Notes on Shore Flies (Diptera: Ephydridae) from Finland and north-western Russia

Jere Kahanpää; Tadeusz Zatwarnicki

doi:10.3897/BDJ.3.e4701

. 2015 Mar 30;(3):e4701. doi: 10.3897/BDJ.3.e4701

Notes on Shore Flies (Diptera: Ephydridae) from Finland and north-western Russia

Jere Kahanpää ^‡,^✉, Tadeusz Zatwarnicki ^§

PMCID: PMC4385901 PMID: 25859131

Abstract Abstract

The recent checklist of the Ephydridae of Finland by Zatwarnicki and Kahanpää (2014) mentioned 13 ephydrid species as new for Finland without further details. This paper presents detailed records for those species and a few other species of interest. Four species are recorded for the first time from Russia. Trimerina indistincta Krivosheina, 2004 is herein considered as a new junior synonym of Trimerina microchaeta Hendel, 1932, syn. nov.

Keywords: New records, new synonym, Trimerina , Scatophila iowana, faunistics

Introduction

Finland is an exceptional country in having a record of regularly published checklists of the order Diptera, including shore flies. Three of such lists have been published during the last century: Frey (1941), Hackman (1980) and Kahanpää and Salmela (2014). The Diptera fauna of the country is well studied in comparison to most other countries, but far from perfectly understood.

The most recent list of the Finnish shore fly fauna is Zatwarnicki and Kahanpää (2014) in Kahanpää and Salmela (2014). The Finnish Ephydridae have attracted relatively little attention: few Finnish dipterologists have studied them and no coordinated study of the fauna has been made. The Finnish Museum of Natural History in Helsinki (MZH) holds by far the largest ephydrid collection in the country. Existing faunal works are essentially based on museum material collected between the years 1900 and 1960, mostly by sweep-netting. After a long hiatus, interest in acalyptrate flies resurged in the 21st century among Finnish amateur entomologists. The newer ephydrid material generally remains in private collections.

Zatwarnicki (1997) included a list of the species known from the country by the mid-1990s. Twelve species have been added to the fauna after 1997. Chronologically first is Notiphila pollinosa Krivosheina by Krivosheina (1998), then Calocoenia paurosoma (Sturtevant & Wheeler) and Parydra arctica Clausen by Krivosheina (2000), Coenia curvicauda (Meigen) by Haarto and Winqvist (2002), Gymnoclasiopa pulchella (Meigen) by Winqvist (2002), Notiphila major Stenhammar by Kahanpää and Winqvist (2003), Trimerina microchaeta Hendel (as T. indistincta Krivosheina) by Krivosheina (2004), Philotelma defectum (Haliday) by Mathis et al. (2009), Atissa pygmaea (Haliday) by Winqvist (2011), and Hecamedoides unispinosus (Collin), Parydra mitis (Cresson) and P. nigritarsis Strobl by Kahanpää (2013). The identity of the North European Glenanthe species was resolved by Zatwarnicki and Mathis (2012).

Thirteen additional species of Ephydridae were recorded for the first time from Finland in the recent checklist (Zatwarnicki and Kahanpää 2014). The checklist format did not allow for the inclusion of record data. This paper provides the previously unpublished records on which the checklist was based. Additionally, one new synonymy is proposed: Trimerina indistincta Krivosheina, 2004 is treated as a junior synonym of Trimerina microchaeta Hendel, 1932 (syn. nov.). After inclusion of all additions and corrections to the fauna, the number of shore fly species recorded from Finland is now 112.

Some material from north-western Russia, adjacent to the Russian-Finnish border, was also examined during the preparation of the Finnish checklist. Four species are recorded from Russia for the first time.

Materials and methods

All the specimens discussed in this paper are deposited at the Zoological Museum of the Finnish Museum of Natural History in Helsinki (MZH) unless otherwise specified. Most MZH specimens have unique specimen identifiers in the form of a URL (Uniform Resource Locator). The URL can be used as a web address: it points to a web page with details of the sample in question. While we cannot guarantee that the web content is available in the future, the address (URL) will remain unique. Standard abbreviations are used for the Finnish biogeographical provinces; see the Fauna Entomologica Scandinavica book series or Heikinheimo and Raatikainen (1971) and Heikinheimo and Raatikainen (1981) for details and maps.

Finnish insect collectors usually use a national uniform grid system (yhtenäiskoordinaatisto, YKJ) for mapping and labeling purposes. The Darwin Core (DC) field "verbatimCoordinates" is used for the YKJ coordinates printed on labels. The DC fields "decimalLatitude" and "decimalLongitude" in the Taxon treatments section are given in World Geodetic System 1984 (WGS84) decimal degrees. Where no collector-supplied coordinates were available, the records were georeferenced by the senior author on the basis of the collecting locality.

The identity of all included species is based on the study of primary types conducted by the junior author, except for Hydrellia tarsata Haliday and Scatophila mesogramma (Loew), for which interpretations are based on Collin (1966) and Sturtevant and Wheeler (1953) respectively.

The order of species follows Zatwarnicki and Kahanpää (2014).

Taxon treatments

Trimerina microchaeta

Hendel, 1932

Trimerina microchaeta Hendel 1932: 11
= Trimerina indistincta Krivosheina 2004: 631 (syn. nov.)

Materials

Type status: Other material. Occurrence: catalogNumber: JKA12-0653; recordedBy: Frey, Richard; individualCount: 1; sex: M; lifeStage: adult; associatedSequences: BOLD:2709504; Taxon: scientificName: Trimerina microchaeta Hendel, 1932; order: Diptera; family: Ephydridae; genus: Trimerina; specificEpithet: microchaeta; scientificNameAuthorship: Hendel, 1932; Location: country: Finland; stateProvince: EH; municipality: Hartola; locality: Hirtesalo; verbatimElevation: 80; verbatimCoordinates: 6839:[3]450; decimalLatitude: 61.66; decimalLongitude: 26.07; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 2012; Event: year: 2002; month: 5; day: 15; habitat: a lake shore; Record Level: institutionCode: Coll. J. Kahanpää; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: JKA12-0654; recordedBy: Frey, Richard; individualCount: 1; sex: F; lifeStage: adult; associatedSequences: BOLD:2709505; Taxon: scientificName: Trimerina microchaeta Hendel, 1932; order: Diptera; family: Ephydridae; genus: Trimerina; specificEpithet: microchaeta; scientificNameAuthorship: Hendel, 1932; Location: country: Finland; stateProvince: ES; municipality: Joutsa; locality: Oravakivensalmi; verbatimElevation: 80; verbatimCoordinates: 6845:[3]451; decimalLatitude: 61.71; decimalLongitude: 26.08; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 2012; Event: year: 2004; month: 6; day: 3; Record Level: institutionCode: Coll. J. Kahanpää; basisOfRecord: PreservedSpecimen

Distribution

Widespread in Europe; also occurs in the Russian Far East (Kamchatka) (Zatwarnicki 2013).

Taxon discussion

After examination of primary types of Trimerina microchaeta Hendel and T. indistincta Krivosheina we suggest to synonymize these two species. Krivosheina (2004) provided three characters to separate T. indistincta from T. microchaeta: a gena-to-eye ratio of 0.17 (versus 0.20 for T. microchaeta), yellow mid and hind coxae (versus all coxae partially dark), mid and hind tibiae entirely yellow (versus mid and hind tibiae with a dark band). These characters separating the two species are not consistent and fall within the variation range of T. microchaeta. Both nominal taxa have essentially the same male terminalia with characteristic postgonostyli (=postsurstyli) bearing short and apically rounded antero-ventral processes as illustrated for T. indistincta by Krivosheina (2004).

Notes

The holotype of Trimerina indistincta is shown in Fig. 1 a.

Figure 1a. — Ephydrid species from Finland. Scale bar = 1.0 mm.

Figure 1b. — Ephydrid species from Finland. Scale bar = 1.0 mm.

Hydrellia albifrons

Fallén, 1813

Materials

Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.16614; recordedBy: Tiensuu, Lauri; individualCount: 1; sex: M; lifeStage: adult; preparations: gen. prep. on pin; Taxon: scientificName: Hydrellia albifrons Fallén, 1813; order: Diptera; family: Ephydridae; genus: Hydrellia; specificEpithet: albifrons; scientificNameAuthorship: Fallén, 1813; Location: country: Finland; stateProvince: V; municipality: Naantali; decimalLatitude: 60.47; decimalLongitude: 22.47; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 20000; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; Event: year: 1939; month: 8; day: 16; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.16615; recordedBy: Tiensuu, Lauri; individualCount: 1; sex: F; lifeStage: adult; Taxon: scientificName: Hydrellia albifrons Fallén, 1813; order: Diptera; family: Ephydridae; genus: Hydrellia; specificEpithet: albifrons; scientificNameAuthorship: Fallén, 1813; Location: country: Finland; stateProvince: V; municipality: Naantali; decimalLatitude: 60.13; decimalLongitude: 22.02; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 20000; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; Event: year: 1949; month: 7; day: 1; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen

Diagnosis

Best identified by structures of the male genitalia as figured by Collin (1966) and Canzoneri and Meneghini (1983).

