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The first paragraph of the Acknowledgements for this paper should have read as follows: We thank the Arthritis Research Campaign for their support, ARC grant reference No. 17552, and also acknowledge the use of data from the Wellcome Trust Case Control Consortium. We are also grateful to the NIHR Manchester Biomedical Research Centre for support and to the NIHR Oxford Comprehensive Biomedical Research Centre for salary support for P.H.