Abstract Abstract
A checklist of the Braulidae (1 species), Camillidae (3 spp.), Diastatidae (10 spp.) and Drosophilidae (67 spp.) of Finland is presented. Campichoetinae is included as a subfamily of Diastatidae.
Keywords: Checklist, Finland, Diptera, biodiversity, faunistics
Introduction
The Ephydroidea is now recognized as a well-supported, probably monophyletic clade of acalyptrate flies (Wiegmann et al. 2011), although the correct placement of some of the included families – especially the very unsual bee louses of family Braulidae – is still being discussed. The relationships of the families within the superfamily are less understood and the families are listed alphabetically in the checklist below. Seven or eight families are currently included in Ephydroidea. Five of them are found in Finland. Four families are covered in this paper; the fifth (Ephydridae itself) has its own treatment in this volume of ZooKeys. The families missing from Finland are the Curtonotidae, Cryptochetidae, and Nannodastiidae with uncertain affinities (Papp and Mathis 2001). Griffiths (1972) proposed a new family, Campichoetidae, for Campichoeta Macquart, 1835. We follow the world catalogue of Diastatidae (Mathis and Barraclough 2011) and keep Campichoetinae as a subfamily of Diastatidae.
World catalogues have been recently published for Diastatidae (Mathis and Barraclough 2011) and Drosophilidae (Brake and Bächli 2008). The North European Drosophilidae were treated in detail by Bächli et al. (2004) and the review of Finnish Drosophila by Hackman (1954) is now obsolete. The Finnish species of Braulidae and Camillidae were last listed by Hackman (1980); the diastatids were treated by Chandler (1987).
Table 1.
Number of species by family.
| Family | Number of species in | Level of knowledge | ||
|---|---|---|---|---|
| World (Pape et al. 2011) | Europe | Finland | ||
| Braulidae | 7 | 3 | 1 | poor |
| Camillidae | 40 | 8 | 3 | average |
| Diastatidae | 49 (Mathis and Barraclough 2011) | 16 | 10 | good |
| Drosophilidae | 4003 | 121 | 67 | good |
Checklist
suborder Brachycera Macquart, 1834
clade Eremoneura Lameere, 1906
clade Cyclorrhapha Brauer, 1863
infraorder Schizophora Becher, 1882
clade Muscaria Enderlein, 1936
parvorder Acalyptratae Macquart, 1835
superfamily Ephydroidea Zetterstedt, 1837
BRAULIDAE Egger, 1853
BRAULA Nitzsch, 1818
Braula coeca Nitzsch, 1818
CAMILLIDAE Frey, 1921
CAMILLA Haliday, 1838
Camilla atrimana Strobl, 1910
= atripes Duda, 1934
= acutipennis misid.
Camilla flavicauda Duda, 1922
= glabrata Collin, 1956
Camilla glabra (Fallén, 1823)
= subfuscipes Collin, 1933
DIASTATIDAE Hendel, 1917
CAMPICHOETINAE Griffiths, 1972
CAMPICHOETA Macquart, 1835
Campichoeta griseola (Zetterstedt, 1855)
Campichoeta obscuripennis (Meigen, 1830)
DIASTATINAE Hendel, 1917
DIASTATA Meigen, 1830
Diastata adusta Meigen, 1830
= unipunctata Zetterstedt, 1847
Diastata boreonigra Chandler, 1987
Diastata costata Meigen, 1830
Diastata flavicosta Chandler, 1987
Diastata fuscula (Fallén, 1820)
Diastata nebulosa (Fallén, 1823)
Diastata ornata Meigen, 1830
Diastata vagans Loew, 1864
DROSOPHILIDAE Rondani, 1856
STEGANINAE Hendel, 1917
AMIOTA Loew, 1862
sg. Amiota Loew, 1862
Amiota albilabris (Roth in Zetterstedt, 1860)
Amiota alboguttata (Wahlberg, 1839)
Amiota rufescens (Oldenberg, 1914)
Amiota subtusradiata Duda, 1934
CACOXENUS Loew, 1858
sg. Paracacoxenus Hardy & Wheeler, 1960
Cacoxenus argyreator Frey, 1932
LEUCOPHENGA Mik, 1886
sg. Neoleucophenga Oldenberg, 1915
Leucophenga quinquemaculata Strobl, 1893
PHORTICA Schiner, 1862
sg. Phortica Schiner, 1862
Phortica variegata Fallén, 1823
STEGANA Meigen, 1830
sg. Stegana Meigen, 1830
Stegana furta (Linnaeus, 1766)
= curvipennis (Fallén, 1823)
sg. Steganina Wheeler, 1960
Stegana baechlii Laštovka & Máca, 1982
Stegana coleoptrata (Scopoli, 1763)
Stegana hypoleuca Meigen, 1830
Stegana longifibula Takada, 1968
Stegana mehadiae Duda, 1924
Stegana nigrithorax Strobl, 1898
Stegana similis Laštovka & Máca, 1982
DROSOPHILINAE Rondani, 1856
CHYMOMYZA Czerny, 1903
Chymomyza amoena (Loew, 1862)
Chymomyza caudatula Oldenberg, 1914
Chymomyza costata (Zetterstedt, 1838)
Chymomyza distincta (Egger, 1862)
Chymomyza fuscimana (Zetterstedt, 1838)
DROSOPHILA Fallén, 1823
sg. Dorsilopha Sturtevant, 1942
Drosophila busckii Coquillett, 1901
sg. Drosophila Fallén, 1823
Drosophila ezoana Takada & Okada, 1957
Drosophila funebris (Fabricius, 1787)
Drosophila histrio Meigen, 1830
Drosophila hydei Sturtevant, 1921
= repleta misid.