Distribution

First recorded from Finland by Zatwarnicki and Kahanpää (2014). Widespread in Europe; also known from North Africa and Pakistan (Zatwarnicki 1988, Zatwarnicki 2013).

Ecology

Larvae of this species feed on pondweeds (Potamogeton) (Zatwarnicki 1988), the common frogbit (Hydrocharis morsus-ranae) (Hering 1926), and possibly also on water plantains (Alisma plantago-aquatica) (see Deonier 1971).

Hydrellia cochleariae

Haliday, 1839

Materials

Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.16690; recordedBy: Palmén, Johan Axel; individualCount: 4; sex: MF; lifeStage: adult; Taxon: scientificName: Hydrellia cochleariae Haliday, 1839; order: Diptera; family: Ephydridae; genus: Hydrellia; specificEpithet: cochleariae; scientificNameAuthorship: Haliday, 1839; Location: country: Finland; stateProvince: U; municipality: Helsinki; locality: Jollas; verbatimLocality: Helsinge; decimalLatitude: 60.16; decimalLongitude: 25.07; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; Event: year: 1864; month: 6; day: 20; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.16691; recordNumber: 4096; recordedBy: Palmén, Johan Axel; individualCount: 5; sex: MF; lifeStage: adult; preparations: gen. prep. on pin; Taxon: scientificName: Hydrellia cochleariae Haliday, 1839; order: Diptera; family: Ephydridae; genus: Hydrellia; specificEpithet: cochleariae; scientificNameAuthorship: Haliday, 1839; Location: country: Finland; stateProvince: U; municipality: Helsinki; locality: Jollas; verbatimLocality: Helsinge; decimalLatitude: 60.16; decimalLongitude: 25.07; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; Event: year: 1864; month: 6; day: 20; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.16690; recordedBy: Meinander, Martin; individualCount: 1; sex: MF; lifeStage: adult; preparations: gen. prep. on pin; Taxon: scientificName: Hydrellia cochleariae Haliday, 1839; order: Diptera; family: Ephydridae; genus: Hydrellia; specificEpithet: cochleariae; scientificNameAuthorship: Haliday, 1839; Location: country: Finland; stateProvince: V; municipality: Korppoo; locality: Jurmo; decimalLatitude: 59.82; decimalLongitude: 21.60; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; Event: year: 1960; month: 8; day: 8; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen

Materials

Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19147; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Finland; stateProvince: EH; municipality: Kangasala; verbatimLocality: Kangasala; decimalLatitude: 61.496; decimalLongitude: 24.0754; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19224; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Finland; stateProvince: EH; municipality: Kangasala; verbatimLocality: Kangasala; decimalLatitude: 61.496; decimalLongitude: 24.0754; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19225; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Finland; stateProvince: EH; municipality: Kangasala; verbatimLocality: Kangasala; decimalLatitude: 61.496; decimalLongitude: 24.0754; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19226; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Finland; stateProvince: EH; municipality: Kangasala; verbatimLocality: Kangasala; decimalLatitude: 61.496; decimalLongitude: 24.0754; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19227; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Finland; stateProvince: V; municipality: Lohja; verbatimLocality: Lojo; decimalLatitude: 60.237; decimalLongitude: 24.0078; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19228; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Finland; stateProvince: V; municipality: Lohja; verbatimLocality: Lojo; decimalLatitude: 60.237; decimalLongitude: 24.0078; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19229; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Russia; municipality: Metsäpirtti; verbatimLocality: Metsäpirtti; decimalLatitude: 60.56; decimalLongitude: 30.52; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19230; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Russia; municipality: Metsäpirtti; verbatimLocality: Metsäpirtti; decimalLatitude: 60.56; decimalLongitude: 30.52; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19231; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Russia; municipality: Metsäpirtti; verbatimLocality: Metsäpirtti; decimalLatitude: 60.56; decimalLongitude: 30.52; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19232; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Russia; municipality: Metsäpirtti; verbatimLocality: Metsäpirtti; decimalLatitude: 60.56; decimalLongitude: 30.52; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19233; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Russia; municipality: Metsäpirtti; verbatimLocality: Metsäpirtti; decimalLatitude: 60.56; decimalLongitude: 30.52; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19234; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Russia; municipality: Metsäpirtti; verbatimLocality: Metsäpirtti; decimalLatitude: 60.56; decimalLongitude: 30.52; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19235; recordedBy: Tiensuu, Lauri; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Russia; municipality: Vaaseni; verbatimLocality: Terijoki; decimalLatitude: 60.96; decimalLongitude: 34.02; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Event: year: 1939; month: 6; day: 8; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19236; recordedBy: Tiensuu, Lauri; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Russia; municipality: Vaaseni; verbatimLocality: Vaaseni; decimalLatitude: 60.96; decimalLongitude: 34.02; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Event: year: 1942; month: 6; day: 15; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19237; recordedBy: Tiensuu, Lauri; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Russia; municipality: Terijoki; verbatimLocality: Terijoki; decimalLatitude: 60.18; decimalLongitude: 29.7; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Event: year: 1939; month: 6; day: 8; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19238; recordedBy: Tiensuu, Lauri; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Russia; municipality: Terijoki; verbatimLocality: Terijoki; decimalLatitude: 60.18; decimalLongitude: 29.7; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Event: year: 1939; month: 6; day: 8; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19239; recordedBy: Tiensuu, Lauri; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Russia; municipality: Terijoki; verbatimLocality: Terijoki; decimalLatitude: 60.18; decimalLongitude: 29.7; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Event: year: 1939; month: 6; day: 8; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19240; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Russia; municipality: Metsäpirtti; verbatimLocality: Metsäpirtti; decimalLatitude: 60.56; decimalLongitude: 30.52; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19241; recordedBy: Tiensuu, Lauri; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Russia; municipality: Terijoki; verbatimLocality: Terijoki; decimalLatitude: 60.18; decimalLongitude: 29.7; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Event: year: 1939; month: 6; day: 8; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19242; recordedBy: Tiensuu, Lauri; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Russia; municipality: Terijoki; verbatimLocality: Terijoki; decimalLatitude: 60.18; decimalLongitude: 29.7; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Event: year: 1939; month: 6; day: 8; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19243; recordedBy: Tiensuu, Lauri; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Finland; stateProvince: V; municipality: Lohja; verbatimLocality: Lojo; decimalLatitude: 60.237; decimalLongitude: 24.0078; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19244; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Russia; municipality: Vaaseni; verbatimLocality: Vaaseni; decimalLatitude: 60.96; decimalLongitude: 34.02; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Event: year: 1942; month: 6; day: 15; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19245; recordedBy: Tiensuu, Lauri; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Russia; municipality: Terijoki; verbatimLocality: Terijoki; decimalLatitude: 60.18; decimalLongitude: 29.7; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Event: year: 1939; month: 6; day: 11; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19246; recordedBy: Tiensuu, Lauri; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Russia; municipality: Vaaseni; verbatimLocality: Vaaseni; decimalLatitude: 60.96; decimalLongitude: 34.02; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Event: year: 1942; month: 6; day: 1; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19247; recordedBy: Tiensuu, Lauri; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Finland; stateProvince: V; municipality: Lohja; verbatimLocality: Lojo; decimalLatitude: 60.237; decimalLongitude: 24.0078; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19248; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Finland; stateProvince: V; municipality: Lohja; verbatimLocality: Lojo; decimalLatitude: 60.237; decimalLongitude: 24.0078; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19249; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Finland; stateProvince: V; municipality: Lohja; verbatimLocality: Lojo; decimalLatitude: 60.237; decimalLongitude: 24.0078; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19250; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Finland; stateProvince: V; municipality: Lohja; verbatimLocality: Lojo; decimalLatitude: 60.237; decimalLongitude: 24.0078; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19251; recordedBy: Tiensuu, Lauri; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Finland; stateProvince: U; municipality: Helsinki; locality: Botanical gardens; verbatimLocality: Helsinki, Hortus Bot.; decimalLatitude: 60.174; decimalLongitude: 24.9508; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Event: year: 1939; month: 6; day: 5; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19252; recordedBy: Storå, Ragnar; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Finland; stateProvince: KP; municipality: Pedersöre; verbatimLocality: Pedersöre; decimalLatitude: 63.531; decimalLongitude: 22.8625; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Event: year: 1958; month: 6; day: 14; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19253; recordedBy: Storå, Ragnar; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Finland; stateProvince: KP; municipality: Pedersöre; verbatimLocality: Pedersöre; decimalLatitude: 63.531; decimalLongitude: 22.8625; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Event: year: 1952; month: 6; day: 28; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19254; recordedBy: Storå, Ragnar; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Finland; stateProvince: KP; municipality: Pedersöre; verbatimLocality: Pedersöre; decimalLatitude: 63.531; decimalLongitude: 22.8625; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Event: year: 1952; month: 6; day: 28; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19255; recordedBy: Storå, Ragnar; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Finland; stateProvince: KP; municipality: Pedersöre; verbatimLocality: Pedersöre; decimalLatitude: 63.531; decimalLongitude: 22.8625; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Event: year: 1952; month: 6; day: 28; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19256; recordedBy: Storå, Ragnar; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Finland; stateProvince: KP; municipality: Pedersöre; verbatimLocality: Pedersöre; decimalLatitude: 63.531; decimalLongitude: 22.8625; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Event: year: 1952; month: 6; day: 28; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19257; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Finland; stateProvince: V; municipality: Vihti; verbatimLocality: Vichtis; decimalLatitude: 60.424; decimalLongitude: 24.3539; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19258; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Finland; stateProvince: EH; municipality: Kangasala; verbatimLocality: Kangasala; decimalLatitude: 61.496; decimalLongitude: 24.0754; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19259; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family: Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus; scientificNameAuthorship: (Collin, 1943); Location: country: Finland; stateProvince: EH; municipality: Kangasala; verbatimLocality: Kangasala; decimalLatitude: 61.496; decimalLongitude: 24.0754; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, Th.; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen

