New combinations in Neotropical Thelypteridaceae

Abstract Abstract

288 new combinations of Neotropical Thelypteridaceae taxa are proposed in order to recognize monophyletic genera, based on the results of the most recent molecular phylogeny of the family, as well as the morphological uniformity of characters for each genus. The new nomenclatural combinations correspond to 186 Amauropelta taxa, 77 species of Goniopteris, and 25 Steiropteris taxa. A key to all native Neotropical genera of the family is also presented.

Introduction

The Thelypteridaceae, comprising approximately 1,000 species, is one of the largest families of ferns (Smith et al. 2006). It has a cosmopolitan distribution, with most species occurring in tropical and subtropical regions (Smith 1992, Smith et al. 2006, 2008). The family is monophyletic as usually construed (Smith and Cranfill 2002, Schuettpelz and Pryer 2007, He and Zhang 2013, and Almeida et al. 2015). However, the classification within the family is controversial, with authors recognizing different numbers of genera: 32 (Pichi Sermolli 1977), 25 (Holttum 1971, 1982, Old World only), 20 (Ching 1978), 5 (Smith 1990a, Smith et al. 2006, 2008), or only one (Morton 1969, Tryon and Tryon 1982).

Recent molecular phylogenetic studies have recovered two main groups in the family: the phegopteroid and the thelypteroid lineages (Smith and Cranfill 2002, He and Zhang 2012, and Almeida et al. 2015). The phegopteroid lineage includes two paleotropical genera, Macrothelypteris (H.Itô) Ching (ca. 10 spp.), Pseudophegopteris Ching (ca. 20 spp.), and one north-temperate genus, Phegopteris Fée (4 spp.). In the thelypteroid lineage, Thelypteris s.s. (Thelypteris palustris Schott, Thelypteris confluens (Thunb.) C.V.Morton) is the basal group and sister to a clade that includes all other non-phegopteroid Thelypteridaceae. This confirms the status of Thelypteris s.s. as a small, isolated segregate genus of two species. The sister group to Thelypteris s.s. comprises two well supported, much larger clades: the amauropeltoid clade, comprising the genera Amauropelta Kunze, Parathelypteris (H. Itô) Ching, Coryphopteris Holttum, and Metathelypteris (H.Itô) Ching, and the cyclosoroid clade, comprising several genera from the Old World and the New World (He and Zhang 2012, and Almeida et al. 2015). Inside the cylosoroid clade, several groups in the Paleotropics are still ill-defined and probably not monophyletic (He and Zhang 2012, Almeida et al. 2015). Thus, the current infrafamilial classification for Thelypteridaceae is not in accordance with growing phylogenetic evidence and principles of monophyly.

The following native Neotropical genera of Thelypteridaceae are recognized: Amauropelta, Christella Lév. (a more precise circumscription still pending), Cyclosorus Link, Goniopteris (C.Presl) Duek., Meniscium Schreber, Stegnogramma Blume, Steiropteris C.Chr., and Thelypteris Schmidel, based on the recent molecular phylogenetic data for this family (Almeida et al. 2015) and the morphological uniformity recognized in previously proposed classifications (Holttum 1971, Pichi Sermolli 1977, Smith 1980, 1990a). With the exception of Amauropelta and Thelypteris, the other six genera belong to the cyclosoroid clade. In the most recent published floras for Neotropical areas, these monophyletic groups of Thelypteridaceae were treated as subgenera of Thelypteris. In the most recent fern classifications (Smith 1990a; Smith et al. 2006, 2008; Christenhusz et al. 2011), Amauropelta was treated within the genus Thelypteris, and Christella, Cyclosorus, Goniopteris, Meniscium, and Steiropteris within the genus Cyclosorus.

Utilizing our most recent results on molecular phylogeny of the family (Almeida et al. 2015), the monophyletic genera accepted here have strong support. They are also easily defined morphologically by characters in the key below.

Amauropelta, predominantly Neotropical, is most closely related to several Old World genera: Coryphopteris, Metathelypteris, and Parathelypteris. Amauropelta is recognized mainly by the laminar base usually gradually or abruptly reduced with one to many pairs of auriculiform or glanduliform pinnae, and by the veins from adjacent segments usually meeting the blade margins above the sinuses on pinnatifid pinnae.

Our results (Almeida et al. 2015) confirm that Steiropteris is monophyletic and sister to all remaining cyclosoroid genera except Stegnogramma. These results include an interesting novelty: Thelypteris polypodioides (Raddi) C.F.Reed and Thelypteris villosa (Link) C.F.Reed, from southern Brazil (previously incertae sedis), nest in Steiropteris with strong support. These two species were treated by Christensen (1913) as belonging to Dryopteris subg. Leptogramma (J.Sm.) C.Chr. (= Stegnogramma).

Goniopteris has long been recognized as a natural group (Christensen 1913, Tryon and Tryon 1982, Smith 1990a). It is also monophyletic (Smith and Cranfill 2002, Schuettpelz and Pryer 2007) and has been accorded generic status, e.g., by Copeland (1947), Brade (1972) and Pichi Sermolli (1977). The presence of furcate or stellate hairs on the blades and/or rhizome apex scales is a useful diagnostic character for Goniopteris (Salino 2002), but this character has apparently been secondarily lost in a few species: Goniopteris mollis Fée (Mexico and Central America; Mickel and Smith 2004; Smith 1995a), Goniopteris macrotis (Hook.) Pic.Serm., Goniopteris semihastata (Kunze) Salino & T.E.Almeida (both endemic to Peru; Smith 1992), Goniopteris holodictya (K.U.Kramer) Salino & T.E.Almeida (French Guiana; Smith 1993), and Goniopteris clypeata (Maxon & C.V.Morton) Salino & T.E.Almeida (Panama, Peru; Smith 1995a, and Colombia).

The narrow circumscription of Cyclosorus used here has already been adopted by some authors (e.g., Pichi Sermolli 1977, Holttum 1971, 1982). This clade (pantropical, with two species, only one of which, Cyclosorus interruptus (Willd.) H.Itô, occurs in the Neotropics) is supported as monophyletic in the results of our molecular phylogeny (Almeida et al. 2015). Most native species previously considered in Thelypteris subg. Cyclosorus (Link) C.V.Morton in some floras (Mickel and Beitel 1988, Mickel and Smith 2004, Proctor 1977, 1985, 1989, Salino and Semir 2002, Smith 1981a,b, 1983, 1988, 1992, 1995a,b) are now treated in the genus Christella, as done by Pichi Sermolli (1977) and Holttum (1974, 1982). However, it appears likely that Christella sensu Holttum is not monophyletic (see Smith and Cranfill 2002; Schuettpelz and Pryer 2007; He and Zhang 2012).

Following Smith (1990a) and Smith et al. (2006, 2008), Mazumdar and Mukhopadhyay (2013) proposed new combinations in Cyclosorus for many species nested in the cyclosoroid clade. However, if this clade is to be considered as a single genus, the oldest generic name is Meniscium (Schreber 1791). In order to retain all the names recombined by Mazumdar and Mukhopadhyay (2013), Mazumdar (2015) subsequently proposed conservation of Cyclosorus against Meniscium. However, we believe conservation is unnecessary, because the cyclosoroid clade, considered by Mazumdar (2015) as a genus, corresponds to at least 10 genera according to recent molecular phylogenetic results (Almeida et al. 2015). We think it preferable that Cyclosorus sensu stricto be construed as comprising only two species (see above). At the same time, we interpret phylogenetic evidence to best support Meniscium as a genus comprising 26 species (Fernandes et al. 2014).

Meniscium, wholly Neotropical, appears to be monophyletic and more closely related to Cyclosorus s.s. (Cyclosorus interruptus), Mesophlebium Holttum, and Ampelopteris Kunze (Almeida et al. 2015). Meniscium is morphologically uniform and characterized by short-creeping rhizomes, pinnate leaves each with a conform apical pinna, and by regularly anastomosing veins resulting in parallel rows of areoles that include either a single free vein or a bisecting vein. Fernandes et al. (2014) made five new combinations for Meniscium species; combinations are already available for other recognized menisciums. Similarly, combinations are available for 38 species of Amauropelta, for nearly 40 species of Goniopteris, as well as for nine species of Steiropteris. New combinations in Amauropelta, Goniopteris, and Steiropteris are established for most remaining Neotropical species, those that appear commonly recognized in existing floras and monographs. We refrain from addressing further Christella sensu Holttum in the Neotropics, because (as mentioned above) existing evidence suggests that it is not monophyletic – most of the neotropical species (sect. Pelazoneuron Holttum) appear to comprise a clade separate from the paleotropical species (sect. Christella). However, sampling of Christella in both the Old World and New World, as well as affiliated cyclosoroid genera circumscribed by Holttum (1982) in the Paleotropics, is still far too meager to construct a viable taxonomy.

