Checklist of British and Irish Hymenoptera - Evanioidea

Introduction

Abbreviations, rationale for species inclusion and additional information are given in the introductory article for the series. This article is the first in a series of 13 checklist sections covering the Hymenoptera of the British Isles. For each genus and species the original combination and a list of synonyms is provided under the heading 'nomenclature'.

A corresponding up-to-date checklist for Evanioidea is maintained on the Hymenoptera of the British Isles site.

Distribution data for the Evanioidea are taken from Crosskey (1951), with additional references given under notes for each taxon. This is a small superfamily of three extant families, but particularly poorly represented in northern Europe. The Evaniidae (sometimes referred to as 'ensign wasps') are, where known, parasitoids/predators of cockroach oothecae (Crosskey 1951; Brown 1973) and are morphologically very distinct from the remaining families. Whilst rather similar in appearance, the Aulacidae and Gasteruptiidae (Fig. 1) differ markedly in biology, with the single British aulacid, Aulacus striatus (Jurine), being a koinobiont endoparasitoid of Xiphydriidae larvae (see E.R. Skinner & W.G. Thompson's 1960 film, 'The Alder Woodwasp and its Insect Enemies', distributed by World Educational Films: http://worldeducationalfilms.com/the-alder-woodwasp-and-its-insect-enemies/) whilst Gasteruption species are essentially predators and cleptoparasites of solitary bee larvae (Malyshev 1964).

Figure 1.

Male Gasteruption jaculator (Linnaeus) (Wilmslow, Cheshire) (courtesy of A.M. Broome).

Checklists

Evanioidea

Aulacidae

Shuckard, 1841

Notes

Taxonomy follows Smith (2001).

Aulacus

Jurine, 1807

• DISPHAERON Dahlbom, 1837

• AULACINUS Westwood, 1868

• PAMMEGISCHIA Provancher, 1882

• DISAULACINUS Kieffer, 1910

• MICRAULACINUS Kieffer, 1910

• NEURAULACINUS Kieffer, 1910

• PARAFOENUS Kieffer, 1910

• PYCNAULACUS Cushman, 1929

Aulacus striatus

Jurine, 1807

• Aulacus arcticus Dahlbom, 1837

• Aulacus exaratus Ratzeburg, 1852

Distribution

England.

Evaniidae

Latreille, 1802

Notes

Taxonomy follows Deans (2005).

Evania

Fabricius, 1775

Evania appendigaster

(Linnaeus, 1758)

• Ichneumon appendigaster Linnaeus, 1758

• Ichneumon niger (de Geer, 1773, Ichneumon)

• Evania laevigata Olivier, 1792

• Evania unicolor Say, 1824

• Evania desjardinsi Blanchard, 1840

• Evania affinis Le Guillou, 1841

• Evania cubae Guérin-Méneville, 1844

• Evania peringueyi Cameron, 1906

Notes

# Presumably an introduction to Britain as it parasitises the oothecae of non-native synanthropic cockroaches (Blattidae). Crosskey (1951) could find no locality data for British specimens and there are no relevant specimens in NHM.

Brachygaster

Leach, 1830

Brachygaster minutus

(Olivier, 1792)

• Evania minuta Olivier, 1792

• Evania fulvipes (Curtis, 1829, Evania)

• Brachygaster rufipes Brullé, 1846 preocc.

• Evania brullei (Westwood, 1851, Evania)

• Brachygaster schlettereri Kieffer, 1912

Distribution

England.

Gasteruptiidae

Ashmead, 1890

Notes

Synonymic data mostly from Fauna Europaea.

Gasteruption

Latreille, 1796

• FOENUS Fabricius, 1798

• RHYDINOFOENUS Bradley, 1909

• DOLICHOFOENUS Kieffer, 1910

• TRICHOFOENUS Kieffer, 1910

• PLUTOFOENUS Kieffer, 1911

Gasteruption assectator

(Linnaeus, 1758)

• Ichneumon assectator Linnaeus, 1758

• Gasteruption affectator misspelling

• Ichneumon annularis (Geoffrey 1785, Ichneumon)

• Foenus incertus (Cresson 1864, Foenus)

• Foenus montanus (Cresson 1864, Foenus)

• Foenus arca (Couper 1870, Foenus)

• Foenus borealis (Thomson 1883, Foenus)

• Foenus fumipennis (Thomson 1883, Foenus)

• Foenus nigritarsis (Thomson 1883, Foenus)

• Gasteruption nitidulum Schletterer, 1885

• Gasteruption brevicauda Kieffer 1904

• Gasteruption micrura Kieffer 1904

• Gasteruption nevadense Kieffer 1904

• Gasteruption nigropectus Kieffer, 1904

• Gasteruption bakeri Kieffer 1910

• Trichofoenus canadensis (Kieffer 1910, Trichofoenus)

• Gasteruption abeillei Kieffer 1912

• Gasteruption aberrans Strand, 1912

• Trichofoenus breviterebrae (Watanabe 1934, Trichofoenus)

• Gasteruption utahensis Townes 1950

• Gasteruption margotae Madl 1987

Distribution

England, Wales, Ireland.

Gasteruption jaculator

(Linnaeus, 1758)

• Ichneumon jaculator Linnaeus, 1758

• Foenus granulithorax (Tournier, 1877, Foenus)

• Foenus obliteratus (Abeille de Perrin, 1879, Foenus)

• Foenus rugidorsus (Costa, 1884, Foenus)

• Gasteruption thomsoni Schletterer, 1885

Distribution

England.

Gasteruption laticeps

(Tournier, 1877)

• Foenus laticeps Tournier, 1877

• Gasteruption foveolatum Schletterer, 1889

• Gasteruption foveolum Szépligeti, 1903

Gasteruption minutum

(Tournier, 1877)

• Foenus minutum Tournier, 1877

• Foenus longigena (Thomson, 1883, Foenus)

Distribution

England, Wales (Alexander 1983).

Gasteruption pedemontanum

(Tournier, 1877)

• Foenus pedemontanum Tournier, 1877

• Gasteruption pedmontanum misspelling

• Foenus terrestre (Tournier, 1877, Foenus)

• Gasteruption trifossulatum Kieffer, 1904

Distribution

England.