Drosophila immigrans Sturtevant, 1921
= tripunctata Becker, 1908 preocc.
Drosophila limbata von Roser, 1840
Drosophila littoralis Meigen, 1830
Drosophila lummei Hackman, 1972
Drosophila montana Stone, Griffen & Patterson, 1941
= ovivororum Lakovaara & Hackman, 1973
Drosophila phalerata Meigen, 1830
Drosophila picta Zetterstedt, 1847
Drosophila repleta Wollaston, 1858
Drosophila subarctica Hackman, 1969
Drosophila testacea von Roser, 1840
Drosophila transversa Fallén, 1823
Drosophila vireni Bächli, Vilela & Haring, 2002
sg. Sophophora Sturtevant, 1942
Drosophila alpina Burla, 1948
Drosophila ambigua Pomini, 1940
Drosophila bifasciata Pomini, 1940
Drosophila eskoi Lakovaara & Lankinen, 1974
Drosophila ingrica Hackman, 1957
Drosophila melanogaster Meigen, 1830
Drosophila obscura Fallén, 1823
Drosophila simulans Sturtevant, 1919
Drosophila subobscura Collin, 1936
Drosophila subsilvestris Hardy & Kaneshiro, 1968
= silvestris Basden, 1954 preocc.
Drosophila tristis Fallén, 1823
HIRTODROSOPHILA Duda, 1924
Hirtodrosophila cameraria (Haliday, 1833)
Hirtodrosophila confusa (Staeger, 1844)
Hirtodrosophila lundstroemi (Duda, 1935)
Hirtodrosophila oldenbergi (Duda, 1924)
Hirtodrosophila trivittata (Strobl, 1893)
LORDIPHOSA Basden, 1961
Lordiphosa fenestrarum (Fallén, 1823)
Lordiphosa nigricolor (Strobl, 1898)
MICRODROSOPHILA Malloch, 1921
sg. Microdrosophila Malloch, 1921
Microdrosophila congesta (Zetterstedt, 1847)
sg. Oxystyloptera Duda, 1924
Microdrosophila zetterstedti Wheeler, 1959
= nigriventris (Zetterstedt, 1847) preocc.
SCAPTODROSOPHILA Duda, 1923
Scaptodrosophila deflexa (Duda, 1924)
= guyenoti (Burla, 1948)
SCAPTOMYZA Hardy, 1849
sg. Hemiscaptomyza Hackman, 1959
Scaptomyza trochanterata Collin, 1953
Scaptomyza unipunctum (Zetterstedt, 1847)
sg. Parascaptomyza Duda, 1924
Scaptomyza pallida (Zetterstedt, 1847)
= disticha (Duda, 1921)
sg. Scaptomyza Hardy, 1849
Scaptomyza consimilis Hackman, 1955
Scaptomyza flava (Fallén, 1823)
= flaveola (Meigen, 1830)
= apicalis Hardy, 1849
Scaptomyza graminum (Fallén, 1823)
Scaptomyza griseola (Zetterstedt, 1847)
Scaptomyza montana Wheeler, 1949
Scaptomyza teinoptera Hackman, 1955
Excluded species
Stegana strobli Mik, 1898 misidentification
Lordiphosa hexasticha (Papp, 1971) not found within present borders
Citation
Kahanpää J (2014) Checklist of the Braulidae, Camillidae, Diastatidae and Drosophilidae of Finland (Diptera, Ephydroidea). In: Kahanpää J, Salmela J (Eds) Checklist of the Diptera of Finland. ZooKeys 441: 333–338. doi: 10.3897/zookeys.441.7057
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