Key to the native Neotropical Thelypteridaceae genera

1	Forked or stellate hairs present at least on the rachises and costae, and/or on the scales at apices of rhizomes, and may be present in other parts of the frond	Goniopteris
–	All hairs unbranched (hairs fasciculate, i.e., basally clustered, in some species of Amauropelta)
2	Laminae 1-pinnate, rarely simple; pinnae entire or with an serrate or undulate margin, rarely shallowly lobed; veins regularly anatomosing, areoles 3–25-seriate between costae and pinna margins	3
–	Laminae 1-pinnate-pinnatifid, rarely 2-pinnate; pinnae incised 1/3 their width or more; veins meeting the margins above sinuses, or connivent at sinuses, or with 1 or 2 pairs of anastomosing below sinuses	4
3	Sori usually in a single row between adjacent costules (except Meniscium andreanum, which often has a double row of sori between main lateral veins), oblong or linear, rarely round; sori exindusiate	Meniscium
–	Sori usually in a double row between adjacent costules, usually round (except Goniopteris holodictya, which sometimes has only one row of somewhat elongate sori between costules); sori indusiate (indusial may be small, or hidden in mature sori!) or exindusiate	Goniopteris
4	Sporangia setose; sori elongate, exindusiate	5
–	Sporangia not setose, or if setose (in eight species of Amauropelta) then sori round; sori round, oblong, or rarely elongate; sori indusiate or exindusiate	6
5	Pinnae broadly adnate and decurrent onto rachises in the distal half of laminae; proximal pinnae not cuneate at bases	Stegnogramma (Stegnogramma pilosa, Stegnogramma burksiorum)
–	Pinnae neither adnate nor decurrent onto rachises, sometimes one distal pair adnate to rachises; proximal pinnae short- to long-cuneate at bases	Steiropteris (Stegnogramma polypodioides, Stegnogramma villosa)
6	Laminae with all veins usually meeting margins above the sinuses	7
–	Laminae with at least some of the proximal veins running to the sinuses or forming an excurrent vein to the sinuses	9
7	Laminae narrowed proximally with (1-) 2−20 pairs of greatly reduced pinnae, the proximal pinnae usually auriculiform or glanduliform, rarely proximal pinnae not reduced	Amauropelta
–	Laminae usually not narrowed proximally, or only slightly so, usually lacking greatly reduced pinnae	8
8	Rhizomes long-creeping; laminae 1-pinnate-pinnatifid to 2-pinnate; veins of sterile laminae 1- or 2-forked, sometimes simple in the fertile laminae	Thelypteris
–	Stems ascending to erect, rarely short-creeping; laminae 1-pinnate-pinnatifid; veins of sterile and fertile laminae simple.. Steiropteris (subg. Glaphyropteris)
9	Rhizomes long-creeping to 3 m or more, blackish, scaleless or nearly so	Cyclosorus
–	Rhizomes short-creeping to ascending or erect, light brown, with conspicuous scales, at least at apices	10
10	Cartilaginous keel (false vein) present below pinna sinuses, keels extending toward costa but not meeting it; aerophores (tuberculate or often threadlike) at pinna bases usually present, absent in a few species; indusia present or absent	Steiropteris (subg. Steiropteris)
–	Cartilaginous keel absent below pinna sinuses; aerophores at pinna bases absent; indusia present, persistent	Christella

New combinations

The names proposed as needing new combinations were obtained from taxonomic and floristic sources such as floras, monographs, or isolated published papers. All names were checked in IPNI.ORG and TROPICOS.ORG. The main floras consulted were ones for Mexico (Smith 1981a, 1988; Mickel and Beitel 1988, Mickel and Smith 2004), Lesser Antilles (Proctor 1977), Jamaica (Proctor 1985), Puerto Rico and the Virgin Islands (Proctor 1989), Cuba (Sánchez et al 2006), Mesoamerica (Smith 1995a), Guatemala (Smith 1981b), Venezuela (Vareschi 1969; Smith 1995b), Guianas (Smith 1993), Ecuador (Smith 1983), Peru (Smith 1992), Argentina (Ponce 1987), and from Brazil (Brade 1971, Sehnem 1979, Salino and Semir 2002, 2003, 2004, Ponce 2007, Salino and Almeida 2010). The monographs of reference were the works of Christensen (1907, 1913, 1920) and Smith (1980). In addition, the following works describing new species were consulted: Smith and Lellinger (1985), Smith (1990b), Salino (2002), Salino and Melo (2000), Smith and Kessler (2008), Matos et al. (2010), and Salino et al. (2011, 2014). The authorship of new combinations was established as follows: species in Venezuela are credited to Alan R. Smith; Alexandre Salino and Thais Elias Almeida are credited with authorship of the remaining species. We choose to cite only the basionym and the name in Thelypteris (when available) for all new combinations; this avoids citation of the often extensive synonymy, information that is available in many recent floristic and taxonomic studies.

New combinations in Amauropelta

Amauropelta achalensis (Hieron.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151259-1

• Aspidium achalense Hieron., Bot. Jahrb. Syst. 22: 371. 1896.

• Thelypteris achalensis (Hieron.) Abbiatti, Darwiniana 13: 566. 1964.

Amauropelta aculeata (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150878-1

• Thelypteris aculeata A.R.Sm., Fl. Ecuador 18: 15. F. 1A, 1B. 1983.

Amauropelta acunae (C.Sánchez & Zavaro)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150879-1

• Thelypteris acunae C.Sánchez & Zavaro, Fontqueria 31: 223. f.1. 1991.

Amauropelta aliena (C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150880-1

• Dryopteris aliena C.Chr., Kongl. Svenska Vetenskapsakad. Handl., ser. 3, 16: 23. 1937.

Amauropelta altitudinis (Ponce)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151260-1

• Thelypteris altitudinis Ponce, Darwiniana 28 (1-4): 345. 1987.

Amauropelta amambayensis (Ponce)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151261-1

• Thelypteris amambayensis Ponce, Candollea 55: 310.

Amauropelta amphioxypteris (Sodiro)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150881-1

• Nephrodium amphioxypteris Sodiro, Recens. Crypt. Vasc. Quit. 51. 1883.

• Thelypteris amphioxypteris (Sodiro) A.R.Sm., Fl. Ecuador 18: 17. 1983.

Amauropelta andicola (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150882-1

• Thelypteris andicola A.R.Sm., Fieldiana, Bot., n.s., 29: 16. 1992.

Amauropelta appressa (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150883-1

• Thelypteris appressa A.R.Sm., Fl. Ecuador 18: 19. 1983.

Amauropelta araucariensis (Ponce)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150884-1

• Thelypteris araucariensis Ponce, Darwiniana 33: 270. 1995.

Amauropelta arborea (Brause)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77151262-1

• Dryopteris arborea Brause, Repert. Spec. Nov. Regni Veg. 13: 294. 1914.

• Thelypteris arborea (Brause) A.R.Sm., Acta Bot. Venez. 14 (3): 7. 1984.

Amauropelta arenosa (A.R.Sm.)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77150885-1

• Thelypteris arenosa A.R.Sm., Fl. Ecuador 18: 21. 1983.

Amauropelta argentina (Hieron.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151263-1

• Aspidium argentinum Hieron., Bot. Jahrb. Syst. 22: 367. 1896.

• Thelypteris argentina (Hieron.) Abbiatti, Revista Mus. La Plata, Secc. Bot. 9 (36-37): 19. 1958.

Amauropelta arrecta (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150886-1

• Thelypteris arrecta A.R.Sm., Fieldiana, Bot., n.s., 29: 36. 1992.

Amauropelta atrorubens (Mett. ex Kuhn)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151264-1

• Aspidium atrorubens Mett. ex Kuhn, Linnaea 36. 112. 1869.

• Thelypteris atrorubens (Mett. ex Kuhn) A.R.Sm., Fieldiana, Bot., n.s. 29: 24. 1992.

Amauropelta atrovirens (C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150887-1

• Dryopteris atrovirens C.Chr., Bull. Herb. Boissier, sér. 2, 7 (4): 263. 1907.

• Thelypteris atrovirens (C.Chr.) C.F.Reed, Phytologia 17 (4): 261. 1968.

Amauropelta aymarae (A.R.Sm. & M.Kessler)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150888-1

• Thelypteris aymarae A.R.Sm. & M.Kessler, Brittonia 60 (1): 49. 2008.

Amauropelta balbisii (Spreng.)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77150889-1

• Polypodium balbisii Spreng., Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 10: 228. 1821.

• Thelypteris balbisii (Spreng.) Ching, Bull. Fan Mem. Inst. Biol. 10: 250. 1941.

Amauropelta barvae (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150890-1

• Thelypteris barvae A.R.Sm., Ann. Missouri Bot. Gard. 77 (1): 118. f. 2A-C. 1990.

