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. 2014 Mar 14;9(3):e91012. doi: 10.1371/journal.pone.0091012

Role of Investment Heterogeneity in the Cooperation on Spatial Public Goods Game

Wu-Jie Yuan 1, Cheng-Yi Xia 2,*
Editor: Matjaž Perc3
PMCID: PMC3954582  PMID: 24632779

Abstract

Public cooperation plays a significant role in the survival and maintenance of biological species, to elucidate its origin thus becomes an interesting question from various disciplines. Through long-term development, the public goods game has proven to be a useful tool, where cooperator making contribution can beat again the free-rides. Differentiating from the traditional homogeneous investment, individual trend of making contribution is more likely affected by the investment level of his neighborhood. Based on this fact, we here investigate the impact of heterogeneous investment on public cooperation, where the investment sum is mapped to the proportion of cooperators determined by parameter Inline graphic. Interestingly, we find, irrespective of interaction networks, that the increment of Inline graphic (increment of heterogeneous investment) is beneficial for promoting cooperation and even guarantees the complete cooperation dominance under weak replication factor. While this promotion effect can be attributed to the formation of more robust cooperator clusters and shortening END period. Moreover, we find that this simple mechanism can change the potential interaction network, which results in the change of phase diagrams. We hope that our work may shed light on the understanding of the cooperative behavior in other social dilemmas.

Introduction

Understanding the sustenance and emergence of cooperation within groups of egoistic agents represents one of long-standing puzzles across various disciplines [1], [2]. A theoretical framework that sheds the light into this open question is the evolutionary game theory [3], [4]. Although the prisoner's dilemma game (PDG) and snowdrift game (SDG), as the metaphors, are often chosen to characterize the social dilemma through the pairwise interactions [5][19], the coordination and management of many common resources, such as environment problems, climate change, public traffic and financial markets, needs to be implemented and solved by multiple sides or group interactions, which is best described by the so-called public goods game (PGG) [20][29]. In the PGG model [30], any participant simultaneously decides to contribute (i.e., cooperate) or not contribute (i.e., defect) to the public pool. After that, all contributions of cooperators are summed and multiplied by a synergy factor Inline graphic, and then uniformly distributed among all participants regardless of their contributions. Nevertheless, in this situation, there is a free-ride problem [31]: why an individual makes contribution if he can reap the benefits without bearing any cost, and why a agent chooses cooperation if he can obtain higher payoff by defecting. Hence, the game theory often predicts a very pessimistic conclusion: all participants take the defective strategy and it eventually leads to the “tragedy of commons” whereby nobody contributes [32].

However, to our surprises, the above-mentioned prediction is different from many experimental findings in the literature [33]. In fact, from the perspectives of the whole group or population, the total cooperation is obviously optimal since it creates the greater collective benefits. Thus, several mechanisms have been put forward to illustrate the evolution of cooperation on the public goods games. For example, a viable mechanism to facilitate the public cooperation is to punish the non-cooperating participants [34][37], but it can also evoke the so-called second-order free rider problem in which the investment of publishers may be exploited by cooperators [38]. Meanwhile, volunteering or voluntary participation [39], reputation mechanism [40] and other theoretical supplements [20] are also proposed as effective means to promote the cooperation on spatial public goods games. In addition, the heterogeneous topology has been ubiquitously found in many natural, social, biological and engineering systems, and this structural heterogeneity creates the social diversity within the system which may greatly promote the collective cooperation [41].

Except for the well-mixing assumption, in reality the spatial PGG also draws a lot of attention of statistical physicists who focused on the phase transition behaviors in the evolution of cooperation [20]. The results indicate that the pattern formation or agglomeration may be responsible for the public cooperation on the spatial PGG [42]. Interestingly, the nonhomogeneous teaching activities play an important role in the promotion of cooperation for the spatial public goods games [43]. Importantly, noise taking place in the strategy adoption can noticeably influence the cooperative behaviors and even substantially change the outlay of cooperators' density in dependence on the noise intensity [44][50]. Qualitatively similar findings are also discovered in the impact of diversity on the cooperation of PGGs [51]. In particular, Szolnoki et al [52] make use of two different types of regular graph, square and honeycomb lattice, to explore the impact of interaction topology and group size on the cooperation in spatial PGG. It is shown that increasing group size leads to indirect linkage or interaction among players participating in the same PGG group, further changes the effective interaction topology and substantially alters the cooperation behaviors on regular lattice without any overlapping structures. However, in most works, the contribution of a cooperator to all PGG groups is still assumed to be identical, which is far from many realistic situations: heterogeneous activity [20]. Inspired by this fact, an interesting question takes place: if we allow cooperators hold heterogeneous investment, how does cooperation varies?