Diagnosis

Two of the North European Hecamedoides species can be identified by the number of short, blunt anteroventral spine-like setae on the fore femur: one in H. unispinosus, several in H. glaucellus (Stenhammar, 1844). Hecamedoides kelmorum Stuke, 2011, a species recently described from Germany, also has a black antenna and a single anteroventral spine (Stuke 2011). Hecamedoides unispinosus and H. kelmorum are best identified by structures of the male genitalia. Two Finnish males (http://id.luomus.fi/GV.19243 and http://id.luomus.fi/GV.19235) were dissected to confirm their identification. The gonite of these specimens is clearly of the H. unispinosus type.

Distribution

A Holarctic species. Recorded from Finland by Kahanpää 2013. New for Russia.

Taxon discussion

Hecamedoides unispinosus is much more common in Finland than H. glaucellus. The Fennica collection of MZH has only three specimens of the latter species: http://id.luomus.fi/GV.19146 from Turku (see Fig. 2 b) and two specimens from Terijoki (=Russia, Leningrad Oblast, Zelenogorsk).

Figure 2a. — *Hecamedoides* species from Finland. Scale bar = 1.0 mm.

Figure 2b. — *Hecamedoides* species from Finland. Scale bar = 1.0 mm.

Pelina norvegica

Dahl, 1975

Materials

Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19162; recordedBy: Frey, Richard; individualCount: 1; sex: M; lifeStage: adult; Taxon: scientificName: Pelina norvegica Dahl, 1975; order: Diptera; family: Ephydridae; genus: Pelina; specificEpithet: norvegica; scientificNameAuthorship: Dahl, 1975; Location: country: Russia; stateProvince: Murmansk Oblast; locality: Yläluostari; verbatimLocality: Petsamo; decimalLatitude: 69.4; decimalLongitude: 31.11; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 09/26/2013; Event: year: 1930; month: 7; day: 12; habitat: riverside; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19163; recordedBy: Frey, Richard; individualCount: 1; sex: M; lifeStage: adult; Taxon: scientificName: Pelina norvegica Dahl, 1975; order: Diptera; family: Ephydridae; genus: Pelina; specificEpithet: norvegica; scientificNameAuthorship: Dahl, 1975; Location: country: Russia; stateProvince: Murmansk Oblast; locality: Yläluostari; verbatimLocality: Petsamo; decimalLatitude: 69.4; decimalLongitude: 31.11; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 09/26/2013; Event: year: 1930; month: 7; day: 12; habitat: riverside; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen

Distribution

Previously recorded from Norway, Germany and Slovakia (Zatwarnicki 2013). New for Russia.

Notes

Specimen http://id.luomus.fi/GV.19162 of Pelina norvegica is shown in Fig. 1 d.

Parydra (Chaetoapnea) mitis

(Cresson, 1930)

Materials

Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.16564; recordedBy: Albrecht, Anders; individualCount: 1; lifeStage: adult; Taxon: scientificName: Parydra mitis (Cresson, 1930); order: Diptera; family: Ephydridae; genus: Parydra; specificEpithet: mitis; scientificNameAuthorship: (Cresson, 1930); Location: country: Finland; stateProvince: U; municipality: Sipoo; locality: Sjöskog; verbatimCoordinates: 6690:3385; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, T.; Event: year: 1975; month: 5; day: 6; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.16565; recordedBy: Tiensuu, Lauri; individualCount: 1; lifeStage: adult; Taxon: scientificName: Parydra mitis (Cresson, 1930); order: Diptera; family: Ephydridae; genus: Parydra; specificEpithet: mitis; scientificNameAuthorship: (Cresson, 1930); Location: country: Russia; municipality: Vaaseni; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, T.; Event: year: 1942; month: 5; day: 2; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.16566; recordedBy: Tiensuu, Lauri; individualCount: 1; lifeStage: adult; Taxon: scientificName: Parydra mitis (Cresson, 1930); order: Diptera; family: Ephydridae; genus: Parydra; specificEpithet: mitis; scientificNameAuthorship: (Cresson, 1930); Location: country: Finland; stateProvince: U; municipality: Helsinki; locality: Herttoniemi; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, T.; Event: year: 1945; month: 5; day: 16; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.16567; recordedBy: Tiensuu, Lauri; individualCount: 1; lifeStage: adult; Taxon: scientificName: Parydra mitis (Cresson, 1930); order: Diptera; family: Ephydridae; genus: Parydra; specificEpithet: mitis; scientificNameAuthorship: (Cresson, 1930); Location: country: Finland; stateProvince: EK; municipality: Vehkalahti; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, T.; Event: year: 1945; month: 4; day: 9; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.16568; recordedBy: Tiensuu, Lauri; individualCount: 1; lifeStage: adult; Taxon: scientificName: Parydra mitis (Cresson, 1930); order: Diptera; family: Ephydridae; genus: Parydra; specificEpithet: mitis; scientificNameAuthorship: (Cresson, 1930); Location: country: Finland; stateProvince: U; municipality: Helsinki; locality: Herttoniemi; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, T.; Event: year: 1945; month: 5; day: 16; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.16569; recordedBy: Lundström, Carl; individualCount: 1; lifeStage: adult; Taxon: scientificName: Parydra mitis (Cresson, 1930); order: Diptera; family: Ephydridae; genus: Parydra; specificEpithet: mitis; scientificNameAuthorship: (Cresson, 1930); Location: country: Finland; stateProvince: PS; municipality: Maaninka; locality: Tuovilanlahti; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, T.; Event: year: 1865; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.16570; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Parydra mitis (Cresson, 1930); order: Diptera; family: Ephydridae; genus: Parydra; specificEpithet: mitis; scientificNameAuthorship: (Cresson, 1930); Location: country: Russia; stateProvince: Republic of Carelia; municipality: Paanajärvi; locality: behind Takalo farm; decimalLatitude: 66.26; decimalLongitude: 29.78; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, T.; Event: year: 1939; month: 17; day: 6; habitat: lake shore; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.16571; recordedBy: Storå, Ragnar; individualCount: 1; lifeStage: adult; Taxon: scientificName: Parydra mitis (Cresson, 1930); order: Diptera; family: Ephydridae; genus: Parydra; specificEpithet: mitis; scientificNameAuthorship: (Cresson, 1930); Location: country: Finland; stateProvince: KP; municipality: Uusikaarlepyy; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, T.; Event: year: 1954; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.16572; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Parydra mitis (Cresson, 1930); order: Diptera; family: Ephydridae; genus: Parydra; specificEpithet: mitis; scientificNameAuthorship: (Cresson, 1930); Location: country: Russia; stateProvince: Republic of Carelia; municipality: Paanajärvi; locality: hotel; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, T.; Event: year: 1939; month: 7; day: 6; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.16573; recordedBy: Storå, Ragnar; individualCount: 1; lifeStage: adult; Taxon: scientificName: Parydra mitis (Cresson, 1930); order: Diptera; family: Ephydridae; genus: Parydra; specificEpithet: mitis; scientificNameAuthorship: (Cresson, 1930); Location: country: Finland; stateProvince: KP; municipality: Uusikaarlepyy; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, T.; Event: year: 1954-1955; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.16574; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Parydra mitis (Cresson, 1930); order: Diptera; family: Ephydridae; genus: Parydra; specificEpithet: mitis; scientificNameAuthorship: (Cresson, 1930); Location: country: Finland; stateProvince: U; municipality: Helsinki; locality: Munkkiniemi; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Zatwarnicki, T.; Event: year: 1941; month: 5; day: 4; habitat: a herb-rich forest with a stream; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen

Distribution

First recorded from Finland by Zatwarnicki and Kahanpää (2014). Widespread in Europe, it is also found in North Africa (Zatwarnicki 2013). The illustrated record by Kahanpää (2013) represents Parydra nigritarsis Strobl, 1893.