Amauropelta basisceletica (C.Sánchez, Caluff & O.Alvarez)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151265-1

• Thelypteris basisceletica C.Sánchez, Caluff & O.Alvarez, Amer. Fern J. 95 (1): 30. f. 1. 2005.

Amauropelta binervata (A.R.Sm.)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77150891-1

• Thelypteris binervata A.R.Sm., Acta Bot. Venez. 14 (3): 6. 1984.

Amauropelta bonapartii (Rosenst.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151266-1

• Dryopteris bonapartii Rosenst., Repert. Spec. Nov. Regni Veg. 7: 303–304. 1909.

• Thelypteris bonapartii (Rosenst.) Alston, J. Wash. Acad. Sci. 48: 233.1958.

Amauropelta brachypoda (Baker)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77150892-1

• Nephrodium brachypodum Baker, Timehri 5: 213. 1886.

• Thelypteris brachypoda (Baker) C.V.Morton, Fieldiana, Bot. 28: 10. 1951.

Amauropelta brachypus (Sodiro)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150893-1

• Nephrodium brachypus Sodiro, Recens. Crypt. Vasc. Quit. 43. 1883.

• Thelypteris brachypus (Sodiro) R.M.Tryon & A.F.Tryon, Rhodora 84: 128. 1982.

Amauropelta brausei (Hieron.)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77150894-1

• Dryopteris brausei Hieron., Hedwigia 46: 337, t. 6, f. 11. 1907.

• Thelypteris brausei (Hieron.) Alston, J. Wash. Acad. Sci. 48: 233. 1958.

Amauropelta burkartii (Abbiatti)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150895-1

• Thelypteris burkartii Abbiatti, Darwiniana 13: 550, f. 4, t. 2. 1964.

Amauropelta campii (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151267-1

• Thelypteris campii A.R.Sm., Fl. Ecuador 18: 27. 1983.

Amauropelta canadasii (Sodiro)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150896-1

• Nephrodium canadasii Sodiro, Recens. Crypt. Vasc. Quit. 48. 1883.

• Thelypteris canadasii (Sodiro) Alston, J. Wash. Acad. Sci 48: 234. 1958.

Amauropelta caucaensis (Hieron.)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77151268-1

• Nephrodium caucaense Hieron., Bot. Jahrb. Syst. 34: 444. 1904.

• Thelypetris caucaensis (Hieron.) Alston, J. Wash. Acad. Sci 48: 233. 1958.

Amauropelta chaparensis (A.R.Sm. & M.Kessler)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150897-1

• Thelypteris chaparensis A.R.Sm. & M.Kessler, Brittonia 60 (1): 51. 2008.

Amauropelta chiriquiana (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150898-1

• Thelypteris chiriquiana A.R.Sm., Ann. Missouri Bot. Gard. 77 (1): 120. f. 3A-C. 1990.

Amauropelta christensenii (Christ ex C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151269-1

• Dryopteris christensenii Christ ex C.Chr., Bull. Herb. Boissier, sér. 2, 7 (4): 263. 1907.

• Thelypteris christensenii (Christ ex C.Chr.) C.F.Reed, Phytologia 17 (4): 267. 1968.

Amauropelta cinerea (Sodiro)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77150900-1

• Nephrodium cinereum Sodiro, Sert. Fl. Ecuad. II. 26. 1908.

• Thelypteris cinerea (Sodiro) A.R.Sm., Phytologia 34: 233. 1976.

Amauropelta cochaensis (C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150901-1

• Dryopteris cochaensis C.Chr., Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10: 152. 1913.

Amauropelta cocos (A.R.Sm. & Lellinger)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150904-1

• Thelypteris cocos A.R.Sm. & Lellinger, Proc. Biol. Soc. Wash. 98 (4): 918. f. 3. 1985.

Amauropelta comptula (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150906-1

• Thelypteris comptula A.R.Sm., Fieldiana, Bot., n.s., 29: 23. 1992.

Amauropelta conformis (Sodiro)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150908-1

• Nephrodium conforme Sodiro, Recens. Crypt. Vasc. Quit. 45. 1883.

• Thelypteris conformis (Sodiro) A.R.Sm., Fl. Ecuador 18: 37. 1983.

Amauropelta consanguinea (Fée)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150910-1

• Aspidium consanguineum Fée, Mém. Foug. 11 (Hist. Foug. Ant.): 76, t. 20, f. 3). 1866.

• Thelypteris consanguinea (Fée) Proctor, Rhodora 61 (732): 306. 1959.

Amauropelta cooleyi (Proctor)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150912-1

• Thelypteris cooleyi Proctor, Rhodora 68: 468. 1966.

Amauropelta corazonensis (Baker)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151270-1

• Nephrodium corazonense Baker, J. Bot. 15: 163. 1877, as carazanensis”.

• Thelypteris corazonensis (Baker) A.R.Sm., Fl. Ecuador 18: 38. 1983.

Amauropelta cornuta (Maxon)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150914-1

• Dryopteris cornuta Maxon, J. Wash. Acad. Sci. 19 (12): 245. f. 1. 1929.

• Thelypteris cornuta (Maxon) Ching, Bull. Fan Mem. Inst. Biol. 10: 251. 1941.

Amauropelta correllii (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150916-1

• Thelypteris correllii A.R.Sm., Fl. Ecuador 18: 39. 1983.

Amauropelta crassiuscula (C.Chr. & Maxon)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150917-1

• Dryopteris crassiuscula C.Chr. & Maxon, Amer. Fern J. 23: 75. 1933.

• Thelypteris crassiuscula (C.Chr. & Maxon) Lellinger, Amer. Fern J. 67(2): 60. 1977.

Amauropelta ctenitoides (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150920-1

• Thelypteris ctenitoides A.R.Sm., Fieldiana, Bot., n.s., 29: 37. 1992.

Amauropelta decrescens (Proctor)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151271-1

• Thelypteris decrescens Proctor, Amer. Fern J. 71(2): 57. 1981.

Amauropelta decurtata (Link)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151422-1

• Asplenium decurtatum Link Fil. sp. 94. 1841.

Amauropelta deflectens (C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150922-1

• Dryopteris deflectens C.Chr., Kongl. Svenska Vetenskapsakad. Handl., ser. 3, 16(2): 25, t. 3, f. 9-11. 1937.

Amauropelta delasotae (A.R.Sm. & Lellinger)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150924-1

• Thelypteris delasotae A.R.Sm. & Lellinger, Proc. Biol. Soc. Wash. 98(4): 919. f. 5. 1985.

Amauropelta demissa (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150926-1

• Thelypteris demissa A.R.Sm., Fieldiana, Bot., n.s., 29: 21. 1992.

Amauropelta denudata (C.Sánchez & Caluff)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150928-1

• Thelypteris denudata C.Sánchez & Caluff, Willdenowia 35: 159. 2005.

Amauropelta dodsonii (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150930-1

• Thelypteris dodsonii A.R.Sm., Fl. Ecuador 18: 41. 1983.

Amauropelta dudleyi (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150932-1

• Thelypteris dudleyi A.R.Sm., Fieldiana, Bot., n.s., 29: 33. 1992.

Amauropelta elegantula (Sodiro)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150933-1

• Nephrodium elegantulum Sodiro, Crypt. Vasc. Quit. 243. 1893.

• Thelypteris elegantula (Sodiro) Alston, J. Wash. Acad. Sci. 48: 233. 1958.

Amauropelta enigmatica (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150936-1

• Thelypteris enigmatica A.R.Sm., Fieldiana, Bot., n.s., 29: 16. 1992.

Amauropelta eriosorus (Fée)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151278-1

• Aspidium eriosorus Fée, Crypt. Vasc. Brésil, 2. 73. T. 101. 1873.

• Thelypteris eriosorus (Fée) Ponce (as T. eriosora), Novon 8: 275. 1998.

Amauropelta euchlora (Sodiro)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77150938-1

• Polypodium euchlorum Sodiro, Recens. Crypt. Vasc. Quit. 58. 1883.

• Thelypteris euchlora (Sodiro) C.F.Reed, Phytologia 17(4): 275. 1968.

Amauropelta euthythrix (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150940-1

• Thelypteris euthythrix A.R.Sm., Fl. Ecuador 18: 44. 1983.

Amauropelta exuta (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151279-1

• Thelypteris exuta A.R.Sm., Fl. Ecuador 18: 44. 1983.

Amauropelta fasciola (A.R.Sm. & M.Kessler)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150942-1

• Thelypteris fasciola A.R.Sm. & M.Kessler, Brittonia 60 (1): 51. 2008.

Amauropelta fayorum (A.R.Sm. & M.Kessler)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150944-1

• Thelypteris fayorum A.R.Sm. & M.Kessler, Brittonia 60(1): 53. 2008.

