmea pengzhouensis

K.Huang & Z.X.Fu sp. nov.

A14A3E45-0B83-5E88-B2B5-F422EC170A56

urn:lsid:ipni.org:names:77368635-1

Type.

China • Sichuan Province, Chengdu City, Pengzhou, Feilaifeng Scenic Area of Sichuan Longmen Mountain National Geological Park, 31.31.193542, 103.917283, ca. 1116 m, 30 April 2025, Ke Huang & Zhixi Fu 8560 (holotype: SCNU!; isotype: SCNU!) (Figs 1–4).

Figure 1.

Figure 1.

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Petrocosmea pengzhouensis sp. nov. A, B. Habitat; C, E. Plant; D, F. Plant and roots (Photos A, B by KH and C–F by XZ).

Figure 4.

Figure 4.

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Close-up image of Petrocosmea pengzhouensis K.Huang & Z.X.Fu, sp. nov.

Figure 2.

Figure 2.

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Petrocosmea pengzhouensis sp. nov. A. Leaves; B, D. Flowers; C. Dissected corollas; E, F. Stamens; G. Pistil and stamens; H. Sepals (Photos by XZ).

Figure 3.

Figure 3.

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Holotype image of Petrocosmea pengzhouensis K.Huang & Z.X.Fu, sp. nov.

Diagnosis.

In terms of traits such as being a rosette-forming herb, possessing a single-flowered cyme, and having a pale purple corolla, P. pengzhouensis most closely resembles P. duclouxii but differs primarily in the following characteristics: leaf shape (ovate to oblique-ovate vs. ovate to nearly orbicular), corolla throat (spotless vs. bearing two deep purple spots), and filament pubescence (translucent-puberulent vs. rust-brown puberulent) (Table 2).

Table 2.

Morphological character comparison amongst Petrocosmea pengzhouensis, P. hexiensis, P. duclouxii, and P. intraglabra.

Character P. pengzhouensis P. duclouxii P. hexiensis P. intraglabra
Leaf shape Ovate to oblique–ovate Ovate to nearly orbicular Ovate–rhombic to rhombic Ovate to nearly orbicular or elliptic
Leaf base Cordate (sometimes oblique) Cordate Truncate Slightly oblique
Leaf number 13–26 5–25 20–60 7–20
Leaf blade indumentum Adaxial: sparsely pilose; abaxial: densely appressed–pubescent Both surfaces sparsely white–pilose Both surfaces densely appressed–pubescent Adaxial: pilose, abaxial: densely puberulent
Leaf margin Repand–crenate Entire or obscurely repand–crenate Repand–crenate Entire near base, crenate-dentate toward apex, apex acute to rounded
Leaf vein 3–4 per side, slightly convex or flat on abaxial side 2–3 per side, slightly convex or flat on abaxial side 3 per side, inconspicuous 2–3 per side, conspicuous
Bracts Absent Present Present Present
Cyme 1–5 cymes 1–7 cymes 6–15 cymes 1–5 cymes
The degree of upper lip cracking Near the middle Near base Near base Near base
Shape of upper lip lobe Ovate Semi–circular Ovate Narrowly ovate
The degree of lower lip cracking Near the middle Near the middle Near the middle Near the middle
Shape of lower lip lobe Long–ovate Round–ovate Ovate Obovate and the top is round
Corolla Tube 4.3–6.4 mm long, throat blue-purple, unspotted Tube 3–5.2 mm long, throat with 2 deep purple spots Tube 4.5–5.5 mm long, throat with 2 deep purple spots Tube 3–4 mm long, throat white, unspotted
Stamen Filaments densely translucent-puberulent Filaments rust–brown puberulent Filaments sparsely pubescent Filaments rust-brown puberulent
Pistil Pistil 4.3–5.4 mm, style pubescent below middle upper part Pistil ca. 6 mm, style sparsely puberulent below middle Pistil ca. 7 mm, style sparsely pubescent below middle Pistil 5.2–6 mm, style pilose near base
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Description.

