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Proposed CrataBL binding GAG motif: eight repeating disaccharide units having two sulfo groups on a hexosamine residue linked to an uronic acid having a carboxyl group. Top: heparin dp 16 Middle: Dis-DS dp 16; Bottom: CS-E dp 16. Sugar symbols: Blue square, GlcNAc; Tan lower segment diamond, IdoA; Yellow square, GalNAc; Blue upper segment diamond, GlcA.