Amauropelta firma (Baker ex Jenman)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150946-1

• Nephrodium firmum Baker ex Jenman., J. Bot. 17: 260. 1879.

• Thelypteris firma (Baker ex Jenman) Proctor, Bull. Inst. Jamaica, Sci. Ser. 5: 60. 1953.

Amauropelta fluminalis (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150948-1

• Thelypteris fluminalis A.R.Sm., Fl. Ecuador 18: 46. 1983.

Amauropelta frigida (Christ)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77150949-1

• Aspidium frigidum Christ, Bull. Herb. Boissier, sér. 2., 6: 160. 1906.

• Thelypteris frigida (Christ) A.R.Sm. & Lellinger, Amer. Fern J. 75(1): 31. 1985.

Amauropelta funckii (Mett.)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77150952-1

• Aspidium funckii Mett., Ann. Sci. Nat., Bot., sér. 5, 2: 246. 1864.

• Thelypteris funckii (Mett.) Alston, J. Wash. Acad. Sci. 48: 233. 1958.

Amauropelta furfuracea (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150954-1

• Thelypteris furfuracea A.R.Sm., Fieldiana, Bot., n.s., 29: 34. 1992.

Amauropelta furva (Maxon)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150956-1

• Dryopteris furva Maxon, J. Wash. Acad. Sci. 34: 24. 1944.

• Thelypteris furva (Maxon) R.M.Tryon, Rhodora 69: 6. 1977.

Amauropelta germaniana (Fée)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150958-1

• Phegopteris germaniana Fée, Mém. Foug. 11 (Hist. Foug. Ant.): 55. 1866.

• Thelypteris germaniana (Fée) Proctor, Rhodora 61: 306. 1959.

Amauropelta glandulosolanosa (C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150960-1

• Dryopteris glandulosolanosa C.Chr., Dansk Bot. Ark. 9(3): 61. 1937.

• Thelypteris glandulosolanosa (C.Chr.) R.M.Tryon, Rhodora 69: 6. 1967.

Amauropelta glutinosa (C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150962-1

• Dryopteris glutinosa C.Chr., Kongl. Svenska Vetenskapsakad. Handl., ser. 3, 16(2): 18. 1937.

• Thelypteris glutinosa (C.Chr.) C.V.Morton, Amer. Fern J. 53(2): 66. 1963.

Amauropelta gomeziana (A.R.Sm. & Lellinger)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150964-1

• Thelypteris gomeziana A.R.Sm. & Lellinger, Proc. Biol. Soc. Wash. 98(4): 921. f 6. 1985.

Amauropelta gracilenta (Jenman)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150965-1

• Polypodium gracilentum Jenman, Bull. Bot. Dept. Jamaica 4. 129. 1897.

• Thelypteris gracilenta (Jenman) Proctor, Amer. Fern J. 71(2): 60. 1981.

Amauropelta gracilis (Heward)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77150968-1

• Gymnogramma gracilis Heward, Mag. Nat. Hist., n.s, 2: 457. 1838.

• Thelypteris gracilis (Heward) Proctor, Bull. Inst. Jamaica, Sci. Ser. 5: 60. 1953.

Amauropelta grayumii (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150970-1

• Thelypteris grayumii A.R.Sm., Ann. Missouri Bot. Gard. 77(1): 122. f 2D-E. 1990.

Amauropelta harrisii (Proctor)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151280-1

• Thelypteris harrisii Proctor, Amer. Fern J. 71: 59. 1981.

Amauropelta hastiloba (C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150972-1

• Dryopteris hastiloba C.Chr., Kongl. Svenska Vetenskapsakad. Handl., ser. 3, 16(2): 20. t 4, f 4-5. 1937.

Amauropelta heineri (C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150974-1

• Dryopteris heineri C.Chr., Repert. Spec. Nov. Regni Veg. 6: 380. 1909.

• Thelypteris heineri (C.Chr.) C.F.Reed, Phytologia 17(4): 282. 1968.

Amauropelta hutchisonii (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150976-1

• Thelypteris hutchisonii A.R.Sm., Fieldiana, Bot., n.s., 29: 29. 1992.

Amauropelta hydrophila (Fée)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150978-1

• Phegopteris hydrophila Fée, Mém. Foug. 11 (Hist. Foug. Ant.): 56. pl 13, f 3. 1866.

• Thelypteris hydrophila (Fée) Proctor, Rhodora 61: 306. 1959.

Amauropelta illicita (Christ)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150980-1

• Dryopteris illicita Christ, Bull. Soc. Bot. Genève sér. 2, 1(5): 225. 1909.

• Thelypteris illicita (Christ) C.F.Reed, Phytologia 17(4) : 284. 1968.

Amauropelta inabonensis (Proctor)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151281-1

• Thelypteris inabonensis Proctor, Amer. Fern J. 75(2): 61. f 3. 1985.

Amauropelta inaequans (C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150981-1

• Dryopteris euchlora (Sodiro) C.Chr. var. inaequans C.Chr., Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10: 150. 1913.

• Thelypteris inaequans (C.Chr.) Lellinger, Amer. Fern J. 67(2): 60. 1977.

Amauropelta inaequilateralis (A.R.Sm. & M.Kessler)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150984-1

• Thelypteris inaequilateralis A.R.Sm. & M.Kessler, Brittonia 60(1): 53. 2008.

Amauropelta insignis (Mett.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151282-1

• Aspidium insigne Mett., Ann. Sci. Nat., Bot., sér. 5, 2: 247. 1864.

• Thelypteris insignis (Mett.) Ching, Bull. Fan Mem. Inst. Biol. 10: 242. 1941.

Amauropelta intromissa (C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150986-1

• Dryopteris intromissa C.Chr., Kungl. Svenska Vetenskapsakad. Handl., ser. 3., 16(2): 22. t 4, f 9-10. 1937.

Amauropelta ireneae (Brade)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150988-1

• Dryopteris ireneae Brade, Sellowia 17: 57. f 4. 1965.

• Thelypteris ireneae (Brade) Lellinger, Amer. Fern. J. 74(2): 60. 1984.

Amauropelta jimenezii (Maxon & C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151283-1

• Dryopteris jimenezii Maxon & C.Chr., Amer. Fern J. 4: 79. 1914.

• Thelypteris jimenezii (Maxon & C.Chr.) C.F.Reed, Phytologia 17(4): 285. 1968.

Amauropelta juergensii (Rosenst.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150990-1

• Nephrodium juergensii Rosenst., Festschrift Albert von Bamberg 63. 1905.

• Thelypteris juergensii (Rosenst.) C.F.Reed, Phytologia 17(4): 286. 1968.

Amauropelta jujuyensis (de la Sota)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150992-1

• Thelypteris jujuyensis de la Sota, Darwiniana 18: 222. f 3, 8c. 1973.

Amauropelta laevigata (Mett. ex Kuhn)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150994-1

• Phegopteris laevigata Mett. ex Kuhn, Linnaea 36. 112. 1869.

• Thelypteris laevigata (Mett. ex Kuhn) R.M.Tryon, Rhodora 69: 6. 1967.

Amauropelta leoniae (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77150996-1

• Thelypteris leoniae A.R.Sm., Fieldiana, Bot., n.s., 29: 18. 1992.

Amauropelta lepidula (Hieron.)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77150997-1

• Dryopteris lepidula Hieron., Hedwigia 46. 328, t. 4, f. 4. 1907.

• Thelypteris lepidula (Hieron.) Alston, J. Wash. Acad. Sci. 48: 233. 1958.

Amauropelta longicaulis (Baker)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151000-1

• Nephrodium longicaule Baker, J. Bot. 19: 204. 1881.

• Thelypteris longicaulis (Baker) C.F.Reed, Phytologia 17(4): 289. 1968.

Amauropelta longipilosa (Sodiro)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151284-1

• Nephrodium longipilosum Sodiro, Anales Univ. Central Quito Ecuador 22(160): 103. 1908.

• Thelypteris longipilosa (Sodiro) C.F.Reed, Phytologia 17(4): 290. 1968.

Amauropelta longisora (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151002-1

• Thelypteris longisora A.R.Sm., Ann. Missouri Bot. Gard. 77(1): 123. f 3D-E. 1990.

Amauropelta loreae (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151003-1

• Thelypteris loreae A.R.Sm., Mem. New York Bot. Gard. 88: 629–630. f 303A-E, 2004.

Amauropelta loretensis (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151006-1

• Thelypteris loretensis A.R.Sm., Fieldiana, Bot., n.s., 29: 30. 1992.

Amauropelta lumbricoides (A.R.Sm. & M.Kessler)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151008-1

• Thelypteris lumbricoides A.R.Sm. & M.Kessler, Brittonia 60(1): 54. 2008.

Amauropelta macra (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151010-1

• Thelypteris macra A.R.Sm., Fl. Ecuador 18: 54. 1983.