Aiming to answer this question, in this work we consider a spatial PGG model, where the cooperator's contribution will be non-uniformly distributed over all PGG groups it involves. While its contribution to each group can be mapped to a function of proportion of cooperations tuned by the parameter Inline graphic. Large-scale numerical simulations demonstrate that the non-homogenous distribution of cooperator's contribution will largely promote the evolution of cooperation. This trait becomes more obvious with the increment of Inline graphic, irrespective of interaction networks. Moreover, we also find that this mechanism changes the potential interaction networks.

Methods

Here, we consider the spatial PGG: all participants are placed on a regular interaction graph which is a square lattice with Von Neumann neighborhood as described in Ref.[52]. Each player will participate in Inline graphic PGG groups where one PGG group is centered around the focal player and other Inline graphic ones are correspondingly centered around Inline graphic nearest neighbors. Among them, Inline graphic is the size of neighborhood, coordination number or number of all nearest neighbors, and Inline graphic.

Initially, each participant or player on site Inline graphic is equally probably designated as a contributor (cooperator, Inline graphic) or a non-contributor (defector, Inline graphic). Then, each player will participate in Inline graphic PGG groups and collect its payoff Inline graphic by accumulating the respective share of each PGG group to which it belongs. In each PGG group, without loss of generality, the contribution of a defector is always fixed to be Inline graphic while the contribution of a cooperator is not Inline graphic, which is the general assumption in many previous reports [34][37], [53]. Here, we hypothesize that the contribution or investment of a cooperator into a PGG group in which it participates depends on the fraction of cooperators inside that group, and this investment quantity Inline graphic can be depicted as follows,

graphic file with name pone.0091012.e019.jpg (1)

where Inline graphic stands for the investment of a cooperator Inline graphic into the Inline graphic centered PGG group, Inline graphic and Inline graphic represent the number of cooperators and all players inside Inline graphic-centered PGG group. Inline graphic is a tunable parameter which characterizes, to some extent, the heterogeneity of distribution of a cooperator's contribution. Noticeably, Inline graphic leads to the traditional PGG model [34][37], [53], and Inline graphic ensures that the investment will be beneficial to the PGG group with higher cooperation level. After that, the investment will be summarized from all cooperators inside Inline graphic-centered PGG group and multiplied by an enhancement factor Inline graphic. At last, the amplified investment will be evenly allocated among all players inside that group. Thus, the net payoff of player Inline graphic from the Inline graphic-centered PGG group will be equal to

graphic file with name pone.0091012.e033.jpg (2)

where Inline graphic denotes the set of player Inline graphic and its Inline graphic nearest neighbors. Subsequently, the total payoff Inline graphic of player Inline graphic can be calculated as follows,

graphic file with name pone.0091012.e039.jpg (3)

Since the system evolves according to the Monte Carlo Simulation (MCS) procedure, the player's strategy will be updated by a specific rule after acquiring the total payoff. Here, within one elementary MCS time step, we will randomly choose a player Inline graphic and one of its nearest neighbors Inline graphic, and calculate their total payoffs Inline graphic and Inline graphic based on the above-described order. Then, player Inline graphic will update its strategy Inline graphic by adopting the strategy Inline graphic of player Inline graphic with the following Fermi probability,

graphic file with name pone.0091012.e048.jpg (4)

where Inline graphic characterizes the uncertainty during the strategy imitation [44], [45]. For Inline graphic, the player Inline graphic will deterministically take the strategy of player Inline graphic if Inline graphic. While for Inline graphic, player Inline graphic can also imitate the strategy of player Inline graphic even if player Inline graphic performs worse as far as the total payoff is concerned, which can mimic the error or non-rationality during the course of decision making to a certain extent. In addition, the strategy is asynchronously updated during the system evolution, that is, the strategy of each player can only have a chance to be updated on average within one full MCS time step.