Notes

Fig. 3 c illustrates specimen http://id.luomus.fi/GV.16525.

Figure 3a. — *Parydra* species from Finland. Scale bar = 1.0 mm.

Figure 3b. — *Parydra* species from Finland. Scale bar = 1.0 mm.

Calocoenia paurosoma

(Sturtevant & Wheeler, 1954)

Materials

Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.18970; recordNumber: 6643; recordedBy: Storå, Ragnar; individualCount: 1; sex: M; lifeStage: adult; Taxon: scientificName: Calocoenia paurosoma (Sturdevant & Wheeler, 1954); order: Diptera; family: Ephydridae; genus: Calocoenia; specificEpithet: paurosoma; scientificNameAuthorship: (Sturdevant & Wheeler, 1954); Location: country: Finland; stateProvince: EnL; municipality: Enontekiö; locality: Kilpisjärvi; decimalLatitude: 69.0; decimalLongitude: 20.9; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 4000; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Krivosheina, Marina G.; dateIdentified: 2004; Event: year: 1954; month: 8; day: 2; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.16463; recordNumber: 1106; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Calocoenia paurosoma (Sturdevant & Wheeler, 1954); order: Diptera; family: Ephydridae; genus: Calocoenia; specificEpithet: paurosoma; scientificNameAuthorship: (Sturdevant & Wheeler, 1954); Location: country: Finland; stateProvince: EnL; municipality: Enontekiö; locality: Kilpisjärvi, Malla; decimalLatitude: 69.09; decimalLongitude: 20.70; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 4000; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 09/26/2013; Event: year: 1929; month: 7; day: 15; habitat: regio subalpina; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.16457; recordNumber: 392; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Calocoenia paurosoma (Sturdevant & Wheeler, 1954); order: Diptera; family: Ephydridae; genus: Calocoenia; specificEpithet: paurosoma; scientificNameAuthorship: (Sturdevant & Wheeler, 1954); Location: country: Finland; stateProvince: Ks; municipality: Kuusamo; locality: Kuusamo town, Niilo-oja; locationRemarks: N side of Niilo-oja; decimalLatitude: 65.97; decimalLongitude: 29.18; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 500; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 09/26/2013; Event: year: 1917; month: 6; day: 9; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.16459; recordNumber: 1115; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Calocoenia paurosoma (Sturdevant & Wheeler, 1954); order: Diptera; family: Ephydridae; genus: Calocoenia; specificEpithet: paurosoma; scientificNameAuthorship: (Sturdevant & Wheeler, 1954); Location: country: Finland; stateProvince: KemLl; municipality: Kittilä; locality: Alakyrö; decimalLatitude: 68.07; decimalLongitude: 24.39; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 09/26/2013; Event: year: 1911; month: 6; day: 19; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.16456; recordNumber: 1153; recordedBy: Lundström, Carl; individualCount: 1; lifeStage: adult; Taxon: scientificName: Calocoenia paurosoma (Sturdevant & Wheeler, 1954); order: Diptera; family: Ephydridae; genus: Calocoenia; specificEpithet: paurosoma; scientificNameAuthorship: (Sturdevant & Wheeler, 1954); Location: country: Finland; stateProvince: PS; municipality: Nilsiä; decimalLatitude: 63.2; decimalLongitude: 28.1; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 09/26/2013; Event: year: 1865; month: 6; day: 27; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.16462; recordNumber: 712; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Calocoenia paurosoma (Sturdevant & Wheeler, 1954); order: Diptera; family: Ephydridae; genus: Calocoenia; specificEpithet: paurosoma; scientificNameAuthorship: (Sturdevant & Wheeler, 1954); Location: country: Russia; stateProvince: Carelian Republic; municipality: Paanajärvi; locality: bog behind Takalo farm; decimalLatitude: 66.26; decimalLongitude: 29.78; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 2000; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 09/26/2013; Event: year: 1937; month: 6; day: 24; habitat: a bog; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.16458; recordNumber: 673; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Calocoenia paurosoma (Sturdevant & Wheeler, 1954); order: Diptera; family: Ephydridae; genus: Calocoenia; specificEpithet: paurosoma; scientificNameAuthorship: (Sturdevant & Wheeler, 1954); Location: country: Russia; stateProvince: Carelian Republic; municipality: Paanajärvi; locality: bog behind Takalo farm; decimalLatitude: 66.26; decimalLongitude: 29.78; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 2000; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 09/26/2013; Event: year: 1937; month: 6; day: 24; habitat: a bog; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.16460; recordNumber: 2492; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Calocoenia paurosoma (Sturdevant & Wheeler, 1954); order: Diptera; family: Ephydridae; genus: Calocoenia; specificEpithet: paurosoma; scientificNameAuthorship: (Sturdevant & Wheeler, 1954); Location: country: Russia; stateProvince: Murmansk Oblast; municipality: Kantalahti; decimalLatitude: 67.2; decimalLongitude: 32.4; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 09/26/2013; Event: year: 1913; month: 6; day: 28; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.16461; recordNumber: 961; recordedBy: Hellén, Wolter; individualCount: 1; lifeStage: adult; Taxon: scientificName: Calocoenia paurosoma (Sturdevant & Wheeler, 1954); order: Diptera; family: Ephydridae; genus: Calocoenia; specificEpithet: paurosoma; scientificNameAuthorship: (Sturdevant & Wheeler, 1954); Location: country: Russia; stateProvince: Murmansk Oblast; municipality: Kantalahti; decimalLatitude: 67.2; decimalLongitude: 32.4; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 09/26/2013; Event: year: 1913; month: 6; day: 27; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen

Diagnosis

At first glance this species resembles the genus Coenia, but it has five dorsocentral setae. Specimen http://id.luomus.fi/GV.16455 is illustrated in Fig. 4 b, d.

Figure 4a. — *Coenia* and *Calocoenia*. Scale bars 0.5 mm (a-b) and 1.0 mm (c-d).

Figure 4b. — *Coenia* and *Calocoenia*. Scale bars 0.5 mm (a-b) and 1.0 mm (c-d).

Distribution

New for Russia. North Holarctic, known in Europe from Norway, Sweden and Finland (Zatwarnicki 2013). First noted from Finland by Krivosheina (2000) from Kilpisjärvi. The new records presented here expand the known European distribution of the species both south- and eastwards.

Coenia vulgata

Krivosheina, 2001

Materials

Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.16464; recordNumber: 174; recordedBy: Wuorentaus, Yrjö; individualCount: 1; sex: M; lifeStage: adult; preparations: gen. prep. on pin; Taxon: scientificName: Coenia vulgata Krivosheina, 2001; order: Diptera; family: Ephydridae; genus: Coenia; specificEpithet: vulgata; scientificNameAuthorship: Krivosheina, 2001; Location: country: Finland; stateProvince: PPe; municipality: Oulu; decimalLatitude: 65.01; decimalLongitude: 25.41; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 10000; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 09/26/2013; Event: year: 1941; month: 4; day: 27; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.16465; recordNumber: 775; recordedBy: Frey, Richard; individualCount: 1; sex: M; lifeStage: adult; Taxon: scientificName: Coenia vulgata Krivosheina, 2001; order: Diptera; family: Ephydridae; genus: Coenia; specificEpithet: vulgata; scientificNameAuthorship: Krivosheina, 2001; Location: country: Finland; stateProvince: PPe; municipality: Hailuoto; locality: Kirkkolahti; decimalLatitude: 64.98; decimalLongitude: 24.72; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 2000; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 09/26/2013; Event: year: 1947; month: 7; day: 14; habitat: coastal meadows; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.16466; recordNumber: 606; recordedBy: Frey, Richard; individualCount: 1; sex: F; lifeStage: adult; Taxon: scientificName: Coenia vulgata Krivosheina, 2001; order: Diptera; family: Ephydridae; genus: Coenia; specificEpithet: vulgata; scientificNameAuthorship: Krivosheina, 2001; Location: country: Finland; stateProvince: PPe; municipality: Hailuoto; locality: Kirkkolahti; decimalLatitude: 64.98; decimalLongitude: 24.72; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 2000; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 09/26/2013; Event: year: 1947; month: 7; day: 12; habitat: coastal meadows, on water; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.16467; recordNumber: 2557; recordedBy: Frey, Richard; individualCount: 1; sex: F; lifeStage: adult; preparations: gen. prep. on pin; Taxon: scientificName: Coenia vulgata Krivosheina, 2001; order: Diptera; family: Ephydridae; genus: Coenia; specificEpithet: vulgata; scientificNameAuthorship: Krivosheina, 2001; Location: country: Finland; stateProvince: PPe; municipality: Hailuoto; locality: Marjaniemi; decimalLatitude: 65.04; decimalLongitude: 24.56; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 1000; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 09/26/2013; Event: year: 1947; month: 7; day: 13; habitat: a coastal bay and a coastal grazing area; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen

Distribution

First recorded from Finland by Zatwarnicki and Kahanpää (2014). Previously known from Archangelsk Oblast (NW European Russia) and Kazakhstan (Krivosheina 2001).

Notes

Paratype specimen http://id.luomus.fi/GV.4185 is illustrated in Fig. 4 a, c.

Scatophila iowana

Wheeler, 1961

Materials

Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/HT.3120; recordedBy: Storå, Ragnar; individualCount: 1; sex: M; lifeStage: adult; Taxon: scientificName: Scatophila iowana Wheeler, 1961; order: Diptera; family: Ephydridae; genus: Scatophila; specificEpithet: iowana; scientificNameAuthorship: Wheeler, 1961; Location: country: Finland; stateProvince: KP; municipality: Pietarsaari; verbatimLocality: Fennia Om Jakobstad; decimalLatitude: 63.69; decimalLongitude: 22.67; geodeticDatum: WGS84; coordinateUncertaintyInMeters: 10000; georeferencedBy: Jere Kahanpää (MZH); Identification: identificationRemarks: Identified as Scatophila modesta Becker by MG Krivosheina in 2003.; Event: year: 1950; month: 7; day: 5; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen

Diagnosis

Scatophila iowana Wheeler has entirely black legs, a yellow haltere, a matt brown mesonotum and a brownish microtomentose abdomen. It lacks the pair of pronounced facial setae typical for S. despecta (Haliday). The wing and mesonotal patterns of Scatophila iowana resemble those of S. caviceps (Stenhammar). Males of these two species can be identified by the shape of the central part of the face: somewhat protruding between well-defined antennal grooves in S. iowana, weakly concave and without antennal grooves in S. caviceps.

The anepisternum of S. iowana is rather uniformly brownish; S. caviceps has a grey anepisternum with two brown spots, one around the base of the anepisternal bristle and one at the ventral margin.

The male terminalia of S. iowana are characterized by the anterior margin of the epandrium, which is slightly concave, and the aedeagus, which consists of three sclerites. In S. caviceps the anterior margin of the epandrium bears a medial projection and the aedeagus is a one-piece structure.

Distribution

First recorded from Finland by Zatwarnicki and Kahanpää (2014). Scatophila iowana is a Holarctic species. It is probably widespread in Europe, but its distribution remains poorly known (Zatwarnicki 2013).

Scatophila mesogramma

(Loew, 1869)

Materials

Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19175; recordNumber: 6405; recordedBy: Storå, Ragnar; individualCount: 1; lifeStage: adult; Taxon: scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family: Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship: (Loew, 1869); Location: country: Finland; stateProvince: KP; municipality: Pietarsaari; verbatimLocality: Jakobstad; decimalLatitude: 63.704; decimalLongitude: 22.634; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; Event: year: 1954; month: 7; day: 27; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19176; recordNumber: 1743; recordedBy: Storå, Ragnar; individualCount: 1; lifeStage: adult; Taxon: scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family: Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship: (Loew, 1869); Location: country: Finland; stateProvince: KP; municipality: Uusikaarlepyy; verbatimLocality: Nykarleby; decimalLatitude: 63.44; decimalLongitude: 22.675; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; Event: year: 1955; month: 7; day: 20; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19177; recordNumber: 7133; recordedBy: Storå, Ragnar; individualCount: 1; lifeStage: adult; Taxon: scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family: Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship: (Loew, 1869); Location: country: Finland; stateProvince: KP; municipality: Uusikaarlepyy; verbatimLocality: Nykarleby; decimalLatitude: 63.436; decimalLongitude: 22.675; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; Event: year: 1955; month: 7; day: 27; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19178; recordNumber: 885; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family: Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship: (Loew, 1869); Location: country: Russia; stateProvince: Republic of Carelia; municipality: Paanajärvi; locality: behind Takalo farm; verbatimLocality: Paanajärvi; decimalLatitude: 66.26; decimalLongitude: 29.78; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; Event: year: 1939; month: 6; day: 17; habitat: lake shore; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19179; recordNumber: 2005; recordedBy: Storå, Ragnar; individualCount: 1; lifeStage: adult; Taxon: scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family: Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship: (Loew, 1869); Location: country: Finland; stateProvince: KP; municipality: Pietarsaari; verbatimLocality: Jakobstad; decimalLatitude: 63.704; decimalLongitude: 22.634; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; Event: year: 1952; month: 8; day: 24; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19180; recordNumber: 2171; recordedBy: Storå, Ragnar; individualCount: 1; lifeStage: adult; Taxon: scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family: Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship: (Loew, 1869); Location: country: Finland; stateProvince: KP; municipality: Pietarsaari; verbatimLocality: Jakobstad; decimalLatitude: 63.704; decimalLongitude: 22.634; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; Event: year: 1952; month: 8; day: 30; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19181; recordNumber: 4726; recordedBy: Storå, Ragnar; individualCount: 1; lifeStage: adult; Taxon: scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family: Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship: (Loew, 1869); Location: country: Finland; stateProvince: KP; municipality: Uusikaarlepyy; verbatimLocality: Nykarleby; decimalLatitude: 63.436; decimalLongitude: 22.675; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; Event: year: 1954; month: 5; day: 24; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19182; recordNumber: 2215; recordedBy: Storå, Ragnar; individualCount: 1; lifeStage: adult; Taxon: scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family: Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship: (Loew, 1869); Location: country: Finland; stateProvince: KP; municipality: Pietarsaari; verbatimLocality: Jakobstad; decimalLatitude: 63.704; decimalLongitude: 22.634; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; Event: year: 1952; month: 8; day: 30; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19183; recordNumber: 7287; recordedBy: Storå, Ragnar; individualCount: 1; lifeStage: adult; Taxon: scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family: Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship: (Loew, 1869); Location: country: Finland; stateProvince: KP; municipality: Uusikaarlepyy; verbatimLocality: Nykarleby; decimalLatitude: 63.436; decimalLongitude: 22.675; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; Event: year: 1955; month: 8; day: 1; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19184; recordNumber: 4557; recordedBy: Storå, Ragnar; individualCount: 1; lifeStage: adult; Taxon: scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family: Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship: (Loew, 1869); Location: country: Finland; stateProvince: KP; municipality: Pietarsaari; verbatimLocality: Jakobstad; decimalLatitude: 63.704; decimalLongitude: 22.634; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; Event: year: 1954; month: 8; day: 15; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19185; recordNumber: 4559; recordedBy: Storå, Ragnar; individualCount: 1; lifeStage: adult; Taxon: scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family: Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship: (Loew, 1869); Location: country: Finland; stateProvince: KP; municipality: Pietarsaari; verbatimLocality: Jakobstad; decimalLatitude: 63.704; decimalLongitude: 22.634; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; Event: year: 1954; month: 8; day: 15; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19186; recordNumber: 4864; recordedBy: Storå, Ragnar; individualCount: 1; lifeStage: adult; Taxon: scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family: Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship: (Loew, 1869); Location: country: Finland; stateProvince: KP; municipality: Uusikaarlepyy; verbatimLocality: Nykarleby; decimalLatitude: 63.436; decimalLongitude: 22.675; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; Event: year: 1954; month: 5; day: 26; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19187; recordNumber: 1308; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family: Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship: (Loew, 1869); Location: country: Finland; stateProvince: InL; municipality: Utsjoki; verbatimLocality: Utsjoki; decimalLatitude: 69.509; decimalLongitude: 27.109; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19188; recordNumber: 2041; recordedBy: Storå, Ragnar; individualCount: 1; lifeStage: adult; Taxon: scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family: Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship: (Loew, 1869); Location: country: Finland; stateProvince: KP; municipality: Pietarsaari; verbatimLocality: Jakobstad; decimalLatitude: 63.704; decimalLongitude: 22.634; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; Event: year: 1952; month: 8; day: 24; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19189; recordNumber: 4859; recordedBy: Storå, Ragnar; individualCount: 1; lifeStage: adult; Taxon: scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family: Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship: (Loew, 1869); Location: country: Finland; stateProvince: KP; municipality: Uusikaarlepyy; verbatimLocality: Nykarleby; decimalLatitude: 63.436; decimalLongitude: 22.675; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; Event: year: 1954; month: 5; day: 26; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19190; recordNumber: 4858; recordedBy: Storå, Ragnar; individualCount: 1; lifeStage: adult; Taxon: scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family: Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship: (Loew, 1869); Location: country: Finland; stateProvince: KP; municipality: Uusikaarlepyy; verbatimLocality: Nykarleby; decimalLatitude: 63.436; decimalLongitude: 22.675; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; Event: year: 1954; month: 5; day: 26; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19191; recordNumber: 4856; recordedBy: Storå, Ragnar; individualCount: 1; lifeStage: adult; Taxon: scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family: Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship: (Loew, 1869); Location: country: Finland; stateProvince: KP; municipality: Uusikaarlepyy; verbatimLocality: Nykarleby; decimalLatitude: 63.436; decimalLongitude: 22.675; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; Event: year: 1954; month: 5; day: 26; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19192; recordNumber: 408; recordedBy: Hellén, Wolter; individualCount: 1; lifeStage: adult; preparations: gen. prep. on pin; Taxon: scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family: Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship: (Loew, 1869); Location: country: Russia; stateProvince: Murmansk Oblast; verbatimLocality: Kola; decimalLatitude: 68.88; decimalLongitude: 33.04; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; Event: year: 1913; month: 7; day: 29; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19193; recordNumber: 3871; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family: Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship: (Loew, 1869); Location: country: Finland; stateProvince: EnL; municipality: Enontekiö; locality: Palojoensuu; verbatimLocality: Enontekis; decimalLatitude: 68.296; decimalLongitude: 22.983; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; Event: year: 1911; month: 7; day: 11; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19194; recordNumber: 4397; recordedBy: Frey, Richard; individualCount: 1; lifeStage: adult; Taxon: scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family: Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship: (Loew, 1869); Location: country: Finland; stateProvince: EnL; municipality: Enontekiö; verbatimLocality: Enontekis; decimalLatitude: 68.379; decimalLongitude: 22.725; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; Event: year: 1911; month: 7; day: 14; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen
Type status: Other material. Occurrence: catalogNumber: http://id.luomus.fi/GV.19195; recordNumber: 2212; recordedBy: Storå, Ragnar; individualCount: 1; lifeStage: adult; Taxon: scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family: Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship: (Loew, 1869); Location: country: Finland; stateProvince: KP; municipality: Pietarsaari; verbatimLocality: Jakobstad; decimalLatitude: 63.704; decimalLongitude: 22.634; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; Identification: identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; Event: year: 1952; month: 8; day: 20; Record Level: institutionCode: MZH; basisOfRecord: PreservedSpecimen