Amauropelta madidiensis (A.R.Sm. & M.Kessler)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151012-1

• Thelypteris madidiensis A.R.Sm. & M.Kessler, Brittonia 60(1): 54. 2008.

Amauropelta malangae (C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151014-1

• Dryopteris malangae C.Chr., Kongl. Svenska Vetenskapsakad. Handl., ser. 3, 16(2): 21. t. 4(6−8). 1937.

• Thelypteris malangae (C.Chr.) C.V.Morton, Amer. Fern J. 53(2): 66. 1963.

Amauropelta melanochlaena (C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151285-1

• Dryopteris melanochlaena C.Chr., Smithsonian Misc. Collect. 52(3): 384. 1909.

• Thelypteris melanochlaena (C.Chr.) C.F.Reed, Phytologia 17(4): 292. 1968.

Amauropelta mertensioides (C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151016-1

• Dryopteris mertensioides C.Chr., Kongel. Danske.Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 4: 328, f. 50. 1907.

• Thelypteris mertensioides (C.Chr.) C.F.Reed, Phytologia 17(4): 292. 1968.

Amauropelta metteniana (Ching)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151017-1

• Thelypteris metteniana Ching, Bull. Fan mem. Inst. Biol. 10: 252. 1941.

Amauropelta micula (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151020-1

• Thelypteris micula A.R.Sm., Fieldiana, Bot., n.s., 29: 33. 1992.

Amauropelta minima (A.R.Sm. & M.Kessler)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151022-1

• Thelypteris minima A.R.Sm. & M.Kessler, Brittonia 60(1): 55. 2008.

Amauropelta minutula (C.V.Morton)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151024-1

• Thelypteris minutula C.V.Morton, Amer. Fern J. 43(4): 173. 1953.

Amauropelta mombachensis (L.D.Gómez)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151286-1

• Thelypteris mombachensis L.D.Gómez, Phytologia 52(3): 155–156. 1982.

Amauropelta mortonii (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151026-1

• Thelypteris mortonii A.R.Sm., Amer. Fern J. 63(3): 125. f 11. 1973.

Amauropelta mosenii (C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151287-1

• Dryopteris mosenii C.Chr., Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 4: 300. f 27. 1907.

• Thelypteris mosenii (C.Chr.) C.F.Reed, Phytologia 17(4): 294. 1968.

Amauropelta muscicola (Proctor)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151028-1

• Thelypteris muscicola Proctor, Rhodora 63: 33. 1961.

Amauropelta namaphila (Proctor)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151030-1

• Thelypteris namaphila Proctor, Amer. Fern J. 75(2): 56. f 1. 1985.

Amauropelta neglecta (Brade & Rosenst.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151032-1

• Dryopteris neglecta Brade & Rosenst., Bol. Mus. Nac. Rio de Janeiro 7(3): 142, t.1, f 3, t. 7. 1931.

• Thelypteris neglecta (Brade & Rosenst.) Ching, Bull. Fan Mem. Inst. Biol. Bot. 10: 253. 1941.

Amauropelta negligens (Jenman)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151033-1

• Nephrodium negligens Jenman, Bull. Bot. Dept. 3: 21. 1896.

• Thelypteris negligens (Jenman) Proctor, Amer. Fern J. 71(2): 58. 1981.

Amauropelta nephelium (A.R.Sm. & M.Kessler)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151036-1

• Thelypteris nephelium A.R.Sm. & M.Kessler, Brittonia 60(1): 57. 2008.

Amauropelta nitens (Desv.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151038-1

• Polypodium nitens Desv., Mém. Soc. Linn. Paris 6: 240. 1827.

• Thelypteris nitens (Desv.) R.M.Tryon, Rhodora 69(777): 7. 1967.

Amauropelta novaeana (Brade)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151288-1

• Dryopteris novaeana Brade, Arq. Inst. Biol. Veg. 3: 2, t. 2, 4(4-6), 6(4-5). 1936.

• Thelypteris novaena (Brade) Ponce, Novon 8(3): 275. 1998.

Amauropelta nubicola (de la Sota)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151040-1

• Thelypteris nubicola de la Sota, Darwiniana 18: 225. pl 4, 8b. 1973.

Amauropelta nubigena (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151042-1

• Thelypteris nubigena A.R.Sm., Proc. Calif. Acad. Sci., ser. 4, 40(8): 229. f 7C. 1975.

Amauropelta oaxacana (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151044-1

• Thelypteris oaxacana A.R.Sm., Amer. Fern J. 63(3): 125. f 7-8. 1973.

Amauropelta ophiorhizoma (A.R.Sm. & Lellinger)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151046-1

• Thelypteris ophiorhizoma A.R.Sm. & Lellinger, Proc. Biol. Soc. Wash. 98(4): 925. f 8. 1985.

Amauropelta oviedoae (C.Sánchez & Zavaro)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151048-1

• Thelypteris oviedoae C.Sánchez & Zavaro, Fontqueria 31: 225. f 2. 1991.

Amauropelta pachyrhachis (Kunze ex Mett.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151289-1

• Aspidium pachyrhachis Kunze ex Mett., Abh. Senckenberg. Naturf. Ges. 2: 367. 1858.

• Thelypteris pachyrhachis (Kunze ex Mett.) Ching, Bull. Fan Mem. Inst. Biol. 10: 253. 1941.

Amauropelta paleacea (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151049-1

• Thelypteris paleacea A.R.Sm., Fl. Ecuador 18: 62. 1983.

Amauropelta patula (Fée)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151052-1

• Gymnogramma patula Fée, Crypt. vasc. Brésil 1: 59, t. 14, f. 3. 1869.

Amauropelta pavoniana (Klotzsch)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151054-1

• Polypodium pavonianum Klotzsch, Linnaea 20. 386. 1847.

• Thelypteris pavoniana (Klotzsch) R.M.Tryon, Rhodora 69(777): 7. 1967.

Amauropelta pelludia (A.R.Sm. & M.Kessler)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151056-1

• Thelypteris pelludia A.R.Sm. & M.Kessler, Brittonia 60(1): 58. 2008.

Amauropelta peradenia (A.R.Sm.)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77151058-1

• Thelypteris peradenia A.R.Sm., Ann. Missouri Bot. Gard. 77(2): 272, f. 11. 1990.

Amauropelta peruviana (Rosenst.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151292-1

• Dryopteris peruviana Rosenst., Repert. Spec. Nov. Regni Veg. 7: 298-299. 1909.

• Thelypteris peruviana (Rosenst.) R.M.Tryon, Rhodora 69(777): 7. 1967.

Amauropelta phacelothrix (Rosenst.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151293-1

• Dryopteris phacelothrix Rosenst., Repert. Spec. Nov. Regni Veg. 11: 56. 1912.

• Thelypteris phacelothrix (Rosenst.) R.M.Tryon, Rhodora 69(777): 7. 1967.

Amauropelta physematioides (Kuhn & Christ ex Krug)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151060-1

• Aspidium physematioides Kuhn & Christ ex Krug, Bot. Jahrb. Syst. 24: 115. 1897.

• Thelypteris physematioides (Kuhn & Christ ex Krug) C.V.Morton, Amer. Fern J. 43(4): 174. 1953.

Amauropelta piedrensis (C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151294-1

• Dryopteris piedrensis C.Chr., Smithsonian Misc. Collect. 52: 372. 1909.

• Thelypteris piedrensis (C.Chr.) C.V.Morton, Amer. Fern J. 53(2): 69. 1963.

Amauropelta pilosissima (C.V.Morton)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77151062-1

• Thelypteris pilosissima C.V.Morton, Fieldiana, Botany 28: 11. 1951.

Amauropelta pilosohispida (Hook.)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77151421-1

• Nephrodium pilosohispidum Hook., Spec. Fil. 4: 105. 1862.

• Thelypteris pilosohispida (Hook.) Alston, J. Wash. Acad. Sci. 48(7): 233. 1958.

Amauropelta pleiophylla (Sehnem)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151295-1

• Dryopteris pleiophylla Sehnem, Fl. Ilustr. Catarin. 1(Aspid.): 245. f 61. 1979.

• Thelypteris pleiophylla (Sehnem) Ponce, Darwiniana 33(1-4): 270. 1995.

Amauropelta podotricha (Sehnem)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151296-1

• Dryopteris podotricha Sehnem, Fl. Ilustr. Catarin. 1(Aspid.): 232. f 57. 1979.

• Thelypteris podotricha (Sehnem) Ponce, Darwiniana 33(1-4): 270. 1995.

Amauropelta proboscidea (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151064-1

• Thelypteris proboscidea A.R.Sm., Fieldiana, Bot., n.s., 29: 36. 1992.

Amauropelta proctorii (A.R.Sm. & Lellinger)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77151297-1

• Thelypteris proctorii A.R.Sm. & Lellinger, Proc. Biol. Soc. Wash. 98(4): 925. f 9. 1985.