Results

We start by presenting how the heterogeneous investment affects the evolution of cooperation on spatial cooperations. Figure 1 depicts the dependence of Inline graphic on the normalized enhancement factor Inline graphic for different tunable parameters Inline graphic. It is obvious, compared to the standard case (Inline graphic), that the evolution of cooperation is greatly promoted when Inline graphic. We can observe the emergence of cooperation from two sides: on one hand, the transition point Inline graphic of normalized synergy factor (Inline graphic) between the full defection (Inline graphic) and mixing state of cooperators and defectors (Inline graphic) is largely reduced, and it means that the players easily transcend the barrier of defection and are prone to cooperate under the low cooperation cost; On the other hand, the critical point Inline graphic from the mixing state (Inline graphic) into the full cooperation (Inline graphic) has also become smaller than that in the traditional case. In particular, the larger Inline graphic, the smaller the critical point. The two critical points are nearly same when Inline graphic and it means the transition from the full defection into the full cooperation is almost abrupt. In addition, according to Eq.(1), our PGG model with a tunable parameter Inline graphic is reduced into the standard case in which each cooperator will make a constant investment quantity, for example, to be Inline graphic in Ref.[52]. Meanwhile, our simulation results with Inline graphic are totally identical with those in [52]. Thus, we can conclude that the investment heterogeneity of a cooperator into different PGG groups are responsible for the promotion of cooperation.

Figure 1. Fraction of cooperators Inline graphic as a function of normalized enhancement factor Inline graphic on square lattice with Von Neumann neighborhood.

Figure 1

Here, Inline graphic, Inline graphic and other linear sizes Inline graphic are also verified not to change the qualitative results of our PGG model. Among them, the fraction of cooperators Inline graphic within the whole population is averaged over Inline graphic MCS time step after discarding the transient steps up to Inline graphic MCS.

To further illustrate the origin favoring the cooperation in our PGG model, Figure 2 depicts the evolution of characteristic snapshots for three different tunable parameters Inline graphic under the prepared initial state. Here each row of patterns correspond to a different Inline graphic, and four columns denote the pattern at Inline graphic, respectively. Under these three cases, the initial distribution of cooperators and defectors are totally same, in which all cooperators are deployed in the middle row of the lattice and defectors are placed on two sides of the lattice. However, the evolutionary pattern exhibits the distinct features for different Inline graphic after enough time steps (Inline graphic). In the standard case (Inline graphic), the investment will be shared in all PGG groups and the free-ride from the defectors thrives so that the cooperators cannot resist the explorations of defectors and tend to be extinct in the end. While for Inline graphic, the cooperator's investment will be heterogeneously allocated among Inline graphic PGG groups, the PGG group with higher cooperation level will obtain the more investment and then this effect will further encourage more players to take the cooperative strategy within this group. The larger Inline graphic, the stronger this effect, hence the cooperators have enough power to create the cooperative (Inline graphic) clusters to inhibit the invasion of defectors. Moreover, the fraction of cooperators in the population will become higher and higher as Inline graphic increases.

Figure 2. Characteristic snapshots of cooperators (red) and defectors (blue) under the prepared initial state for different times steps.

Figure 2

From top row to bottom panel, the tunable parameter Inline graphic is set to be 0, 0.5 and 1.0, respectively. In all dynamical patterns, the synergy factor Inline graphic is 0.6, lattice size Inline graphic is 200 and strategy adoption uncertainty Inline graphic is 0.1.

Moreover, it is also interesting to examine the time course of cooperation under this novel mechanism. According to the recent investigate prediction [19], [54], [55]: a typical evolution process can be divided into two evident periods: the enduring (END) period and the expanding (EXP) period. The shorter the END period (the more perfect the formation of cooperator cluster), the higher the final cooperation level. We expect that under the present mechanism the increment of Inline graphic is beneficial for END period and cooperators can converge into more robust clusters. Figure 3 shows the results obtained for Inline graphic under different Inline graphic value. What we first glance at is the evolution trend of Inline graphic (namely, the traditional version), there exists only END period and cooperators die out. However, with the increment of Inline graphic, the tide changes: END periods becomes shorter and shorter (the end of END period is marked by arrows), and at the same time the remainder cooperators form clusters and can expand in the subsequent EXP period. Obviously, when Inline graphic is sufficient large (Inline graphic), the initial invasion of defection becomes very difficult, hence END periods vanishes and cooperators directly organize clusters. Along this line, the cooperator clusters can expand effectively and reach full cooperation state faster. To further support this point, we also feature the corresponding snapshots under the same time steps. It is obvious that the lager the value of Inline graphic, the larger the reminder cooperator cluster at the end of END period, which finally leads to larger clusters. This observation validates our guess: heterogeneous investment can change the evolution trend (i.e., accelerate the END period) and promotes the formation of cooperator clusters, which is the basic guarantee for the maintenance of cooperation.

