Abstract Abstract
A Finnish checklist of the sphaeroceroid fly families Chyromyidae and Heleomyzidae is provided.
Keywords: Species list, Finland, Diptera, biodiversity, faunistics
Introduction
The superfamily Sphaeroceroidea is a medium-sized one, with two families of moderate diversity, Sphaeroceridae (1550 species) and Heleomyzidae (~720 species), and the small family Chyromyidae. The enigmatic afrotropical Mormotomyia hirsuta Austen, 1936 was once placed near Sphaeroceridae but it is now seen as an ephydroid fly (Kirk-Spriggs et al. 2011). McAlpine (2007) has proposed an alternative concept for Sphaeroceroidea with Sphaeroceridae and Heleomyzidae united as a single family called Heteromyzidae. This proposal has not gained significant support and for the purposes of this checklist the traditional concept of family Sphaeroceridae is retained.
There is no general agreement on the relationships of various heleomyzid tribes. Several different schemes for subfamilies have been proposed (see McAlpine 2007, McAlpine and Woodley 2010). Some taxa treated here as heleomyzids (primarily Trixoscelidinae, Chiropteromyzinae, Heteromyzinae and Borboropsini) may deserve full family status. As a conservative approach this checklist follows Marshall (2012) and keeps them as subfamilies and tribes. The heleomyzid subfamilies and tribes are listed alphabetically.
The Finnish chyromyids are small yellow flies with (at least while alive) iridescent blue or green eyes. Chyromyids are rarely collected and little is known about their ecology or the proper place of the family within Sphaeroceroidea. They may actually be a specialized lineage arising from within Heleomyzidae sensu lato.
Two of the three sphaeroceroid families are treated in this paper. The largest, Sphaeroceridae, is covered in a separate paper in this issue of ZooKeys. The Finnish species of Heleomyzidae and Chyromyidae were last listed by Hackman (1980).
Table 1.
Number of species by family.
| Family | Number of species in | Level of knowledge | ||
|---|---|---|---|---|
| World (Pape et al. 2011) | Europe | Finland | ||
| Chyromyidae | 138 | 59 | 4 | poor–average |
| Heleomyzidae | 727 | 175 | 61 | average |
Checklist
suborder Brachycera Macquart, 1834
clade Eremoneura Lameere, 1906
clade Cyclorrhapha Brauer, 1863
infraorder Schizophora Becher, 1882
clade Muscaria Enderlein, 1936
parvorder Acalyptratae Macquart, 1835
superfamily Sphaeroceroidea Macquart, 1835
CHYROMYIDAE Hendel, 1916
CHYROMYINAE Hendel, 1916
CHYROMYA Robineau-Desvoidy, 1830
Chyromya flava (Linnaeus, 1758)
Chyromya oppidana (Scopoli, 1763)
GYMNOCHIROMYIA Hendel, 1933
Gymnochiromyia flavella (Zetterstedt, 1848)
= minima (Becker, 1904)
Gymnochiromyia inermis (Collin, 1933)
HELEOMYZIDAE Westwood, 1840
BORBOROPSINAE Griffiths, 1972
BORBOROPSIS Czerny, 1902
Borboropsis puberula (Zetterstedt, 1838)
= fulviceps (Strobl, 1898)
CHIROPTEROMYZINAE Frey, 1952
CHIROPTEROMYZA Frey, 1952
Chiropteromyza broersei (de Meijere, 1946)
= wegelii Frey, 1952
NEOSSOS Malloch, 1927
= Ornitholeria Frey, 1930
Neossos nidicola (Frey, 1930)
HETEROMYZINAE Fallén, 1820
HETEROMYZA Fallén, 1820
Heteromyza atricornis Meigen, 1830
Heteromyza oculata Fallén, 1820
Heteromyza rotundicornis (Zetterstedt, 1846)
TEPHROCHLAMYS Loew, 1862
Tephrochlamys flavipes (Zetterstedt, 1838)
Tephrochlamys rufiventris (Meigen, 1830)
= lapponica (Czerny, 1924)
Tephrochlamys steniusi Frey, 1930
Tephrochlamys tarsalis (Zetterstedt, 1847)
HELEOMYZINAE Westwood, 1840
tribe Heleomyzini Westwood, 1840
GYMNOMUS Loew, 1863
Gymnomus amplicornis (Czerny, 1924)
HELEOMYZA Fallén, 1810
= Helomyza Fallén, 1820 emend.
= Leria Robineau-Desvoidy, 1830
sg. Heleomyza Fallén, 1810
Heleomyza borealis Boheman, 1865
= czernyi Collart, 1933
= modesta misid.
Heleomyza hackmani Frey, 1950
Heleomyza pleuralis (Becker, 1907)
Heleomyza serrata (Linnaeus, 1758)
MORPHOLERIA Garrett, 1921
sg. Spanoparea Czerny, 1924
Morpholeria dudai (Czerny, 1924)
Morpholeria kerteszii Czerny, 1924
Morpholeria obscuriventris (Zetterstedt, 1847)
Morpholeria ruficornis (Meigen, 1830)
NEOLERIA Malloch, 1919
Neoleria inscripta (Meigen, 1830)
= minuta (Zetterstedt, 1838)
Neoleria prominens (Becker, 1897)
= tibialis misid.