Amauropelta prolatipedis (Lellinger)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77151298-1

• Thelypteris prolatipedis Lellinger, Amer. Fern J. 67(2): 60, 1977.

• Dryopteris coarctata var. longipes C.Chr., Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 4: 292. 1907.

Amauropelta ptarmiciformis (C.Chr. & Rosenst. ex Rosenst.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151065-1

• Dryopteris ptarmiciformis C.Chr. & Rosenst. ex Rosenst., Repert. Spec. Nov. Regni Veg. 12: 472. 1913.

• Thelypteris ptarmiciformis (C.Chr. & Rosenst. ex Rosenst.) C.F.Reed, Phytologia 17(4): 307. 1968.

Amauropelta pteroidea (Klotzsch)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77151068-1

• Polypodium pteroideum Klotzsch, Linnaea 20. 389. 1847.

• Thelypteris pteroidea (Klotzsch) R.M.Tryon, Rhodora 69(777): 8. 1967.

Amauropelta pusilla (Mett.)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77151070-1

• Aspidium pusillum Mett., Ann. Sci. Nat., Bot., sér. 5, 2: 245. 1864.

• Thelypteris pusilla (Mett.) Ching, Bull. Fan Mem. Inst. Biol. 10: 254. 1941.

Amauropelta raddii (Rosenst.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151072-1

• Dryopteris raddii Rosenst., Hedwigia 56: 367. 1915.

• Thelypteris raddii (Rosenst.) Ponce, Darwiniana 33(1-4): 266. 1995.

Amauropelta randallii (Maxon & C.V.Morton ex C.V.Morton)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151074-1

• Thelypteris randallii Maxon & C.V.Morton ex C.V.Morton, Amer. Fern J. 53(2): 69. 1963.

Amauropelta recumbens (Rosenst.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151299-1

• Dryopteris recumbens Rosenst., Hedwigia 46. 123. 1906.

• Thelypteris recumbens (Rosenst.) C.F.Reed, Phytologia 17: 308. 1968.

Amauropelta reducta (C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151076-1

• Dryopteris reducta C.Chr., Kongl. Svenska Vetenskapsakad. Handl., ser. 3, 16(2): 18, t. 2, f. 1-3. 1937.

Amauropelta regnelliana (C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151300-1

• Dryopteris regnelliana C.Chr., Kongel. Danske.Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 4: 284, f. 12. 1907.

• Thelypteris regnelliana (C.Chr.) Ponce, Darwiniana 33(1-4): 264. 1995.

Amauropelta retrorsa (Sodiro)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151078-1

• Nephrodium retrorsum Sodiro, Recens. Crypt. Vasc. Quit. 51. 1883.

• Thelypteris retrorsa (Sodiro) A.R.Sm., Fl. Ecuador 18: 71. 1983.

Amauropelta rheophyta (Proctor)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151080-1

• Thelypteris rheophyta Proctor, Amer. Fern J. 75(2): 58. f 2. 1985.

Amauropelta rigescens (Sodiro)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151081-1

• Nephrodium rigescens Sodiro, Crypt. Vasc. Quit. 239. 1893.

• Thelypteris rigescens (Sodiro) A.R.Sm., Fl. Ecuador 18: 72. 1983.

Amauropelta rivularioides (Fée)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151084-1

• Aspidium rivularioides Fée, Crypt. Vasc. Brésil 1: 145, t. 50, f. 1. 1869.

• Thelypteris rivularioides (Fée) Abbiatti, Revista Mus. La Plata, Secc. Bot. 9(36-37): 19. 1958.

Amauropelta roraimensis (Baker)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77151301-1

• Polypodium roraimense Baker, Timehri 5: 214. 1886.

• Thelypteris roraimensis (Baker) C.F.Reed, Phytologia 17(4): 310. 1968.

Amauropelta rosenstockii (C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151086-1

• Dryopteris rosenstockii C.Chr., Kongel. Danske.Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 4: 304, f. 30. 1907.

• Thelypteris rosenstockii (C.Chr.) R.M.Tryon, Rhodora 69(777): 8. 1977.

Amauropelta rosulata (A.R.Sm. & M.Kessler)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151088-1

• Thelypteris rosulata A.R.Sm. & M.Kessler, Brittonia 60(1): 60. 2008.

Amauropelta rudiformis (C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151090-1

• Dryopteris rudiformis C.Chr., Univ. Calif. Publ. Bot. 19(9): 305. 1941.

• Thelypteris rudiformis (C.Chr.) A.R.Sm., Fl. Ecuador 18: 74. 1983.

Amauropelta rufa (Poir.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151302-1

• Polypodium rufum Poir., Encycl. 5: 532. 1804.

• Thelypteris rufa (Poir.) A.R.Sm. Fl. Ecuador 18: 77. 1983.

Amauropelta ruiziana (Klotzsch)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151092-1

• Polypodium ruizianum Klotzsch, Linnaea 20: 385. 1847.

• Thelypteris ruiziana (Klotzsch) A.R.Sm. Fieldiana, Bot., n.s. 29: 35. 1992.

Amauropelta rupestris (Klotzsch)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77151094-1

• Leptogramma rupestris Klotzsch, Linnaea 20. 415. 1847.

• Thelypteris rupestris (Klotzsch) C.F.Reed, Phytologia 17(4): 310. 1968.

Amauropelta rupicola (C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151303-1

• Dryopteris rupicola C.Chr., Repert. Spec. Nov. Regni Veg. 15: 24. 1917.

• Thelypteris rupicola (C.Chr.) Ching, Bull. Fan Mem. Inst. Biol. 10: 254. 1941.

Amauropelta rustica (Fée)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151096-1

• Phegopteris rustica Fée, Mém. Foug. 11 (Hist. Foug. Ant.): 55, t. 13, f. 1. 1866.

• Thelypteris rustica (Fée) Proctor, Rhodora 61(732): 306. 1959[1960].

Amauropelta sanctae-catharinae (Rosenst.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151097-1

• Dryopteris sanctae-catharinae Rosenst., Hedwigia 46: 126. 1906.

• Thelypteris sanctae-catharinae (Rosenst.) Ponce, Darwiniana 33(1-4): 280. 1995.

Amauropelta saxicola (Sw.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151100-1

• Polypodium saxicola Sw., Kongl. Vetensk. Acad. Handl. 1817: 59, t. 3, f. 5. 1817.

• Thelpyteris saxicola (Sw.) C.F.Reed, Phytologia 17(4): 312. 1968.

Amauropelta scalpturoides (Fée)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151102-1

• Phegopteris scalpturoides Fée, Mém. foug. 11 (Hist. foug. Ant.): 51. 1866.

• Thelypteris scalpturoides (Fée) C.F.Reed, Phytologia 17(4): 312. 1968.

Amauropelta sellensis (C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151104-1

• Dryopteris sellensis C.Chr., Kungl. Svenska Vetenskapsakad. Handl., ser. 3, 16(2): 24. t. 3(7−8). 1937.

Amauropelta semilunata (Sodiro)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151106-1

• Nephrodium semilunatum Sodiro, Recens. Crypt. Vasc. Quit. 46. 1883.

• Thelyteris semilunata (Sodiro) A.R.Sm., Fl. Ecuador 18: 79. 1983.

Amauropelta shaferi (Maxon & C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151304-1

• Dryopteris shaferi Maxon & C.Chr., Amer. Fern J. 4: 77. 1914.

• Thelypteris shaferi (Maxon & C.Chr.) Duek, Adansonia, sér. 2, 11: 719. 1971 [1972].

Amauropelta soridepressa (Salino & V.A.O.Dittrich)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151108-1

• Thelypteris soridepressa Salino & V.A.O.Dittrich, Amer. Fern J. 98(4): 199. f 1. 2009.

Amauropelta steyermarkii (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151110-1

• Thelypteris steyermarkii A.R.Sm., Fl. Ecuador 18: 79. 1983.

Amauropelta stierii (Rosenst.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151305-1

• Gymnogramma stierii Rosenst., Festschrift Albert von Bamberg 64. 1905.

• Thelypteris stierii (Rosenst.) C.F.Reed, Phytologia 17(4): 316. 1968.

Amauropelta straminea (Sodiro)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151112-1

• Nephrodium stramineum Sodiro, Recens. Crypt. Vasc. Quit. 43. 1883.

• Thelypteris chimboracensis A.R.Sm., Fl. Ecuador 18: 34. 1993.

Amauropelta strigillosa (A.R.Sm. & Lellinger)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151113-1

• Thelypteris strigillosa A.R.Sm. & Lellinger, Proc. Biol. Soc. Wash. 98(4): 927. F 10. 1985.

