Abstract Abstract
A checklist of the Diptera superfamilies Conopoidea (Conopidae), Nerioidea (Micropezidae, Pseudopomyzidae) and Diopsoidea (Megamerinidae, Psilidae, Strongylophthalmyiidae, Tanypezidae) from Finland in presented. Myopa vicaria Walker, 1849 is formally recorded for the first time from the country.
Keywords: Checklist, Finland, Diptera, biodiversity, faunistics
Introduction
Four fly families form the superfamily Nerioidea. Two of them, Micropezidae and Pseudopomyzidae, occur in Finland. Neriids are mostly tropical and cypselomatids (sensu stricto) have only been found in the Australasian and Oriental regions. The pseudopomyzids were included in Cypselosomatidae Hendel, 1931 in the previous checklist (Hackman 1980) and are still often included in Cypselosomatidae sensu lato. The Finnish micropezid fauna is relatively well known. One or two species of Cnodacophora Czerny, 1930 and Rainieria Rondani, 1843 could be present but are so far undetected.
Diopsoidea consists of seven to ten families (depending on the source), none particularly species-rich. Only four are found in Finland: Psilidae, Tanypezidae, Strongylophthalmyiidae and Megamerinidae. Tanypezidae and Strongylophthalmyiidae are here treated as separate families following Lonsdale (2013). Megamerinidae has been assigned to several superfamilies, most often Diopsoidea following McAlpine (1989), but also Nerioidea, Sciomyzoidea, and recently Opomyzoidea (Wiegmann et al. 2011). These families have raised no particular interest among Finnish dipterists. The genus Chamaepsila Hendel, 1917 in particular needs more attention. It is likely that a few Finnish species remain unrecognised in this genus.
The systematic position of the Conopidae is unclear. It has traditionally been considered basal within Schizophora. Wiegmann et al. (2011) listed Conopidae within Sciomyzoidea while others (see Skevington et al. 2010) place the family near Tephritoidea. Within the family the recent classification of Gibson and Skevington (2013) is followed. The Finnish conopid fauna is now relatively well-understood and few additional species are expected to occur in the country.
The Finnish species of these families were last listed by Hackman (1980).
Table 1.
Number of species by family.
| Family | Number of species in | Level of knowledge | ||
|---|---|---|---|---|
| World (Pape et al. 2011) | Europe | Finland | ||
| Conopidae | ~796 | 83 | 19 | good |
| Nerioidea: | ||||
| Micropezidae | 573 | 22 | 6 | average |
| Pseudopomyzidae | 18 | 1 | 1 | good |
| Diopsoidea: | ||||
| Psilidae | 320 | 47 | 29 | average |
| Tanypezidae | 27 (Lonsdale 2013) | 1 | 1 | good |
| Strongylophthalmyiidae | 47 (Lonsdale 2013) | 2 | 2 | good |
| Megamerinidae | 15 | 1 | 1 | good |
Checklist part 1: Conopoidea
suborder Brachycera Macquart, 1834
clade Eremoneura Lameere, 1906
clade Cyclorrhapha Brauer, 1863
infraorder Schizophora Becher, 1882
? clade Archischiza Enderlein, 1936
superfamily Conopoidea Latreille, 1802
CONOPIDAE Latreille, 1802
CONOPINAE Latreille, 1802
CONOPS Linnaeus, 1758
sg. Conops Linnaeus, 1758
Conops quadrifasciatus De Geer, 1776
Conops strigatus Wiedemann, 1824
Conops vesicularis Linnaeus, 1761
PHYSOCEPHALA Schiner, 1861
Physocephala nigra (De Geer, 1776)
ZODIONINAE Rondani, 1856
ZODION Latreille, 1796
Zodion cinereum (Fabricius, 1794)
= notatum Meigen, 1804
SICINAE Zimina, 1960
SICUS Scopoli, 1763
Sicus ferrugineus (Linnaeus, 1761)
MYOPINAE Macquart, 1834
MYOPA Fabricius, 1775
Myopa buccata (Linnaeus, 1758)
Myopa fasciata Meigen, 1804
Myopa hirsuta Stuke & Clements, 2008
= strandi auct. nec Duda, 1940
Myopa occulta Wiedemann, 1824
= nigrifrons von Bonsdorff, 1866
Myopa tessellatipennis Motschulsky, 1859
= polystigma auct. nec Rondani, 1857
Myopa testacea (Linnaeus, 1767)
Myopa vicaria Walker, 1849
= villosa Ringdahl, 1945
= strandi Duda, 1940
MYOPOTTA Zimina, 1969
Myopotta pallipes (Wiedemann, 1824)
THECOPHORA Rondani, 1845
= Occemyia Robineau-Desvoidy, 1853
Thecophora cinerascens (Meigen, 1804)
= pusilla (Meigen, 1824)
= atra misid.
Thecophora distincta (Wiedemann, 1824)
= melanopa misid.
Thecophora fulvipes (Robineau-Desvoidy, 1830)
= sundewalli (Zetterstedt, 1844)
Thecophora jakutica Zimina, 1974
= atra misid.
DALMANNIINAE Hendel, 1916
DALMANNIA Robineau-Desvoidy, 1830
Dalmannia dorsalis (Fabricius, 1794)
= punctata misid.