Perennial rosette-forming herbs. Roots fibrous, clustered. Leaves herbaceous, 13–26, basal, densely rosulate; inner leaves subsessile or with petioles to 2.2 cm long in outer leaves, densely pubescent and sparsely villous; leaf blade 0.4–1.8 cm long, 0.3–1.2 cm wide, ovate to oblique-ovate in outer leaves, apex obtuse to rounded, base cordate (sometimes oblique), margin crenate, adaxially sparsely villous, abaxially densely pubescent or sparsely villous; lateral veins 3–4 per side, slightly raised or flat abaxially. Cymes 1–5, each 1-flowered; peduncle 0.5–4.3 cm long, densely pubescent or sparsely villous. Bracts absent. Calyx zygomorphic, 5-parted to base; lobes narrowly lanceolate, 1.9–2.7 mm long, externally pubescent. Flowers pale purple, 8.7–13.8 mm long; corolla externally densely puberulent, internally glabrous; tube 4.3–6.4 mm long, throat 4–5 mm in diameter, base 2.5–3 mm in diameter, subcampanulate, throat blue-purple, unspotted; adaxial lip conspicuously shorter, 8–9 mm long, 2-lobed to the middle, lobes ovate, 2–2.3 mm long, 2–2.5 mm wide, slightly reflexed; abaxial lip 11–13.8 mm long, 3-lobed to the middle, lobes long-ovate, 2.6–3.0 mm long, 3.6–4.2 mm wide, slightly reflexed. Stamens 2, ca. 2.5 mm long; filaments adnate, ca. 1 mm above corolla base, 1.8–2.3 mm long, 0.3–0.5 mm in diameter, densely translucent-puberulent; anthers ovate, dehiscing longitudinally, ca. 1 mm long, 1.0–1.1 mm wide; thecae 2, parallel. Pistil 4.3–5.4 mm long; ovary densely villous, ovoid, ca. 1.3 mm long, 1.2–1.5 mm in diameter; style 3–4.1 mm long, 0.3 mm in diameter, sparsely pubescent below middle-upper portion, slightly geniculate (ca. 45°) above base; stigma capitate, ca. 0.4 mm in diameter. Capsule straight relative to pedicel, 2.9–3.5 mm long, 1.2–1.3 mm in diameter, oblong to ovoid, dehiscing loculicidally to base.

Phenology.

Flowering from April to May, fruiting in the wild is unknown; only capsules of the previous year were observed.

Etymology.

The new taxon is named after the type locality, Pengzhou, Chengdu City, China.

Vernacular name.

The Chinese name is “péng zhōu shí hú dié” (彭州石蝴蝶).

Distribution and habitat.

To date, only a single population has been found, exclusively in the Feilaifeng Scenic Area of the Longmen Mountain National Geological Park in Pengzhou, Chengdu City, Sichuan Province, China. Petrocosmea pengzhouensis was found growing on rocks in limestone caves at an elevation of ca. 1116 m (Fig. 5).

Figure 5.

Figure 5.

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Distribution of Petrocosmea pengzhouensis and P. duclouxii in Sichuan Province, China (red star indicates P. pengzhouensis, red dots indicate P. duclouxii).

Conservation status and IUCN Red List category.

In the type locality, a population of approximately 100 mature individuals was discovered. This habitat is located within the local Feilaifeng Scenic Area, and it is highly likely that additional populations exist in the region. Before further investigations, this species should be assessed as “Data Deficient” (DD) according to the IUCN standards (IUCN 2024).

Discussion

Morphologically, we observed that specimens collected from moist limestone cliffs in Pengzhou exhibited high similarity to Petrocosmea duclouxii, P. hexiensis, and P. intraglabra, particularly P. duclouxii. A detailed morphological comparison is provided in Table 2. Petrocosmea duclouxii and P. pengzhouensis are perennial rosette herbs with pubescent leaves on both surfaces and cymose inflorescences bearing a single flower, along with pale purple corollas. However, the newly discovered population also displayed distinct differences from the latter species. P. duclouxii is characterized by ovate to nearly orbicular leaves, two deep purple spots at the throat, and rust-brown puberulent filaments (Qiu et al. 2012). The newly discovered species from Pengzhou has ovate to oblique-ovate leaves, a spotless throat, and translucent-puberulent filaments. According to the Plant Photo Bank of China (PPBC, http://ppbc.iplant.cn/list21?latin=Petrocosmea%20duclouxii&pro=%E5%9B%9B%E5%B7%9D&sel=like), the Chinese Virtual Herbarium (CVH, https://www.cvh.ac.cn/spms/list.php?&taxonName=Petrocosmea%20duclouxii), and literature records (Li and Wang 2005; Qiu et al. 2012), the distribution of P. duclouxii specimens in Sichuan Province is primarily concentrated in Huidong County of the Liangshan Yi Autonomous Prefecture, as well as Mount Emei, Leshan Giant Buddha, and Longchi Sixigou in Leshan City. These locations exhibit significant geographical isolation from Pengzhou City.

In this study, the ML tree (Fig. 6) revealed that the new species formed a highly supported clade (BS = 100) with P. intraglabra and P. duclouxii, and this clade was sister to P. melanophthalma (BS = 90). Similarly, in the BI tree (Fig. 7), the same three species clustered together (PP = 1) and were adjacent to P. melanophthalma (PP = 0.99). This consistent topological structure is supported by previous research findings (Qiu et al. 2015), confirming that both P. intraglabra and P. duclouxii are closely related to P. pengzhouensis.

Figure 6.

Figure 6.

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Phylogenetic reconstruction of 37 Petrocosmea species based on chloroplast regions (rps16 and trnT-L) and ITS sequences. Petrocosmea pengzhouensis is highlighted in red. Numbers at branches represent ML bootstrap (ML-BS) values; “-” indicates nodes with support values <70.

Figure 7.

Figure 7.

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Phylogenetic reconstruction of 37 Petrocosmea species based on ITS sequences. The phylogenetic position of Petrocosmea pengzhouensis is marked in red font. The numbers on the branches represent Bayesian posterior probabilities (PP) values.

