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[Preprint]. 2025 Dec 25:2024.08.12.605506. [Version 2] doi: 10.1101/2024.08.12.605506

Circulating extracellular vesicle microRNAs mediate immune modulation of social behavior in mice

Ken Matoba, Eisuke Dohi, Phoebe A Garcia, Jose Francis-Oliveira, Mirmohammadali Mirramezanializamini, Inssaf Berkiks, Frida Anguiano, Jana H Badrani, Oluwaseun Fatoba, Eric Y Choi, Julia See, Md Sorwer Alam Parvez, Takahiro Kochi, Norimichi Ito, Rei Mitani, Indigo VL Rose, Takashi Imai, David K Crossman, Mikhail V Pletnikov, Kenneth W Witwer, Minae Niwa, Shin-ichi Kano
PMCID: PMC12767323  PMID: 41497581

Abstract

Extracellular vesicles (EVs) are cell-derived small membrane vesicles and circulate throughout the body, but the impact of circulating EVs on brain function and behavior remains elusive. Here, we report that wild-type (WT) mouse blood, particularly EVs, increases sociability in socially impaired immunodeficient Rag1 -/- mice, mimicking the effects of WT T cell transfer. These EVs localized to neurons and regulated PKCε expression, GABA A receptor synaptic localization, and inhibitory postsynaptic signaling in prefrontal cortex (PFC) pyramidal neurons. Injection of Rag1 -/- EVs supplemented with miR-23a-3p and miR-103-3p enhanced synaptic function and sociability in Rag1 -/- mice. T cells secreted miR-23a-3p via EVs, and Mir23a -/- T cells failed to increase sociability. Similar beneficial effects of WT blood EVs were observed in additional mouse models with sociability deficits, Cntnap2 -/- and Shank3 -/- mice. These findings uncover a previously unrecognized role of EV miRNAs in mediating immune modulation of synaptic function and social behavior, revealing a novel molecular pathway for immune-neuron communication.

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