Figure 3. Top panel: time courses depicting the evolution of cooperation for different values of Inline graphic under the random distribution of strategies.

Figure 3

Note that the arrows denote the end of enduring (END) period and the beginning of expanding (EXP) period. Bottom panel: the evolution snapshots of different values of Inline graphic. From top to bottom, the values of Inline graphic are 0, 0.5, 1.0 and 2.0, respectively. The colore code is the same with Fig.2: cooperator (red) and defector (blue). From left to right, the time steps are 0, 10, 100, 1000 for each value of Inline graphic.

The robustness of cooperation against the noise can be discussed through the full Inline graphic phase diagram in Fig. 4. Here we depict the relationship between critical synergy factor Inline graphic and noise strength Inline graphic which are both normalized by the group size (Inline graphic). It is observed that the phase boundaries display the distinct behaviors for three different Inline graphic. In the traditional case (i.e., Inline graphic), the boundary between the full defection Inline graphic and mixed phase Inline graphic (i.e., lower boundary Inline graphic) will monotonically decrease as Inline graphic, but the boundary separating the mixed phase Inline graphic from full cooperation Inline graphic (i.e., upper boundary Inline graphic) takes on a bell-shaped outlay which means that there is an optimal noise strength ensuring the coexistence of cooperators and defectors. As Inline graphic, the lower boundary is nearly similar to the traditional one and only the critical value is much smaller, which means the cooperator is easier to sustain in the sea of defectors. However, the upper boundary will continuously increase in the limit of strong selection (i.e. Inline graphic), and introducing the noise will benefit the emergence of cooperators. In particular, the upper and lower boundaries will almost collapse into the same curve under the weak selection limit (i.e. Inline graphic), which means that it is impossible for the cooperators and defectors to coexist within the same population. Thus, the heterogeneous investment will lead to the phase transitions which differ from the traditional case, and is responsible for the promotion of cooperation in the spatial PGG model. Moreover, we can observe that optimal cooperation dies out with the increment of Inline graphic, while the actual topology network does not have any change. Similar to the previous literatures [45], [46], we deduce that this may caused by the fact that the topology interaction network has changed. The square lattice obviously lacks overlapping triangles and thus enables the observation of an optimal Inline graphic, trimming the likelihood of who will act as a investor seems to effectively enhance linkage among essentially disconnected triplets and thus precludes the same observation.

Figure 4. Full normalized Inline graphic phase diagrams for different tunable parameters Inline graphic.

Figure 4

From top to bottom, panels (a), (b), (c) correspond to Inline graphic, respectively. The lattice size is Inline graphic and PGG group size is fixed to be Inline graphic.

Finally, it is still of interest to examining the universality of this simple mechanism under different topologies. To answer this question, in Fig. 5 we consider one type of triangular lattice depicted as in Ref.[52], in which the coordination number Inline graphic is Inline graphic so that each player will participate in Inline graphic different PGG groups. It can be clearly shown that the promotion of cooperation is very obvious and similar to that on the square lattices. Again, under the heterogeneous payoff distribution mechanism, the fraction of cooperators can be greatly elevated when compared to the traditional PGG model in which the total payoff will be equally allocated within a group. Likely, the critical synergy factors Inline graphic or Inline graphic will also be decreased into a lower value, below which the cooperators will completely disappear, or the cooperators and defectors will coexist. Thus, irrespective of the interaction networks, the heterogeneous investment can be regarded as an universally effective in promoting the evolution of cooperation.

Figure 5. Fraction of cooperators Inline graphic as a function of normalized enhancement factor Inline graphic on triangular lattices in which Inline graphic and Inline graphic as that in Ref.[52].

Figure 5

Other parameters are identical with those in Fig.1.

