Abstract Abstract
A checklist of the Lonchopteridae and Phoridae recorded from Finland is presented.
Keywords: Checklist, Finland, Diptera, Lonchopteridae, Phoridae
Introduction
The superfamily Phoroidea includes at least the fly families Phoridae, Lonchopteridae, and the small family Ironomyiidae known only from Australia. The flat-footed fly families Platypezidae and Opetiidae, treated in a separate paper in this volume, are placed either in their own basal superfamily Platypezoidea or included in Phoroidea (see Cumming et al. 1995, Woodley et al. 2009 and Wiegmann et al. 2011).
Lonchoptera Meigen, 1803 is the only currently recognized genus in Lonchopteridae. Five Lonchoptera species have been added to the Finnish fauna after the checklist of Hackman (1980) (Andersson 1991, Kahanpää 2013).
The scuttle flies (family Phoridae) may be the largest single family in Diptera. The true number of species in this family has been estimated to be over 30000, the vast majority still undescribed (Marshall 2012). The ‘megagenus’ Megaselia Rondani, 1856 contains about 50% of the species (Marshall 2012). Other phorid genera are relatively better known in Finland but none of them can be called well studied. It is likely that at least a hundred (or hundreds of) Finnish species remain unnoticed, most of them in Megaselia.
The checklist for Megaselia is mostly based on previously published records. The collections of Finnish Museum of Natural History (MZH) were examined by the author during the preparation of this checklist and a few obvious mistakes were taken into account. Most of the material in MZH was identified by H. Schmitz (between years 1915 and 1940) or E. Beyer (in 1958). The lists for other phorid genera are based on recently examined specimens from both MZH and private collections. Due to the difficulties encountered in identifying Megaselia, the list for this genus should definitely be considered preliminary. A revision of the Finnish material is sorely needed.
The Finnish species of these families were last listed by Hackman (1980).
Number of species (Lonchopteridae, Phoridae):
World: 63, 4105 species (Pape et al. 2011)
Europe: 13, 656 species
Finland: 8, ~224–234 species
Faunistic knowledge level in Finland: good, poor
Checklist
suborder Brachycera Macquart, 1834
clade Eremoneura Lameere, 1906
clade Cyclorrhapha Brauer, 1863
superfamily Phoroidea Curtis, 1833
LONCHOPTERIDAE Macquart, 1835
LONCHOPTERA Meigen, 1803
= Dipsa Fallén, 1810
Lonchoptera bifurcata (Fallén, 1810)
= furcata (Fallén, 1823)
Lonchoptera fallax de Meijere, 1906
Lonchoptera impicta Zetterstedt, 1848
Lonchoptera lutea Panzer, 1809
Lonchoptera meijerei Collin, 1938
Lonchoptera nigrociliata Duda, 1927
Lonchoptera nitidifrons Strobl, 1898
Lonchoptera scutellata Stein, 1890
PHORIDAE Curtis, 1833
AENIGMATIAS Meinert, 1890
Aenigmatias lubbockii (Verrall, 1877)
ANEVRINA Lioy, 1864
Anevrina thoracica (Meigen, 1804)
Anevrina unispinosa (Zetterstedt, 1860)
= fennica (Becker, 1901)
Anevrina urbana (Meigen, 1830)
BOROPHAGA Enderlein, 1924
Borophaga agilis (Meigen, 1830)
Borophaga carinifrons (Zetterstedt, 1848)
Borophaga femorata (Meigen, 1830)
Borophaga incrassata (Meigen, 1830)
Borophaga subsultans (Linnaeus, 1767)
= okellyi Schmitz, 1937
CHAETOPLEUROPHORA Schmitz, 1922
Chaetopleurophora erythronota (Strobl, 1892)
Chaetopleurophora spinosissima (Strobl, 1892)
CONICERA Meigen, 1830
Conicera dauci (Meigen, 1830)
= atra Meigen, 1830
Conicera floricola Schmitz, 1938
= similis auct. nec (Haliday, 1833)
= minuscula Schmitz, 1953
Conicera schnittmanni Schmitz, 1926
Conicera similis (Haliday, 1833)
= pauxilla Schmitz, 1920
? Conicera tarsalis Schmitz, 1920
Conicera tibialis Schmitz, 1925
= fallens Schmitz, 1948
DIPLONEVRA Lioy, 1864
= Diploneura emend.