Phylogenetic analysis also revealed a distant relationship between P. pengzhouensis and P. duclouxii, supported by distinct chloroplast gene counts and genomic features (Fig. 8, Table 3). The complete chloroplast genome of P. pengzhouensis (153,088 bp; total GC content: 35.57%) comprises an 82,885 bp large single-copy (LSC) region (GC: 35.57%), an 18,181 bp small single-copy (SSC) region (GC: 30.65%), and two 24,639 bp inverted repeat (IR) regions (GC: 43.01%). In comparison, P. duclouxii exhibits a slightly larger chloroplast genome (153,320 bp; total GC content: 37.60%) with an 84,322 bp LSC (GC: 35.60%), an 18,312 bp SSC (GC: 31.21%), and two 25,343 bp IRs (GC: 43.25%). Notably, both species display elevated GC content in their IR regions relative to single-copy regions, with P. duclouxii possessing marginally expanded IRs.

Figure 8.

Figure 8.

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Gene map of the plastomes of Petrocosmea pengzhouensis.

Table 3.

Characteristics of complete chloroplast genomes of P. pengzhouensis and P. duclouxii.

Speciese Genome size·(bp) LSC size (bp) IRa/Rb size·(bp) SSC Size (bp) Total. GC Content (%) GC Content in·LSC (%) GC Content in·IRa/IRb (%) GC Content in·SSC (%)
Petrocosmea pengzhouensis 153, 088 82, 885 24, 639 18, 181 37.61 35.57 43.01 30.65
Petrocosmea duclouxii 153, 320 84, 322 25, 343 18, 312 37.60 35.60 43.25 31.21
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In summary, morphologically, P. pengzhouensis closely resembles P. duclouxii, P. hexiensis, and P. intraglabra, particularly P. duclouxii, yet distinct differences remain. Molecular evidence supports P. pengzhouensis as a close relative of P. duclouxii and P. intraglabra. These findings confirm that P. pengzhouensis represents a new species within the genus Petrocosmea. Further molecular and morphological studies can help clarify the taxonomic status and evolutionary relationships of this species.

Supplementary Material

XML Treatment for Petrocosmea pengzhouensis

Acknowledgments

We would like to thank the anonymous reviewers for their valuable comments and suggestions.

Citation

Zheng X, Qu T, Chen X, Zhang Y, Wang J, Li L, Wu W, Wang Y, Li B, Huang K, Fu Z (2025) Petrocosmea pengzhouensis (Gesneriaceae), a new species from Sichuan Province, China. PhytoKeys 262: 145–160. https://doi.org/10.3897/phytokeys.262.164617

Funding Statement

This study was financially supported by the National Natural Science Foundation of China (No. 32000158), the National Science & Technology Fundamental Resources Investigation Program of China (No. 2021XJKK0702), the Foundation of Sustainable Development Research Center of Resources and Environment of Western Sichuan, Sichuan Normal University (No. 2020CXZYHJZX03), the Key Laboratory of Chemistry in Ethnic Medicinal Resources (Yunnan Minzu University), and State Ethnic Affairs Commission & Ministry of Education (No. MZY2301).

Contributor Information

Ke Huang, Email: 542176689@qq.com.

Zhixi Fu, Email: fuzx2017@sicnu.edu.cn.

Additional information

Conflict of interest

The authors have declared that no competing interests exist.

Ethical statement

No ethical statement was reported.

Use of AI

No use of AI was reported.

Funding

This study was financially supported by the National Natural Science Foundation of China (No. 32000158), the National Science & Technology Fundamental Resources Investigation Program of China (No. 2021XJKK0702), the Foundation of Sustainable Development Research Center of Resources and Environment of Western Sichuan, Sichuan Normal University (No. 2020CXZYHJZX03), the Key Laboratory of Chemistry in Ethnic Medicinal Resources (Yunnan Minzu University), and the State Ethnic Affairs Commission & Ministry of Education (No. MZY2301).

Author contributions

XZ: Conceptualization, writing – original draft; TQ: software, formal analysis; XC: formal analysis, data curation; YZ: data curation; JW: data curation; LL: data curation; WW: data curation; YW: data curation; BL: data curation; KH: resources, writing – review and editing; ZF: conceptualization, writing – review and editing. All authors have read and approved the final manuscript.

Author ORCIDs

Xinyi Zheng https://orcid.org/0009-0003-2300-5525

Tianmeng Qu https://orcid.org/0009-0008-0185-5306

Xinyu Chen https://orcid.org/0009-0008-6347-2490

Yanru Zhang https://orcid.org/0009-0009-0450-2001

Yujie Wang https://orcid.org/0009-0004-0732-2827

Bo Li https://orcid.org/0000-0002-5131-8639

Ke Huang https://orcid.org/0009-0006-2057-5452

Zhixi Fu https://orcid.org/0000-0002-2789-6287

Data availability

All of the data that support the findings of this study are available in the main text.

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Associated Data

This section collects any data citations, data availability statements, or supplementary materials included in this article.

Supplementary Materials

XML Treatment for Petrocosmea pengzhouensis
phytokeys.262.164617-treatment1.xml (22KB, xml)

Data Availability Statement

All of the data that support the findings of this study are available in the main text.

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