Conclusions and Discussions

To sum, a novel spatial PGG model with heterogeneous payoff investment is proposed to illustrate the public cooperation behavior among selfish agents. Distinguishing from the traditional version, the investment of cooperator for a given PGG group (i.e., Inline graphic) is mapped to the fraction of cooperators inside tuned by a parameter Inline graphic. (i.e., Inline graphic). In this case, a cooperator tends to give a larger investment share into a group with the higher cooperation level. On one hand, the cooperative groups will receive the more investment; On the other hand, the cooperator who gives this group will also obtain the higher payoff. This duplicate mechanism to promote the cooperation encourages more players to adopt the cooperation strategy and enhance the collective cooperation. Large scale numerical simulations display that the cooperation can be greatly promoted under the heterogeneous resource investment scheme, regardless of the interaction topology and initial state. Compared with the traditional PGG model, the critical values Inline graphic and Inline graphic will be largely reduced, which will be beneficial to the cooperators to resist the defective temptation. In particular, the phase diagrams are largely changed and the coexistence region between cooperators and defectors are absent under the weak selection limit. Altogether, current findings help to understand the persistence of cooperation within many real-world systems.

Funding Statement

This project is partially supported by the National Natural Science Foundation of China (NSFC) under Grant Nos. 61374169 and 11005047, the Young University Teacher's Fund of Anhui Province in China under Grant No. 2008jql071, and the Young Fund of Huaibei Normal University under Grant No. 2013xqz17. The funders had no role in study design,data collection and analysis, decision to publish, or preparation of the manuscript.