Diplonevra florescens (Turton, 1801)
= florea (Fabricius, 1794) preocc.
= abdominalis (Fallén, 1823)
Diplonevra concinna (Meigen, 1830)
Diplonevra freyi Schmitz, 1927
Diplonevra funebris (Meigen, 1830)
= rostralis (Schmitz, 1918)
Diplonevra glabra Schmitz, 1927
= parcepilosa Schmitz, 1927
Diplonevra nitidula (Meigen, 1830)
Diplonevra oldenbergi Schmitz, 1920
DOHRNIPHORA Dahl, 1898
Dohrniphora cornuta (Bigot, 1857)
GYMNOPHORA Macquart, 1835
Gymnophora arcuata (Meigen, 1830)
Gymnophora healeyae Disney, 1980
Gymnophora nigripennis Schmitz, 1926
Gymnophora quartomollis Schmitz, 1920
HYPOCERA Lioy, 1864
Hypocera mordellaria (Fallén, 1823)
MEGASELIA Rondani, 1856
= Aphiochaeta Brues, 1903
= Plastophora Brues, 1905
Megaselia aequalis (Wood, 1909)
Megaselia affinis (Wood, 1909)
= proxima (Lundbeck, 1920)
Megaselia albicaudata (Wood, 1910)
Megaselia albiclava (Schmitz, 1926)
Megaselia altifrons (Wood, 1909)
Megaselia analis (Lundbeck, 1920)
? Megaselia angelicae (Wood, 1910)
Megaselia angularis (Schmitz, 1924)
Megaselia angusta (Wood, 1909)
= angustata emend.
= pulicaria auct. nec (Fallén, 1923)
Megaselia annulipes (Schmitz, 1921)
Megaselia armata (Wood, 1909)
Megaselia atrosericea Schmitz, 1927
Megaselia auricoma Schmitz, 1927
Megaselia baltica (Schmitz, 1924)
Megaselia barbulata (Wood, 1909)
= depilata (Lundbeck, 1921)
? Megaselia basispinata (Lundbeck, 1920)
Megaselia basiveluta Schmitz, 1935
Megaselia beckeri (Wood, 1909)
= laticrus Schmitz, 1927
Megaselia berndseni (Schmitz, 1919)
= pygmaeoides (Lundbeck, 1921)
Megaselia bovista (Gimmerthal, 1848)
Megaselia brevicostalis (Wood, 1910)
Megaselia breviseta (Wood, 1912)
Megaselia breviterga (Lundbeck, 1920)
= similata (Lundbeck, 1921)
Megaselia campestris (Wood, 1908)
Megaselia ciliata (Zetterstedt, 1848)
Megaselia cinereifrons (Strobl, 1910)
Megaselia clara (Schmitz, 1921)
Megaselia coaetanea Schmitz, 1929
Megaselia coccyx Schmitz, 1965
? Megaselia collini (Wood, 1909)
Megaselia conformis (Wood, 1909)
Megaselia costalis (von Roser, 1840)
Megaselia curvivenia Schmitz, 1928
Megaselia dahli (Becker, 1901)
Megaselia discreta (Wood, 1909)
= nudiventris (Wood, 1909)
Megaselia diversa (Wood, 1909)
Megaselia dubitalis (Wood, 1908)
Megaselia eccoptomera Schmitz, 1927
Megaselia eisfelderae Schmitz, 1948
Megaselia elongata (Wood, 1914)
Megaselia erecta (Wood, 1910)
Megaselia exarcuata Schmitz, 1927
Megaselia excavata Schmitz, 1927
Megaselia excorticata Disney, 2009
Megaselia fennicola Beyer, 1958
Megaselia flammula Schmitz, 1928
Megaselia flava (Fallén, 1823)
Megaselia flavicoxa (Zetterstedt, 1848)
Megaselia fumata (Malloch, 1909)
Megaselia funeralis Schmitz, 1928
Megaselia fungivora (Wood, 1909)
Megaselia fusca (Wood, 1909)
Megaselia fuscipalpis (Lundbeck, 1920)
Megaselia fuscoides Schmitz, 1934
Megaselia fuscovariana Schmitz, 1933
Megaselia giraudii (Egger, 1862)
Megaselia glabrifrons (Wood, 1909)
Megaselia groenlandica (Lundbeck, 1901)