Distribution

New to Finland and Russia. First recorded from Finland by Zatwarnicki and Kahanpää (2014). Holarctic; in Europe it is known from Finland, Sweden, Denmark, Great Britain, Belgium and Germany (Zatwarnicki 2013).

Discussion

Notes on previous misidentifications

The species identified as Discocerina pulicaria [auct. nec Haliday] by Frey (1933), Frey (1937), Frey (1949), Lindberg and Saris (1952) is actually Ditrichophora fuscella (Stenhammar, 1844). Similarly, Allotrichoma laterale sensu Frey (1940b), Frey (1940a) is A. bezzii Becker, 1896. Previous records of Philygria posticata by Frey 1930 represent Ph. femorata (Stenhammar).

Supplementary Material

XML Treatment for Trimerina microchaeta

BDJ.3.e4701-treatment1.xml^{(21.7KB, xml)}

XML Treatment for Hydrellia albifrons

BDJ.3.e4701-treatment2.xml^{(18.2KB, xml)}

XML Treatment for Hydrellia cochleariae

BDJ.3.e4701-treatment3.xml^{(23.7KB, xml)}

XML Treatment for Hydrellia fulviceps

BDJ.3.e4701-treatment4.xml^{(11.4KB, xml)}

XML Treatment for Hydrellia laticeps

BDJ.3.e4701-treatment5.xml^{(86KB, xml)}

XML Treatment for Hydrellia mutata

BDJ.3.e4701-treatment6.xml^{(36.7KB, xml)}

XML Treatment for Hydrellia subalbiceps

BDJ.3.e4701-treatment7.xml^{(10.5KB, xml)}

XML Treatment for Hydrellia tarsata

BDJ.3.e4701-treatment8.xml^{(11.3KB, xml)}

XML Treatment for Allotrichoma bezzii

BDJ.3.e4701-treatment9.xml^{(149.1KB, xml)}

XML Treatment for Ditrichophora calceata

BDJ.3.e4701-treatment10.xml^{(17.4KB, xml)}

XML Treatment for Hecamedoides unispinosus

BDJ.3.e4701-treatment11.xml^{(162.9KB, xml)}

XML Treatment for Pelina norvegica

BDJ.3.e4701-treatment12.xml^{(16.6KB, xml)}

XML Treatment for Parydra (Chaetoapnea) mitis

BDJ.3.e4701-treatment13.xml^{(53.5KB, xml)}

XML Treatment for Calocoenia paurosoma

BDJ.3.e4701-treatment14.xml^{(52.4KB, xml)}

XML Treatment for Coenia vulgata

BDJ.3.e4701-treatment15.xml^{(28.9KB, xml)}

XML Treatment for Scatophila iowana

BDJ.3.e4701-treatment16.xml^{(11.6KB, xml)}

XML Treatment for Scatophila mesogramma

BDJ.3.e4701-treatment17.xml^{(103.3KB, xml)}

Acknowledgements

We would like to thank Dr. Wayne N. Mathis, Dr. Lorenzo Munari, Dr. Daniel Whitmore and Dr. Owen Lonsdale for their constructive comments on an earlier version of this paper.