References

  • 1. Axelrod R, Hamilton WD (1981) The evolution of cooperation. Science 21: 1390–1396. [DOI] [PubMed] [Google Scholar]
  • 2.Axelrod R (1984) The Evolution of Cooperation. Basic Books, New York.
  • 3.Nowak MA (2006) Evolutionary Dynamics: Exploring the Equations of Life. Harvard University Press, Cambridge, MA.
  • 4.Weibull JW (1995) Evolutionary Game Theory. MIT Press, Boston, MA.
  • 5. Szabó G, Fáth G (2007) Evolutionary games on graphs. Phys Rep 446: 97. [Google Scholar]
  • 6. Roca CP, Cuesta JA, Sáchez A (2009) Evolutionary game theory: Temporal and spatial effects beyond replicator dynamics. Phys Life Rev 6: 208–249. [DOI] [PubMed] [Google Scholar]
  • 7. Perc M, Szolnoki A (2010) Coevolutionary games: A mini review. Biosystems 99: 109–125. [DOI] [PubMed] [Google Scholar]
  • 8. Wang Z, Perc M (2010) Aspiring to the fittest and promotion of cooperation in the prisoners dilemma game. Phys Rev E 82: 021115. [DOI] [PubMed] [Google Scholar]
  • 9. Perc M, Wang Z (2010) Heterogeneous aspirations promote cooperation in the prisoner's dilemma game. PLoS ONE 5: e15117. [DOI] [PMC free article] [PubMed] [Google Scholar]
  • 10. Ma ZQ, Xia CY, Sun SW, Wang L, Wang HB, et al. (2011) Heterogeneous link weight promotes the cooperation in spatial prisoner's dilemma. Int J Mod Phys C 22: 1257–1268. [Google Scholar]
  • 11. Vainstein MH, Silva ATC, Arenzon JJ (2007) Does mobility decrease cooperation? J Theor Biol 244: 722–729. [DOI] [PubMed] [Google Scholar]
  • 12. Wang Z, Wang Z, Zhu XD, Arenzon JJ (2012) Cooperation and age structure in spatial games. Phys Rev E 85: 011149. [DOI] [PubMed] [Google Scholar]
  • 13. Shigaki K, Tanimoto J, Wang Z, Kokubo S, Hgishima A, et al. (2012) Direct reciprocity in spatial populations enhances R-reciprocity as well ss ST-reciprocity. PLoS ONE 8: e71961. [DOI] [PMC free article] [PubMed] [Google Scholar]
  • 14. Tanimoto J, Brede M, Yamauchi A (2012) Network reciprocity by coexisting learning and teaching strategies. Phys Rev E 85: 032101. [DOI] [PubMed] [Google Scholar]
  • 15. Meloni S, Buscarino A, Fortuna L, Frasca M, Gomez-Gardeñes J, et al. (2009) Effects of mobility in a population of prisoners dilemma players. Phys Rev E 79: 067101. [DOI] [PubMed] [Google Scholar]
  • 16. Wang Z, Szolnoki A, Perc M (2012) If players are sparse social dilemmas are too: Importance of percolation for evolution of cooperation. Sci Rep 2: 369. [DOI] [PMC free article] [PubMed] [Google Scholar]
  • 17. Wang Z, Yin ZY, Wang L, Xia CY (2012) Inferring reputation promotes the evolution of cooperation in spatial social dilemma games. PLoS ONE 7: e40218. [DOI] [PMC free article] [PubMed] [Google Scholar]
  • 18. Du WB, Cao XB, Hu MB, Wang WX (2009) Asymmetriccostinsnowdrift game onscale- freenetworks. Europhys Lett 87: 60004. [Google Scholar]
  • 19. Shigaki K, Tanimoto J, Wang Z, Kokubo S, Hagishima A, et al. (2012) Referring to the social performance promotes cooperation in spatial prisoner's dilemma games. Phys Rev E 86: 031141. [DOI] [PubMed] [Google Scholar]
  • 20. Perc M, Gómez-Gardeñes J, Szolnoki A, Floría LM, Moreno Y (2013) Evolutionary dynamics of group interactions on structured populations: a review. JR Soc Interface 10: 20120997. [DOI] [PMC free article] [PubMed] [Google Scholar]
  • 21. Santos FC, Santos MD, Pacheco JM (2008) Social diversity promotes the emergence of cooperation in public goods games. Nature 454: 213–216. [DOI] [PubMed] [Google Scholar]
  • 22. Wang Z, Szolnoki A, Perc M (2012) Evolution of public cooperation on interdependent networks: The impact of biased utility functions. Europhys Lett 97: 48001. [Google Scholar]
  • 23. Zhang HF, Liu RR, Wang Z, Yang HX, Wang BH (2011) Aspiration-induced reconnection in spatial public-goods game. Europhys Lett 94: 18006. [Google Scholar]
  • 24. Wang Z, Xu ZJ, Huang JH, Zhang LZ (2010) Maintenance of cooperation induced by punishment in public goods games. Chin Phys B 19: 100204. [Google Scholar]
  • 25. Gómez-Gardeñes J, Romance M, Criado R, Vilone D, Sánchez A (2011) Evolutionary games defined at the network mesoscale: The public goods game. Chaos 21: 016113. [DOI] [PubMed] [Google Scholar]
  • 26. Szolnoki A, Wang Z, Perc M (2012) Wisdom of groups promotes cooperation in evolutionary social dilemmas. Sci Rep 2: 576. [DOI] [PMC free article] [PubMed] [Google Scholar]
  • 27. Wang Z, Szolnoki A, Perc M (2012) Percolation threshold determines the optimal population density for public cooperation. Phys Rev E 85: 037101. [DOI] [PubMed] [Google Scholar]
  • 28. Jin Q, Wang Z, Wang Z, Wang YL (2012) Strategy changing penalty promotes cooperation in spatial prisoner's dilemma game. Chaos, Solitons & Fractals. 45: 395. [Google Scholar]