Megaselia halterata (Wood, 1910)
= plurispinosa (Lundbeck, 1920)
Megaselia hectochaeta Schmitz, 1974
Megaselia hilaris Schmitz, 1927
Megaselia hirsuta (Wood, 1910)
Megaselia hirticaudata (Wood, 1910)
Megaselia hirticrus (Schmitz, 1918)
Megaselia humeralis (Zetterstedt, 1838)
= cubitalis (Becker, 1901)
Megaselia hyalipennis (Wood, 1912)
Megaselia infraposita (Wood, 1909)
Megaselia ignobilis (Schmitz, 1919)
Megaselia intonsa Schmitz, 1948
Megaselia involuta (Wood, 1910)
Megaselia latifemorata (Becker, 1901)
Megaselia limburgensis (Schmitz, 1918)
Megaselia longicostalis (Wood, 1912)
Megaselia longifurca (Lundbeck, 1921)
= spinolabella Disney, 1989
Megaselia longipalpis (Wood, 1910)
Megaselia longiseta (Wood, 1909)
Megaselia lucifrons (Schmitz, 1918)
= subnitida (Lundbeck, 1920)
Megaselia luminifrons (Schmitz, 1926)
Megaselia lutea (Meigen, 1830)
Megaselia major (Wood, 1912)
Megaselia mallochi (Wood, 1909)
Megaselia manicata (Wood, 1910)
Megaselia maura (Wood, 1910)
Megaselia meconicera (Speiser, 1925)
Megaselia meigeni (Becker, 1901)
? Megaselia miki Schmitz, 1929
Megaselia minor (Zetterstedt, 1848)
= angustifrons (Wood, 1912)
Megaselia minuta (Aldrich, 1892)
= luminosa Schmitz, 1952
Megaselia mixta (Schmitz, 1918)
Megaselia nasoni (Malloch, 1914)
= coaequalis (Schmitz, 1919)
Megaselia nigra (Meigen, 1830)
= albidihalteralis (Felt, 1896)
Megaselia nigriceps (Loew, 1866)
= projecta (Becker, 1901)
Megaselia nigripalpis (Lundbeck, 1920)
Megaselia nudipleura (Beyer, 1958)
Megaselia obscuripennis (Wood, 1909)
Megaselia offuscata (Schmitz, 1921)
Megaselia opacicornis Schmitz, 1949
Megaselia parnassia Disney, 1986
Megaselia parva (Wood, 1909)
Megaselia pectoralis (Wood, 1910)
Megaselia pectunculata Schmitz, 1927
Megaselia picta (Lehmann, 1822)
Megaselia pleuralis (Wood, 1909)
Megaselia plurispinulosa (Zetterstedt, 1860)
Megaselia posticata (Strobl, 1898)
Megaselia praeacuta (Schmitz, 1919)
Megaselia prodroma (Lundbeck, 1921)
Megaselia producta (Schmitz, 1921)
= sordescens auct. nec Schmitz, 1927
Megaselia propinqua (Wood, 1909)
Megaselia pubecula Schmitz, 1927
Megaselia pulicaria (Fallén, 1923)
= sinuata Schmitz, 1926
Megaselia pumila (Meigen, 1830)
= atripes (Brues, 1915)
Megaselia pusilla (Meigen, 1830)
Megaselia pygmaea (Zetterstedt, 1848)
Megaselia quadriseta (Schmitz, 1918)
= badia Schmitz, 1938
? Megaselia raetica Schmitz, 1934
Megaselia robertsoni Disney, 2008
Megaselia robusta Schmitz, 1928
Megaselia rubricornis (Schmitz, 1919)
Megaselia rudis (Wood, 1909)
Megaselia ruficornis (Meigen, 1830)
Megaselia rufipes (Meigen, 1804)
Megaselia scalaris (Loew, 1866)
Megaselia scutellaris (Wood, 1909)
= scutellariformis (Schmitz, 1926)
Megaselia sepulchralis (Lundbeck, 1920)
Megaselia setulipalpis Schmitz, 1938
Megaselia sordida (Zetterstedt, 1838)
= eminens Schmitz, 1953
= semiscaura Schmitz, 1927
? Megaselia specularis Schmitz, 1935
Megaselia spinicincta (Wood, 1910)