References

Becker Theodor. Dipterologische Studien IV. Ephydridae. Berliner Entomologische Zeitschrift. 1896;41:91–276. doi: 10.1002/mmnd.48018960505. [DOI] [Google Scholar]
Canzoneri Silvano, Meneghini Dino. Fauna d'Italia 20: Ephydridae - Canaceidae. Edizioni Calderini; Bologna: 1983. 337. [Google Scholar]
Collin J. E. A contribution towards the knowledge of the male genitalia of species of Hydrellia (Diptera, Ephydridae) Bollettino del Museo Civico di Storia Naturale di Venezia. 1966;16[1963]:7–18. [Google Scholar]
Deonier D. L. A systematic and ecological study of Nearctic Hydrellia (Diptera: Ephydridae) Smithsonian Contributions to Zoology. 1971;68:1–147. doi: 10.5479/si.00810282.68. [DOI] [Google Scholar]
Fischer M. Die paläarktischen Opiinae (Madenwespen) der Zoologischen Staatssammlung München. Entomofauna. 1997;18(14):137–196. [Google Scholar]
Frey R. [Ephydrid-fauna] Notulae entomologicae. 1930;10:114–115. [Google Scholar]
Frey R. En anmärkningsvärd insektstandort vid Viisjoki (Ik) Notulae entomologicae. 1933;13:81–87. [Google Scholar]
Frey R. Einige Massenvorkomnisse von Insekten an der Südküste Finnlands während des Sommers 1935. Ein Beitrag zur Verbreitungsökologie der Insekten. Acta Societatis Pro Fauna et Flora Fennica. 1937;60:406–453. [Google Scholar]
Frey R. De inhemska entomologiska samlingarnas tillväxt under verksamhetsåret 1939-1940. Memoranda Societatis pro Fauna et Flora Fennica. 1940;16:150–152. [Google Scholar]
Frey R. [Månadsmöte- Kuukausikokous] Notulae entomologicae. 1940;20:24. [Google Scholar]
Frey Richard. DipteraBrachycera (excl. Muscidae, Tachinidae) In: Frey Richard, Tiensuu Lauri, Storå Ragnar., editors. Enumeratio Insectorum Fenniae. V I Diptera. Helsingin hyönteisvaihtoyhdistys r.y; Helsinki: 1941. 63 [Google Scholar]
Frey R. Anteckningar om dipterfaunan på Karlo (Hailuoto) sommaren 1947. Memoranda Societatis pro Fauna et Flora Fennica. 1949;24:69–80. [Google Scholar]
Haarto A., Winqvist K. Faunistic notes. Sahlbergia. 2002;7(1):35–37. [Google Scholar]
Hackman W. A check list of the Finnish Diptera II. Cyclorrhapha. Notulae entomologicae. 1980;60:117–162. [Google Scholar]
Heikinheimo Osmo, Raatikainen Mikko. The recording of localities of biological finds in Finland. Annales entomologici fennici. 1971;37(1a):1–27. [Google Scholar]
Heikinheimo Osmo, Raatikainen Mikko. Ruutukoordinaattien ja paikannimien käyttö Suomessa. Notulae entomologicae. 1981;61(3):133–154. [Google Scholar]
Hendel F. Entomologische Ergebnisse der schwedischen Kamtchatka-Expedition 1920-1922. 34. Diptera, Brachycera 3. Arkiv för Zoologi. 1932;23A(7):1–12. [Google Scholar]
Hering E. M. Bestimmunstabellen der Blattminen von Europa: einschliesslich des Mittelmeerbeckens und der Kanarischen Inseln. Band. 1-2. Dr. W. Junk’s-Gravenhage; Haag: 1957. 1185 (vol. 1&2) [Google Scholar]
Hering Martin. Minenstudien VI. Zeitschrift für Morphologie und Ökologie der Tiere. 1926;4(3):502–539. doi: 10.1007/BF00408466. [DOI] [Google Scholar]
Kahanpää J. Viisikymmentä Suomelle uutta kärpäslajia (Diptera: Brachycera) Sahlbergia. 2013;19(1):63–71. [Google Scholar]
Kahanpää Jere, Salmela Jukka. Checklist of the Diptera of Finland. Pensoft Publishers; Sofia: 2014. 408. [Google Scholar]
Kahanpää J., Winqvist K. Rare flies recorded in Finland during the year 2002 (Diptera) Sahlbergia. 2003;8:92–95. [Google Scholar]
Krivosheina M. G. A revision of the shore-fly genus Notiphila Fallén of Palaearctic (Diptera, Ephydridae) An International Journal of Dipterological Research. 1998;9(1):31–63. [Google Scholar]
Krivosheina M. G. New Records of Shore Flies (Diptera: Ephydridae) An International Journal of Dipterological Research. 2000;11(4):171–174. [Google Scholar]
Krivosheina M. G. New Data on Palaearctic Species of the Genus Coenia (Diptera, Ephydridae) Zoologicheskii Zhurnal. 2001;80(11):1367–1371. [Google Scholar]
Krivosheina M. G. New data on shore flies of the genus Trimerina (Diptera, Ephydridae) in the Palaearctic with description of a new species from Finland. Zoologicheskii Zhurnal. 2004;83(5):631–634. [Google Scholar]
Lindberg H., Saris N. -E. Insektfaunan i Pisavaara Naturpark (Finland, Prov. Ob) Acta Societatis pro Fauna et Flora Fennica. 1952;69(2):1–82. [Google Scholar]
Mathis W. N., Zatwarnicki T., Kubátová-Hiršová T. A Revision of the Shore-fly Genus Philotelma Becker (Diptera: Ephydridae) Insect Systematics and Evolution. 2009;40:121–158. doi: 10.1163/187631209X416714. [DOI] [Google Scholar]
Przhiboro A. A. New data on Hydrellia tarsata Haliday (Diptera: Ephydridae) Zoosystematica Rossica. 2004;13(2):239–240. [Google Scholar]
Robbins R. A provisional atlas of the leaf miners of Warwickshire, with notes on others occurring in the Midlands. Warwickshire Museum Service; 1990. 181 [Google Scholar]
Stuke Jens-Hermann. Eine kritische Liste der aus Deutschland nachgewiesenen Ephydridae mit der Beschreibung einer neuen Art (Diptera) Entomologische Zeitschrift. 2011;121(3):115–126. [Google Scholar]
Sturtevant A. H., Wheeler M. R. Synopses of Nearctic Ephydridae (Diptera) Transactions of the American Entomological Society. 1953;79(4):151–257. [Google Scholar]
Winqvist K. Faunistic notes. Species new to the Finnish fauna. Diptera. Sahlbergia. 2002;7:36–37. [Google Scholar]
Winqvist K. Ruissalon kärpäsistä (Diptera, Brachycera) W-album. 2011;11:3–23. [Google Scholar]
Zatwarnicki T. Materials to the knowledge of the genus Hydrellia. Polskie Pismo Entomologiczne. 1988;58:587–634. [Google Scholar]
Zatwarnicki Tadeusz. Fauna Europaea: Ephydridae. Pape Thomas, Beuk Paul., editors. http://www.faunaeur.org Fauna Europaea: Diptera, version 2.6.2. 2013
Zatwarnicki Tadeusz, Kahanpää Jere. Checklist of the family Ephydridae of Finland (Insecta, Diptera) http://dx.doi.org/10.3897/zookeys.441.7448. ZooKeys. 2014;441:339–346. doi: 10.3897/zookeys.441.7448. [DOI] [PMC free article] [PubMed] [Google Scholar]
Zatwarnicki T., Mathis W. N. Heterogeneity in shore flies - the case of Glenanthe Haliday in the Old World (Diptera: Ephydridae) Annales de la Société Entomologique de France. 2012;47(3):418–443. [Google Scholar]

Associated Data

This section collects any data citations, data availability statements, or supplementary materials included in this article.

Supplementary Materials

XML Treatment for Trimerina microchaeta

BDJ.3.e4701-treatment1.xml^{(21.7KB, xml)}

XML Treatment for Hydrellia albifrons

BDJ.3.e4701-treatment2.xml^{(18.2KB, xml)}

XML Treatment for Hydrellia cochleariae

BDJ.3.e4701-treatment3.xml^{(23.7KB, xml)}

XML Treatment for Hydrellia fulviceps

BDJ.3.e4701-treatment4.xml^{(11.4KB, xml)}

XML Treatment for Hydrellia laticeps

BDJ.3.e4701-treatment5.xml^{(86KB, xml)}

XML Treatment for Hydrellia mutata

BDJ.3.e4701-treatment6.xml^{(36.7KB, xml)}

XML Treatment for Hydrellia subalbiceps

BDJ.3.e4701-treatment7.xml^{(10.5KB, xml)}

XML Treatment for Hydrellia tarsata

BDJ.3.e4701-treatment8.xml^{(11.3KB, xml)}

XML Treatment for Allotrichoma bezzii

BDJ.3.e4701-treatment9.xml^{(149.1KB, xml)}

XML Treatment for Ditrichophora calceata

BDJ.3.e4701-treatment10.xml^{(17.4KB, xml)}

XML Treatment for Hecamedoides unispinosus

BDJ.3.e4701-treatment11.xml^{(162.9KB, xml)}

XML Treatment for Pelina norvegica

BDJ.3.e4701-treatment12.xml^{(16.6KB, xml)}

XML Treatment for Parydra (Chaetoapnea) mitis

BDJ.3.e4701-treatment13.xml^{(53.5KB, xml)}

XML Treatment for Calocoenia paurosoma

BDJ.3.e4701-treatment14.xml^{(52.4KB, xml)}

XML Treatment for Coenia vulgata

BDJ.3.e4701-treatment15.xml^{(28.9KB, xml)}

XML Treatment for Scatophila iowana

BDJ.3.e4701-treatment16.xml^{(11.6KB, xml)}

XML Treatment for Scatophila mesogramma

BDJ.3.e4701-treatment17.xml^{(103.3KB, xml)}

[B1229864] Becker Theodor. Dipterologische Studien IV. Ephydridae. Berliner Entomologische Zeitschrift. 1896;41:91–276. doi: 10.1002/mmnd.48018960505. [DOI] [Google Scholar]

[B1226789] Canzoneri Silvano, Meneghini Dino. Fauna d'Italia 20: Ephydridae - Canaceidae. Edizioni Calderini; Bologna: 1983. 337. [Google Scholar]

[B533304] Collin J. E. A contribution towards the knowledge of the male genitalia of species of Hydrellia (Diptera, Ephydridae) Bollettino del Museo Civico di Storia Naturale di Venezia. 1966;16[1963]:7–18. [Google Scholar]

[B1226285] Deonier D. L. A systematic and ecological study of Nearctic Hydrellia (Diptera: Ephydridae) Smithsonian Contributions to Zoology. 1971;68:1–147. doi: 10.5479/si.00810282.68. [DOI] [Google Scholar]