Amauropelta struthiopteroides (C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151306-1

• Dryopteris struthiopteroides C.Chr., Smithsonian Misc. Collect. 52(3): 388. 1909.

• Thelypteris struthiopteroides (C.Chr.) C.F.Reed, Phytologia 17(4): 316. 1968.

Amauropelta subscandens (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151116-1

• Thelypteris subscandens A.R.Sm., Ann. Missouri Bot. Gard. 77(1): 123. f 4. 1990.

Amauropelta subtilis (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151118-1

• Thelypteris subtilis A.R.Sm., Fl. Ecuador 18: 80. 1983.

Amauropelta supina (Sodiro)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151307-1

• Nephrodium supinum Sodiro, Crypt. Vasc. Quit. 241. 1893.

• Thelypteris supina (Sodiro) A.R.Sm., Fl. Ecuador 18: 82. 1983.

Amauropelta tablana (Christ)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151120-1

• Aspidium tablanum Christ, Bull. Herb. Boissier, sér. 2, 5: 727. 1905.

• Thelypteris tablana (Christ) A.R.Sm., Amer. Fern J. 63(3): 127. 1973.

Amauropelta tamandarei (Rosenst.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151122-1

• Dryopteris tamandarei Rosenst., Hedwigia 56. 365. 1915.

• Thelypteris tamandarei (Rosenst.) Ponce, Novon 8(3): 277. 1998.

Amauropelta tapantensis (A.R.Sm. & Lellinger)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151124-1

• Thelypteris tapantensis A.R.Sm. & Lellinger, Proc. Biol. Soc. Wash. 98(4): f 11. 927. 1985.

Amauropelta tenerrima (Fée)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151126-1

• Aspidium tenerrimum Fée, Crypt. Vasc. Brésil 1: 134. t 43, f 1. 1869.

• Thelypteris tenerrima (Fée) C.F.Reed, Phytologia 17(4): 319. 1968.

Amauropelta trelawniensis (Proctor)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151308-1

• Thelypteris trelawniensis Proctor, Amer. Fern J. 71(2): 58. 1981.

Amauropelta uncinata (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151128-1

• Thelypteris uncinata A.R.Sm., Fl. Ecuador 18: 86. 1983.

Amauropelta vattuonei (Hicken)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151129-1

• Dryopteris vattuonei Hicken, Darwiniana 1: 100. 1924.

• Thelypteris vattuonei (Hicken) Abbiatti, Darwiniana 13: 566. 1964.

Amauropelta venturae (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151132-1

• Thelypteris venturae A.R.Sm., Mem. New York Bot. Gard. 88: 638. f 302A-C. 2004.

Amauropelta vernicosa (A.R.Sm. & Lellinger)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151134-1

• Thelypteris vernicosa A.R.Sm. & Lellinger, Proc. Biol. Soc. Wash. 98(4): 929. f 12. 1985.

Amauropelta villana (L.D.Gómez)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151136-1

• Thelypteris villana L.D.Gómez, Rev. Biol. Trop. 17(1): 106. f 1-4. 1970.

Amauropelta yungensis (A.R.Sm. & M.Kessler)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151138-1

• Thelypteris yungensis A.R.Sm. & M.Kessler, Brittonia 60(1): 61. 2008.

Amauropelta zurquiana (A.R.Sm. & Lellinger)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151140-1

• Thelypteris zurquiana A.R.Sm. & Lellinger, Proc. Biol. Soc. Wash. 98(4): 929. f 13. 1985.

New combinations in Goniopteris

Goniopteris abdita (Proctor)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151141-1

• Thelypteris abdita Proctor Amer. Fern J. 75(2): 63. f 4. 1985.

Goniopteris abrupta (Desv.)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77151144-1

• Polypodium abruptum Desv., Mém. Soc. Linn. Paris 6: 239. 1827.

• Thelypteris abrupta (Desv.) Proctor, Rhodora 61: 305. 1960.

Goniopteris alan-smithiana (L.D.Gómez)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151354-1

• Thelypteris alan-smithiana L.D.Gómez, Phytologia 50(7): 458. 1982.

Goniopteris amazonica (Salino & R.S.Fernandes)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151310-1

• Thelypteris amazonica Salino & R.S.Fernandes, Nordic J. Bot. 29(5): 611. f 1-2. 2011.

Goniopteris ancyriothrix (Rosenst.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151311-1

• Dryopteris ancyriothrix Rosenst., Repert. Spec. Nov. Regni Veg. 7: 305-306. 1909.

• Thelypteris ancyriothrix (Rosenst.) A.R.Sm., Fl. Ecuador 18: 140. 1983.

Goniopteris anoptera (Kunze ex Kuhn)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151312-1

• Aspidium anopteron Kunze ex Kuhn, Linnaea 36. 113. 1869.

• Thelypteris anoptera (Kunze ex Kuhn) C.F.Reed, Phytologia 17(4): 260. 1968.

Goniopteris aureola (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151148-1

• Thelypteris aureola A.R.Sm., Ann. Missouri Bot. Gard. 77(1): 118. f 1D-E. 1990.

Goniopteris beckeriana (F.B.Matos, A.R.Sm. & Labiak)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151150-1

• Thelypteris beckeriana F.B.Matos, A.R.Sm. & Labiak, Brittonia 62(2): 149. f 1. 2010.

Goniopteris berlinii (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151152-1

• Thelypteris berlinii A.R.Sm., Novon 16: 431. 2006.

Goniopteris bibrachiata (Jenman)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151154-1

• Nephrodium bibrachiatum Jenman, Gard. Chron., ser. 3, 15: 230. 1894.

• Thelypteris bibrachiata (Jenman) Proctor, Ferns Jamaica 337. 1985.

Goniopteris biformata (Rosenst.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151355-1

• Dryopteris biformata Rosenst., Repert. Spec. Nov. Regni Veg. 7: 300-301. 1909.

• Thelypteris biformata (Rosenst.) R.M.Tryon, Rhodora 69(777): 5. 1967.

Goniopteris blanda (Fée)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151156-1

• Phegopteris blanda Fée, Mém. Foug. 8: 91. 1857.

• Thelypteris blanda (Fée) C.F.Reed, Phytologia 17(4): 264. 1968.

Goniopteris calypso (L.D.Gómez)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151157-1

• Thelypteris calypso L.D.Gómez, Brenesia 8: 98. 1976.

Goniopteris chocoensis (A.R.Sm. & Lellinger)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151160-1

• Thelypteris chocoensis A.R.Sm. & Lellinger, Proc. Biol. Soc. Wash. 98(4): 916. f 1. 1985.

Goniopteris clypeata (Maxon & C.V.Morton)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151162-1

• Dryopteris clypeata Maxon & C.V.Morton, Bull. Torrey Bot. Club 66: 52. 1939.

• Thelypteris clypeata (Maxon & C.V.Morton) K.U.Kramer, Acta Bot. Neerl. 18: 141. 1969.

Goniopteris cordata (Fée)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151164-1

• Phegopteris cordata Fée, Mém. Foug. 5 (Gen. filic.): 244. 1852.

• Thelypteris cordata (Fée) Proctor, Bull. Inst. Jamaica, Sci. Ser. 5: 59. 1953.

Goniopteris costaricensis

Salino & T.E.Almeida nom. nov.

urn:lsid:ipni.org:names:77151356-1

• Thelypteris crenata A.R.Sm. & Lellinger, Proc. Biol. Soc. Wash. 98(4): 919. f 4. 1985.

• Goniopteris costaricensis This needs a new name in Goniopteris because of the prior Goniopteris crenata C. Presl, 1836.

Goniopteris crassipila (Caluff & C.Sánchez)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151314-1

• Thelypteris crassipila Caluff & C.Sánchez, Willdenowia 34: 513. 2004.

Goniopteris croatii (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151168-1

• Thelypteris croatii A.R.Sm., Ann. Missouri Bot. Gard. 77(1): 122. f 1A-C. 1990.

Goniopteris curta (Christ)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77151170-1

• Dryopteris curta Christ, Bull. Herb. Boissier, sér. 2, 7(4): 263. 1907.

• Thelypteris curta (Christ) C.F.Reed, Phytologia 17(4): 270. 1968.

Goniopteris dissimulans (Maxon & C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151172-1

• Dryopteris dissimulans Maxon & C.Chr., Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10: 215. 1913.

• Thelypteris dissimulans (Maxon & C.Chr.) C.F.Reed, Phytologia 17(4): 273. 1968.

Goniopteris equitans (Christ)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151173-1

• Nephrodium equitans Christ, Bull. Herb. Boissier, sér. 2, 6(2): 163. 1906.

• Thelypteris equitans (Christ) C.F.Reed, Phytologia 17(4): 275. 1968.

Goniopteris erythrothrix (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151176-1

• Thelypteris erythrothrix A.R.Sm., Fieldiana, Bot., n.s., 29: 59. 1992.