Checklist part 2: Nerioidea and Diopsoidea
suborder Brachycera Macquart, 1834
clade Eremoneura Lameere, 1906
clade Cyclorrhapha Brauer, 1863
infraorder Schizophora Becher, 1882
clade Muscaria Enderlein, 1936
parvorder Acalyptratae Macquart, 1835
superfamily Nerioidea Westwood, 1840
MICROPEZIDAE Blanchard, 1840
CALOBATINAE Bigot, 1853
CALOBATA Meigen, 1803
Calobata petronella (Linnaeus, 1761)
NERIA Robineau-Desvoidy, 1830
Neria cibaria (Linnaeus, 1761)
Neria commutata (Czerny, 1930)
= nigricornis auct. nec Zetterstedt, 1838
Neria ephippium (Fabricius, 1794)
Neria nigricornis (Zetterstedt, 1838)
= helleni (Frey, 1918)
= nitidicollis (Frey, 1947)
MICROPEZINAE Blanchard, 1840
MICROPEZA Meigen, 1803
Micropeza corrigiolata (Linnaeus, 1767)
PSEUDOPOMYZIDAE McAlpine, 1966
PSEUDOPOMYZA Strobl, 1893
Pseudopomyza atrimana (Meigen, 1830)
superfamily Diopsoidea Billberg, 1820
MEGAMERINIDAE Hendel, 1913
MEGAMERINA Rondani, 1861
= Lissa Meigen, 1826 preocc.
Megamerina dolium (Fabricius, 1805)
= loxocerina (Fallén, 1820)
PSILIDAE Macquart, 1835
CHYLIZINAE Rondani, 1856
CHYLIZA Fallén, 1820
sg. Chyliza Fallén, 1820
Chyliza annulipes Macquart, 1835
= fuscipennis auct. nec Robineau-Desvoidy, 1830
Chyliza leptogaster (Panzer, 1798)
= scutellata (Fabricius, 1798)
Chyliza nova Collin, 1944
Chyliza vittata Meigen, 1826
PSILINAE Macquart, 1835
tribe Loxocerini Macquart, 1835
IMANTIMYIA Frey, 1925
Imantimyia albiseta (Schrank, 1803)
= ichneumonea (Linnaeus, 1758) nom. dubium (see Notes)
Imantimyia fulviventris (Meigen, 1826)
Imantimyia nigrifrons (Macquart, 1835)
Imantimyia sylvatica (Meigen, 1826)
LOXOCERA Meigen, 1803
sg. LOXOCERA Meigen, 1803
Loxocera aristata (Panzer, 1801)
= ichneumonea (Linnaeus, 1758) nom. dubium (see Notes)
tribe Psilini Macquart, 1835
CHAMAEPSILA Hendel, 1917
= Tetrapsila Frey, 1925
Chamaepsila atra (Meigen, 1826)
Chamaepsila bicolor (Meigen, 1826)
= nigromaculata (Strobl, 1909)
Chamaepsila buccata (Fallén, 1826)
= gracilis (Meigen, 1826)
Chamaepsila humeralis (Zetterstedt, 1847)
Chamaepsila limbatella (Zetterstedt, 1847)
Chamaepsila morio (Zetterstedt, 1835)
Chamaepsila nigra (Fallén, 1820)
Chamaepsila nigricornis (Meigen, 1826)
Chamaepsila nigrosetosa Frey, 1925
Chamaepsila obscuritarsis (Loew, 1856)
Chamaepsila pallida (Fallén, 1820)
Chamaepsila pectoralis (Meigen, 1826)
Chamaepsila rosae (Fabricius, 1794)
Chamaepsila rufa (Meigen, 1826)
Chamaepsila unilineata (Zetterstedt, 1847)
PSILA Meigen, 1803
sg. FREYOPSILA Shatalkin, 1986
Psila sibirica (Frey, 1925)
sg. PSILA Meigen, 1803
Psila fimetaria (Linnaeus, 1761)
Psila merdaria Collin, 1944
PSILOSOMA Zetterstedt, 1860
Psilosoma audouini (Zetterstedt, 1835)
Psilosoma lefebvrei (Zetterstedt, 1835)
STRONGYLOPHTHALMYIIDAE Hendel, 1917
STRONGYLOPHTHALMYIA Heller, 1902
Strongylophthalmyia pictipes Frey, 1935
Strongylophthalmyia ustulata (Zetterstedt, 1847)
TANYPEZIDAE Rondani, 1856
TANYPEZA Fallén, 1820
Tanypeza longimana Fallén, 1820
Excluded species
Dalmannia punctata (Fabricius, 1794) not found within present borders.
Notes
Chamaepsila unilineata (Zetterstedt, 1847) and Chamaepsila nigrosetosa Frey, 1925 were synonymized with Chamaepsila pallida but they have recently been reinstated as valid species (Shatalkin and Merz 2010).
Loxocera ichneumonea (Linnaeus, 1758). This name has been used for Loxocera aristata (Panzer) and Loxocera albiseta (Schrank). Unfortunately no type material is known. Linné’s original description is insufficient and may in fact belong to a hoverfly (see Buck and Marshall 2006).
Myopa vicaria Walker, 1849. One of the common Myopa species through Finland, but previously unreported due to difficulties in identification. Recorded from provinces Ab, N, Ka, St, Ta, Sa, Kl, Om, Ok, Lkoc and Li.
Citation
Kahanpää J, Stuke J-H (2014) Checklist of the superfamilies Conopoidea, Diopsoidea and Nerioidea of Finland (Insecta, Diptera). In: Kahanpää J, Salmela J (Eds) Checklist of the Diptera of Finland. ZooKeys 441: 251–257. doi: 10.3897/zookeys.441.7227
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