[B1226295] Fischer M. Die paläarktischen Opiinae (Madenwespen) der Zoologischen Staatssammlung München. Entomofauna. 1997;18(14):137–196. [Google Scholar]

[B591615] Frey R. [Ephydrid-fauna] Notulae entomologicae. 1930;10:114–115. [Google Scholar]

[B591625] Frey R. En anmärkningsvärd insektstandort vid Viisjoki (Ik) Notulae entomologicae. 1933;13:81–87. [Google Scholar]

[B591635] Frey R. Einige Massenvorkomnisse von Insekten an der Südküste Finnlands während des Sommers 1935. Ein Beitrag zur Verbreitungsökologie der Insekten. Acta Societatis Pro Fauna et Flora Fennica. 1937;60:406–453. [Google Scholar]

[B591655] Frey R. De inhemska entomologiska samlingarnas tillväxt under verksamhetsåret 1939-1940. Memoranda Societatis pro Fauna et Flora Fennica. 1940;16:150–152. [Google Scholar]

[B591645] Frey R. [Månadsmöte- Kuukausikokous] Notulae entomologicae. 1940;20:24. [Google Scholar]

[B1451096] Frey Richard. DipteraBrachycera (excl. Muscidae, Tachinidae) In: Frey Richard, Tiensuu Lauri, Storå Ragnar., editors. Enumeratio Insectorum Fenniae. V I Diptera. Helsingin hyönteisvaihtoyhdistys r.y; Helsinki: 1941. 63 [Google Scholar]

[B591709] Frey R. Anteckningar om dipterfaunan på Karlo (Hailuoto) sommaren 1947. Memoranda Societatis pro Fauna et Flora Fennica. 1949;24:69–80. [Google Scholar]

[B465969] Haarto A., Winqvist K. Faunistic notes. Sahlbergia. 2002;7(1):35–37. [Google Scholar]

[B465979] Hackman W. A check list of the Finnish Diptera II. Cyclorrhapha. Notulae entomologicae. 1980;60:117–162. [Google Scholar]

[B1430144] Heikinheimo Osmo, Raatikainen Mikko. The recording of localities of biological finds in Finland. Annales entomologici fennici. 1971;37(1a):1–27. [Google Scholar]

[B1430154] Heikinheimo Osmo, Raatikainen Mikko. Ruutukoordinaattien ja paikannimien käyttö Suomessa. Notulae entomologicae. 1981;61(3):133–154. [Google Scholar]

[B1145046] Hendel F. Entomologische Ergebnisse der schwedischen Kamtchatka-Expedition 1920-1922. 34. Diptera, Brachycera 3. Arkiv för Zoologi. 1932;23A(7):1–12. [Google Scholar]

[B1226325] Hering E. M. Bestimmunstabellen der Blattminen von Europa: einschliesslich des Mittelmeerbeckens und der Kanarischen Inseln. Band. 1-2. Dr. W. Junk’s-Gravenhage; Haag: 1957. 1185 (vol. 1&2) [Google Scholar]

[B1226315] Hering Martin. Minenstudien VI. Zeitschrift für Morphologie und Ökologie der Tiere. 1926;4(3):502–539. doi: 10.1007/BF00408466. [DOI] [Google Scholar]

[B466798] Kahanpää J. Viisikymmentä Suomelle uutta kärpäslajia (Diptera: Brachycera) Sahlbergia. 2013;19(1):63–71. [Google Scholar]

[B1145036] Kahanpää Jere, Salmela Jukka. Checklist of the Diptera of Finland. Pensoft Publishers; Sofia: 2014. 408. [Google Scholar]

[B465989] Kahanpää J., Winqvist K. Rare flies recorded in Finland during the year 2002 (Diptera) Sahlbergia. 2003;8:92–95. [Google Scholar]

[B465999] Krivosheina M. G. A revision of the shore-fly genus Notiphila Fallén of Palaearctic (Diptera, Ephydridae) An International Journal of Dipterological Research. 1998;9(1):31–63. [Google Scholar]

[B466009] Krivosheina M. G. New Records of Shore Flies (Diptera: Ephydridae) An International Journal of Dipterological Research. 2000;11(4):171–174. [Google Scholar]

[B465247] Krivosheina M. G. New Data on Palaearctic Species of the Genus Coenia (Diptera, Ephydridae) Zoologicheskii Zhurnal. 2001;80(11):1367–1371. [Google Scholar]

[B466019] Krivosheina M. G. New data on shore flies of the genus Trimerina (Diptera, Ephydridae) in the Palaearctic with description of a new species from Finland. Zoologicheskii Zhurnal. 2004;83(5):631–634. [Google Scholar]

[B591719] Lindberg H., Saris N. -E. Insektfaunan i Pisavaara Naturpark (Finland, Prov. Ob) Acta Societatis pro Fauna et Flora Fennica. 1952;69(2):1–82. [Google Scholar]

[B466049] Mathis W. N., Zatwarnicki T., Kubátová-Hiršová T. A Revision of the Shore-fly Genus Philotelma Becker (Diptera: Ephydridae) Insect Systematics and Evolution. 2009;40:121–158. doi: 10.1163/187631209X416714. [DOI] [Google Scholar]

[B504781] Przhiboro A. A. New data on Hydrellia tarsata Haliday (Diptera: Ephydridae) Zoosystematica Rossica. 2004;13(2):239–240. [Google Scholar]

[B1226334] Robbins R. A provisional atlas of the leaf miners of Warwickshire, with notes on others occurring in the Midlands. Warwickshire Museum Service; 1990. 181 [Google Scholar]

[B589246] Stuke Jens-Hermann. Eine kritische Liste der aus Deutschland nachgewiesenen Ephydridae mit der Beschreibung einer neuen Art (Diptera) Entomologische Zeitschrift. 2011;121(3):115–126. [Google Scholar]

[B533314] Sturtevant A. H., Wheeler M. R. Synopses of Nearctic Ephydridae (Diptera) Transactions of the American Entomological Society. 1953;79(4):151–257. [Google Scholar]

[B466113] Winqvist K. Faunistic notes. Species new to the Finnish fauna. Diptera. Sahlbergia. 2002;7:36–37. [Google Scholar]

[B466029] Winqvist K. Ruissalon kärpäsistä (Diptera, Brachycera) W-album. 2011;11:3–23. [Google Scholar]

[B1226305] Zatwarnicki T. Materials to the knowledge of the genus Hydrellia. Polskie Pismo Entomologiczne. 1988;58:587–634. [Google Scholar]

[B1226271] Zatwarnicki Tadeusz. Fauna Europaea: Ephydridae. Pape Thomas, Beuk Paul., editors. http://www.faunaeur.org Fauna Europaea: Diptera, version 2.6.2. 2013

[B1145026] Zatwarnicki Tadeusz, Kahanpää Jere. Checklist of the family Ephydridae of Finland (Insecta, Diptera) http://dx.doi.org/10.3897/zookeys.441.7448. ZooKeys. 2014;441:339–346. doi: 10.3897/zookeys.441.7448. [DOI] [PMC free article] [PubMed] [Google Scholar]

[B466039] Zatwarnicki T., Mathis W. N. Heterogeneity in shore flies - the case of Glenanthe Haliday in the Old World (Diptera: Ephydridae) Annales de la Société Entomologique de France. 2012;47(3):418–443. [Google Scholar]

PERMALINK

Notes on Shore Flies (Diptera: Ephydridae) from Finland and north-western Russia

Jere Kahanpää

Tadeusz Zatwarnicki

Abstract Abstract

Introduction

Materials and methods

Taxon treatments

Trimerina microchaeta

Materials

Distribution

Taxon discussion

Notes

Figure 1a.

Figure 1b.

Figure 1c.

Figure 1d.

Hydrellia albifrons

Materials

Diagnosis

Distribution

Ecology

Hydrellia cochleariae

Materials

Diagnosis

Distribution

Ecology

Notes

Hydrellia fulviceps

Materials

Diagnosis

Distribution

Ecology

Notes

Hydrellia laticeps

Materials

Diagnosis

Distribution

Hydrellia mutata

Materials

Diagnosis

Distribution

Ecology

Notes

Hydrellia subalbiceps

Materials

Diagnosis

Distribution

Ecology

Hydrellia tarsata

Materials

Diagnosis

Distribution

Biology

Notes

Allotrichoma bezzii

Materials

Distribution

Notes

Ditrichophora calceata

Materials

Distribution

Notes

Hecamedoides unispinosus

Materials

Diagnosis

Distribution

Taxon discussion

Figure 2a.

Figure 2b.

Pelina norvegica

Materials

Distribution

Notes

Parydra (Chaetoapnea) mitis

Materials

Distribution

Notes

Figure 3a.

Figure 3b.