Neoleria ruficauda (Zetterstedt, 1847)
Neoleria ruficeps (Zetterstedt, 1838)
SCOLIOCENTRA Loew, 1862
sg. Chaetomus Czerny, 1924
Scoliocentra confusa (Wahlgren, 1918)
Scoliocentra flavotestacea (Zetterstedt, 1838)
sg. Leriola Gorodkov, 1962
Scoliocentra brachypterna (Loew, 1873)
Scoliocentra nigrinervis (Wahlgren, 1918)
sg. Scoliocentra Loew, 1862
Scoliocentra dupliciseta (Strobl, 1894)
Scoliocentra scutellaris (Zetterstedt, 1838)
Scoliocentra villosa (Meigen, 1830)
tribe Oecotheini Gorodkov, 1972
ECCOPTOMERA Loew, 1862
Eccoptomera infuscata Wahlgren, 1918
Eccoptomera longiseta (Meigen, 1830)
Eccoptomera marginicornis Czerny, 1924
Eccoptomera microps (Meigen, 1830)
Eccoptomera obscura (Meigen, 1830)
Eccoptomera ornata Loew, 1862
Eccoptomera pallescens (Meigen, 1830)
OECOTHEA Haliday, 1837
Oecothea fenestralis (Fallén, 1820)
tribe Orbelliini Gorodkov, 1972
ORBELLIA Robineau-Desvoidy, 1830
Orbellia nivicola Frey, 1913
SUILLIINAE Wahlgren, 1917
SUILLIA Robineau-Desvoidy, 1830
= Allophyla Loew, 1862
Suillia affinis (Meigen, 1830)
Suillia apicalis (Loew, 1862)
Suillia atricornis (Meigen, 1830)
Suillia bicolor (Zetterstedt, 1838)
Suillia femoralis (Loew, 1862)
Suillia flava (Meigen, 1830)
Suillia flavifrons (Zetterstedt, 1838)
= nudipes (Czerny, 1932)
Suillia fuscicornis (Zetterstedt, 1847)
Suillia humilis (Meigen, 1830)
= inornata (Loew, 1862)
Suillia laevifrons (Loew, 1862)
Suillia lineitergum (Pandellé, 1901)
= stroblii (Czerny, 1904)
Suillia lurida (Meigen, 1830)
Suillia mikii (Pokorny, 1886)
Suillia nemorum (Meigen, 1830)
Suillia pallida (Fallén, 1820)
Suillia parva (Loew, 1862)
= collini Hackman, 1972
= flavifrons auct. nec (Zetterstedt, 1838)
Suillia quadrilineata Czerny, 1924
Suillia vaginata (Loew, 1862)
TRIXOSCELIDINAE Hendel, 1916
TRIXOSCELIS Rondani, 1856
Trixoscelis frontalis (Fallén, 1823)
? = canescens misid. (see Notes)
Trixoscelis marginella (Fallén, 1823)
Trixoscelis obscurella (Fallén, 1823)
Trixoscelis similis Hackman, 1970
Notes
Chyromya oppidana (Scopoli, 1763). Found only inside houses and farm buildings in Finland.
Orbellia nivicola Frey, 1913. This species was synonymized with Orbellia myiopiformis R.-D. by Storå (1958), but Frey (1958) defended its validity. The status of Orbellia nivicola as a species needs verification.
Trixoscelis canescens (Loew, 1865). This species was originally described on the basis of a single female. Soós (1979) examined the type and revived the name from synonymy with Trixoscelis frontalis. Woźnica (2008) provided an illustrated diagnosis for Trixoscelis canescens and synonymized Trixoscelis gigans Carles-Tolrá, 2001 and Trixoscelis fumipennis Papp, 2005 with it. The species was recently recorded from Finland by Flinck and Kahanpää (2013). Specimens with darkened costal veins and dorsal abdominal surfaces, both proposed diagnostic characters of Trixoscelis canescens, are common among Finnish Trixoscelis frontalis material (see Fig. 8 in Flinck and Kahanpää 2013). Finnish males with these features have genitalia identical with those illustrated for Trixoscelis frontalis by Hackman (1970) and quite unlike the genitalia of Trixoscelis gigans (= fumipennis Papp). The male specimen mentioned in Flinck and Kahanpää (2013) was later dissected and it belongs to Trixoscelis frontalis. The external characters (darkened costa and dorsum of abdomen) can not be used to reliably separate Trixoscelis canescens from Trixoscelis frontalis. The Finnish records of Trixoscelis canescens are probably all misidentifications of Trixoscelis frontalis.
Citation
Kahanpää J (2014) Checklist of the fly families Chyromyidae and Heleomyzidae (Diptera) of Finland. In: Kahanpää J, Salmela J (Eds) Checklist of the Diptera of Finland. ZooKeys 441: 319–324. doi: 10.3897/zookeys.441.7507
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