Megaselia spinigera (Wood, 1908)
Megaselia styloprocta (Schmitz, 1921)
Megaselia subcarpalis (Lundbeck, 1920)
Megaselia subpalpalis (Lundbeck, 1920)
Megaselia subpleuralis (Wood, 1909)
Megaselia subtumida (Wood, 1909)
Megaselia sulphuripes (Meigen, 1830)
Megaselia superciliata (Wood, 1910)
Megaselia superfurcata Schmitz, 1928
Megaselia sylvatica (Wood, 1910)
= impolluta (Schmitz, 1920)
Megaselia tarsalis (Wood, 1910)
Megaselia tenuiventris Schmitz, 1927
Megaselia tignorum Disney, 2009
Megaselia tumida (Wood, 1909)
= setifer Lundbeck, 1920
Megaselia unicolor (Schmitz, 1919)
Megaselia valvata Schmitz, 1935
Megaselia variana Schmitz, 1926
Megaselia vestita (Wood, 1914)
Megaselia winqvisti Disney, 2011
Megaselia woodi (Lundbeck, 1922)
Megaselia xanthozona (Strobl, 1892)
= euryprocta Schmitz, 1957
Megaselia zonata (Zetterstedt, 1838)
MENOZZIOLA Schmitz, 1927
Menozziola obscuripes (Schmitz, 1927)
METOPINA Macquart, 1835
Metopina galeata (Haliday, 1833)
= inaequalis Schmitz, 1927
Metopina oligoneura (Mik, 1867)
Metopina pileata Schmitz, 1936
MICROSELIA Schmitz, 1934
Microselia forsiusi (Schmitz, 1927)
PHALACROTOPHORA Enderlein, 1912
Phalacrotophora berolinensis Schmitz, 1920
Phalacrotophora beuki Disney in Disney & Beuk, 1997
Phalacrotophora fasciata (Fallén, 1823)
PHORA Latreille, 1796
= Trineura Meigen, 1803
Phora artifrons Schmitz, 1920
Phora atra (Meigen, 1804)
= aterrima (Fabricius, 1794) preocc.
Phora bullata Schmitz, 1927
Phora convallium Schmitz, 1928
Phora convergens Schmitz, 1920
Phora dubia (Zetterstedt, 1848)
= schineri (Becker, 1901)
Phora edentata Schmitz, 1920
Phora hamata Schmitz, 1927
Phora holosericea Schmitz, 1920
Phora hyperborea Schmitz, 1927
Phora obscura (Zetterstedt, 1848)
Phora occidentata Malloch, 1912
= zetterstedti Schmitz, 1927
Phora penicillata Schmitz, 1920
Phora praepandens Schmitz, 1927
Phora pubipes Schmitz, 1920
Phora stictica Meigen, 1830
Phora tincta Schmitz, 1920
PLECTANOCNEMA Schmitz, 1926
Plectanocnema nudipes (Becker, 1901)
PSEUDACTEON Coquillett, 1907
Pseudacteon fennicus Schmitz, 1927
Pseudacteon formicarum (Verrall, 1877)
SPINIPHORA Malloch, 1909
Spiniphora bergenstammi Mik, 1864
Spiniphora dorsalis (Becker, 1901)
Spiniphora excisa (Becker, 1901)
Spiniphora maculata (Meigen, 1830)
= helicivora (Dufour, 1841)
TRIPHLEBA Rondani, 1856
= Citrago Schmitz, 1924
Triphleba admirabilis Schmitz, 1927
? Triphleba aequalis (Schmitz, 1919)
? Triphleba autumnalis (Becker, 1901)
Triphleba bicornuta (Strobl, 1910)
= uncinata (Schmitz, 1918)
Triphleba citreiformis (Becker, 1901)
Triphleba distinguenda (Strobl, 1892)
= unicalcarata (Becker, 1901)
Triphleba excisa (Lundbeck, 1921)
Triphleba gracilis (Wood, 1907)
Triphleba hyalinata (Meigen, 1830)
? Triphleba inaequalis Schmitz, 1943
Triphleba intermedia (Malloch, 1908)
Triphleba lugubris (Meigen, 1830)
Triphleba luteifemorata (Wood, 1906)
Triphleba nudipalpis (Becker, 1901)
