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Serum antibody titres to β-gal 28 days after intramuscular injection of fish with different formulations. Antibody titres to β-gal were measured by ELISA as previously been described.7 The titre was the last dilution to give a binding twice the conjugate control, i.e. > 0·336.15 *VR1223 was empty (see text). Values in parenthesis represent geometric mean titres.