  • 29. Xia CY, Meloni S, Moreno Y (2012) Effects of environment knowledge on agglomeration and cooperation in spatial Public Goods Games. Adv Compl Sys 15: 1250056. [Google Scholar]
  • 30.Ostrom E (1990) Governing the Commons: The Evolution of Institutions for Collective Action. Cambridge University Press, Cambridge, UK.
  • 31. Roca CP, Helbing D (2011) Emergence of social cohesion in a model society of greedy, mobile individuals. Proc Natl Acad Sci USA 108: 11370–11374. [DOI] [PMC free article] [PubMed] [Google Scholar]
  • 32. Hardin G (1968) The tragedy of the commons. Science 162: 1243–1248. [PubMed] [Google Scholar]
  • 33. Fehr E, Gintis H (2007) Human motivation and social cooperation: Experimental and analytical foundations. Annu Rev Sociol 33: 43–64. [Google Scholar]
  • 34. Fehr E, Gächter S (2002) Altruistic Punishment in Humans. Nature (London) 415: 137–140. [DOI] [PubMed] [Google Scholar]
  • 35. Boyd R, Gintis H, Bowles S, Richerson PJ (2003) The evolution of altruistic punishment. Proc Natl Acad Sci USA 100: 3531–3535. [DOI] [PMC free article] [PubMed] [Google Scholar]
  • 36. Helbing D, Szolnoki A, Perc M, Szabó G (2010) Punish, but not too hard: how costly punishment spreads in the spatial public goods game. New J Phys 12: 083005. [Google Scholar]
  • 37. Wang Z, Xu ZJ, Zhang LZ (2010) Punishment in optional public goods games. Chin Phys B 19: 110201. [Google Scholar]
  • 38. Panchanathan K, Boyd R (2004) Indirect reciprocity can stabilize cooperation without the second-order free rider problem. Nature (London) 432: 499–502. [DOI] [PubMed] [Google Scholar]
  • 39. Hauert C, Monte SD, Hofbauer J, Sigmund K (2002) Volunteering as red queen mechanism for cooperation in public goods games. Science 296: 1129–1132. [DOI] [PubMed] [Google Scholar]
  • 40. Nowak MA, Sigmund K (1998) Evolution of indirect reciprocity by image scoring. Nature (London) 393: 573–577. [DOI] [PubMed] [Google Scholar]
  • 41. Santos FC, Santos MD, Pacheco JM (2008) Social diversity promotes the emergence of cooperation in public goods games. Nature (London) 454: 213–216. [DOI] [PubMed] [Google Scholar]
  • 42. Szabó G, Hauert C (2002) Phase transitions and volunteering in spatial public goods games. Phys Rev Lett 89: 118101. [DOI] [PubMed] [Google Scholar]
  • 43. Guan JY, Wu ZX, Wang YH (2007) Effects of inhomogeneous activity of players and noise on cooperation in spatial public goods games. Phys Rev E 76: 056101. [DOI] [PubMed] [Google Scholar]
  • 44. Szabó G, Tőke C (1998) Evolutionary prisoner's dilemma game on a square lattice. Phys Rev E 58: 69–73. [Google Scholar]
  • 45. Szabó G, Vukov J, Szolnoki A (2005) Phase diagrams for an evolutionary prisoner's dilemma game on two-dimensional lattices. Phys Rev E 72: 047107. [DOI] [PubMed] [Google Scholar]
  • 46. Vukov J, Szabó G, Szolnoki A (2006) Cooperation in the noisy case: Prisoner's dilemma game on two types of regular random graphs. Phys Rev E 73: 067103. [DOI] [PubMed] [Google Scholar]
  • 47. Szolnoki A, Perc M (2013) Correlation of positive and negative reciprocity fails to confer an evolutionary advantage: Phase transitions to elementary strategies. Phys Rev X 3: 041021. [Google Scholar]
  • 48. Wang Z, Szolnoki A, Perc M (2013) Optimal interdependence between networks for the evolution of cooperation. Sci Rep 3: 2470. [DOI] [PMC free article] [PubMed] [Google Scholar]
  • 49. Zhang GQ, Sun QB, Wang L (2013) Noise-induced enhancement of network reciprocity in social dilemmas. Chaos Soliton & Fractals 51: 31. [Google Scholar]
  • 50. Wang Z, Szolnoki A, Perc M (2013) Interdependent network reciprocity in evolutionary games. Sci Rep 3: 1183. [DOI] [PMC free article] [PubMed] [Google Scholar]
  • 51. Yang HX, Wang WX, Wu ZX, Lai YC, Wang BH (2009) Diversity-optimized cooperation on complex networks. Phys Rev E 79: 056107. [DOI] [PubMed] [Google Scholar]
  • 52. Szolnoki A, Perc M, Szabó G (2009) Topology-independent impact of noise on cooperation in spatial public goods games. Phys Rev E 80: 056109. [DOI] [PubMed] [Google Scholar]
  • 53. Xu ZJ, Wang Z, Song HP, Zhang LZ (2010) Self-adjusting rule in spatial voluntary public goods games. Europhys Lett 90: 20001. [Google Scholar]
  • 54. Wang Z, Kokubo S, Tanimoto J, Fukuda E, Shigaki K (2013) Insight into the so-called spatial reciprocity. Phys Rev E 88: 042145. [DOI] [PubMed] [Google Scholar]
  • 55. Shigaki K, Wang Z, Tanimoto J, Fukuda E (2013) Effect of Initial Fraction of Cooperators on Cooperative Behavior in Evolutionary Prisoners Dilemma Game. PLoS ONE 8: e76942. [DOI] [PMC free article] [PubMed] [Google Scholar]

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