Triphleba opaca (Meigen, 1830)
Triphleba pachyneurella (Schmitz, 1919)
Triphleba palposa (Zetterstedt, 1848)
Triphleba renidens Schmitz, 1927
Triphleba subcompleta Schmitz, 1927
Triphleba trinervis (Becker, 1901)
VERUANUS Schmitz, 1927
Veruanus oldenbergi Schmitz, 1919
= memorabilis Schmitz, 1927
Excluded species
Megaselia cirriventris Schmitz, 1929 mistake?
Megaselia crassipes (Wood, 1909) misidentified
Megaselia digitalis Schmitz, 1957 mistake?
Megaselia indifferens (Lundbeck, 1920) not found within present borders
Megaselia lapponica Schmitz, 1928 not found within present borders
Megaselia luteipes (Schmitz, 1918) misidentified
Megaselia palmeni (Becker, 1901) not found within present borders
Megaselia septentrionalis (Schmitz, 1919) not found within present borders
Megaselia simulans (Wood, 1912) not found within present borders ?
Triphleba gilvipes Schmitz, 1943 not found within present borders
Notes
Conicera tarsalis, Megaselia collini. Records of these species are based on single females identified by H. Schmitz.
Megaselia basispinata, Megaselia miki, Megaselia raetica. Species recorded from Finland by Hackman (1980) and Disney (1991) but no Finnish specimens or published records with details could be located.
Megaselia cirriventris, Megaselia digitalis. These species were recorded from ‘Fennia’ and ‘Lappland’ by Schmitz (1958), but all specimens listed are from the Abisko region in northen Sweden. It seems likely that Schmitz erroneously assigned the material from Swedish Lappland to Finland.
Megaselia luteipes. Examined specimens belong to Megaselia sp. not luteipes (females) (Espoo, leg. W. Hackman) and possibly Megaselia sp. 24 of Disney (2013) (Finland, Muonio, leg. J. Sahlberg and Russia, Kola, Kantalahti, leg. R. Frey).
Megaselia pulicaria agg. This difficult complex of species was revised by Disney (1999). The Finnish material has not been re-examined. The Finnish species involved are Megaselia angusta, Megaselia costalis, Megaselia longicostalis, Megaselia longifurca, Megaselia pulicaria, Megaselia subtumida and Megaselia tumida.
Megaselia simulans. Recorded only(?) from Kuusamo, a municipality split between Finland and Russia in 1944. Most records preceding the split are from the Russian side. No material was found in MZH.
Megaselia specularis. Recorded, with some doubt, from Finland by Hackman (1963). No material could be found in MZH.
Triphleba aequalis. Schmitz (1943) mention a Finnish male stored in MZH, but no such specimen could be found.
Triphleba autumnalis, Triphleba inaequalis. Finnish according to Disney (1991), but no specimens or detailed records could be located.
Citation
Kahanpää J (2014) Checklist of the families Lonchopteridae and Phoridae of Finland (Insecta, Diptera). In: Kahanpää J, Salmela J (Eds) Checklist of the Diptera of Finland. ZooKeys 441: 213–223. doi: 10.3897/zookeys.441.7197
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