Goniopteris gonophora (Weath.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151357-1

• Dryopteris gonophora Weath., Lloydia 2: 164. f. 1(1−3). 1939.

• Thelypteris gonophora (Weath.) A.R.Sm., Ann. Missouri Bot. Gard. 77(1): 203. 1990.

Goniopteris hildae (Proctor)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151315-1

• Thelypteris hildae Proctor, Amer. Fern J. 75(2): 67. f 6. 1985.

Goniopteris holodictya (K.U.Kramer)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151316-1

• Thelypteris holodictya K.U.Kramer, Acta Bot. Neerl. 18: 140. f. 2. 1969.

Goniopteris hondurensis (L.D.Gómez)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151358-1

• Thelypteris hondurensis L.D.Gómez, Phytologia 50(7): 458. 1982.

Goniopteris imitata (C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151317-1

• Dryopteris imitata C.Chr., Kongl. Svenska Vetenskapsakad. Handl., ser. 3, 16(2): 29, t. 6, f. 1-4. 1937.

• Thelypteris imitata (C.Chr.) Alain, Moscosoa 3: 46. 1978.

Goniopteris indusiata (Salino)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151182-1

• Thelypteris indusiata Salino, Phytotaxa 156(5): 280. f 1-3. 2014.

Goniopteris jamesonii (Hook.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151184-1

• Nephrodium jamesonii Hook., Sp. fil. 4: 66. 1862.

• Thelypteris jamesonii (Hook.) R.M.Tryon, Rhodora 69(777): 6. 1967.

Goniopteris jarucoensis (Caluff & C.Sánchez)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151186-1

• Thelypteris jarucoensis Caluff & C.Sánchez, Willdenowia 34: 515. 2004.

Goniopteris killipii (A.R.Sm. & Lellinger)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151188-1

• Thelypteris killipii A.R.Sm. & Lellinger, Proc. Biol. Soc. Wash. 98(4): 923. f 7. 1985.

Goniopteris leonina (Caluff & C.Sánchez)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151189-1

• Thelypteris leonina Caluff & C.Sánchez, Willdenowia 34: 517. 2004.

Goniopteris levyi (E.Fourn.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151359-1

• Aspidium levyi E.Fourn., Bull. Soc. Bot. France 19: 255. 1872.

• Thelypteris levyi (E.Fourn.) C.V.Morton, Contr. U.S. Natl. Herb. 38: 37. 1967.

Goniopteris liebmannii (Maxon & C.V.Morton)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151192-1

• Polypodium meniscioides Liebm., Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 5, 1: 211. 1849.

• Dryopteris liebmannii Maxon & C.V.Morton, Bull. Torrey Bot. Club 65: 348. 1938.

• Thelypteris meniscioides (Liebm.) C.F.Reed, Phytologia 17: 292. 1968.

• Goniopteris liebmannii (Maxon & C.V.Morton) The earliest species ephitet, meniscioides, has been used already in Goniopteris – Goniopteris meniscioides Fée, a synonym of Ampelopteris prolifera (Retz.) Copel., a Paleotropical species.

Goniopteris littoralis (Salino)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151360-1

• Thelypteris littoralis Salino, Brittonia 54(4): 332. f 1. 2002.

Goniopteris lugubriformis (Rosenst.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151319-1

• Dryopteris lugubriformis Rosenst., Repert. Spec. Nov. Regni Veg. 7: 299. 1909.

• Thelypteris lugubriformis (Rosenst.) R.M.Tryon, Rhodora 69(777): 7. 1967.

Goniopteris martinezii (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151194-1

• Thelypteris martinezii A.R.Sm., Mem. New York Bot. Gard. 88: 654-655. f 308I. 2004.

Goniopteris minor (C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151196-1

• Dryopteris nicaraguensis (E.Fourn.) C.Chr. var. minor C.Chr., Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10: 252. 1913.

• Thelypteris minor (C.Chr.) A.R.Sm., Phytologia 34(4): 232. 1976.

Goniopteris minutissima (Caluff & C.Sánchez)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151198-1

• Thelypteris minutissima Caluff & C.Sánchez, Willdenowia 34: 511. 2004.

Goniopteris montana (Salino)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151361-1

• Thelypteris montana Salino, Brittonia 54(4): 334. f 2D-F. 2002.

Goniopteris multigemmifera (Salino)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151321-1

• Thelypteris multigemmifera Salino, Brittonia 54(4): 336. f 3. 2002.

Goniopteris munchii (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151200-1

• Thelypteris munchii A.R.Sm., Amer. Fern J. 63(3): 120. f 9-10. 1973.

Goniopteris nicaraguensis (E.Fourn.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151202-1

• Phegopteris nicaraguensis E.Fourn., Bull. Soc. Bot. France 19: 252. 1872.

• Thelypteris nicaraguensis (E.Fourn.) C.V.Morton, Contr. U.S. Natl. Herb. 38(2): 55. 1967.

Goniopteris nigricans (Ekman & C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151204-1

• Dryopteris nigricans Ekman & C.Chr., Kongl. Svenska Vetenskapsakad. Handl., ser. 3, 16(2): 31, t. 7, f. 1-2. 1937.

Goniopteris oroniensis (L.D.Gómez)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151325-1

• Thelypteris oroniensis L.D.Gómez, Amer. Fern J. 68(1): 9. f 1. 1978.

Goniopteris paranaensis (Salino)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151323-1

• Thelypteris paranaensis Salino, Brittonia 54(4): 337. F 2A-C. 2002.

Goniopteris paucijuga (Klotzsch)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77151205-1

• Aspidium paucijugum Klotzsch, Linnaea 20: 368. 1847.

• Thelypteris paucijuga (Klotzsch) A.R.Sm., Univ. Calif. Publ. Bot. 59: 98. 1971

Goniopteris paucipinnata (Donn.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151362-1

• Nephrodium fendleri D.C.Eaton var. paucipinnatum Donn.Sm., Bot. Gaz. 12: 134. 1887.

• Thelypteris paucipinnata (Donn.Sm.) C.F.Reed, Phytologia 17(4): 302. 1968.

Goniopteris pellita (Willd.)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77151324-1

• Aspidium pellitum Willd., Sp. Pl. ed. 4. 5: 242. 1810.

• Thelypteris pellita (Willd.) Proctor & Lourteig, Bradea 5(40): 384. 1990.

Goniopteris peripae (Sodiro)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151326-1

• Nephrodium peripae Sodiro, Recens. Crypt. Vasc. Quit. 52. 1883.

• Thelypteris peripae (Sodiro) C.F.Reed, Phytologia 17(4): 303. 1968.

Goniopteris pilonensis (A.R.Sm. & M.Kessler)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151210-1

• Thelypteris pilonensis A.R.Sm. & M.Kessler, Brittonia 60(1): 58. 2008.

Goniopteris pinnatifida (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151212-1

• Thelypteris pinnatifida A.R.Sm., Fl. Ecuador 18: 110. 1983.

Goniopteris praetermissa (Maxon)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151214-1

• Dryopteris praetermissa Maxon, Proc. Biol. Soc. Wash. 57: 20. 1944.

• Thelypteris praetermissa (Maxon) A.R.Sm., Phytologia 34(3): 232. 1976.

Goniopteris redunca (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151216-1

• Thelypteris redunca A.R.Sm., Ann. Missouri Bot. Gard. 77(1): 123. f 1F-H. 1990.

Goniopteris resiliens (Maxon)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151218-1

• Dryopteris resiliens Maxon, Field Mus. Publ. Bot. 17: 302. 1938.

• Thelypteris resiliens (Maxon) A.R.Sm., Amer. Fern J. 63(3): 121. 1973.

Goniopteris retroflexa (L.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151327-1

• Polypodium retroflexum L., Sp. Pl. 1089. 1753.

• Thelypteris retroflexa (L.) Proctor & Lourteig, Bradea 5(40): 385. 1990.

Goniopteris rhachiflexuosa (Riba)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151329-1

• Thelypteris rhachiflexuosa Riba, Amer. Fern J. 79(3): 122. f 1. 1989.

Goniopteris ×rolandii (C.Chr.)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77151328-1

• Dryopteris ×rolandii C.Chr., Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10: 258. 1913.

• Thelypteris ×rolandii (C.Chr.) R.M.Tryon, Rhodora 69: 8. 1967.

Goniopteris sapechoana (A.R.Sm. & M.Kessler)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151220-1

• Thelypteris sapechoana A.R.Sm. & M.Kessler, Brittonia 60(1): 60. 2008.

Goniopteris schaffneri (Fée)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151330-1

• Nephrodium schaffneri Fée, Mém. Foug. 8: 108. 1857.

• Thelypteris schaffneri (Fée) C.F.Reed, Phytologia 17(4): 312. 1968.

Goniopteris schippii (Weath.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151331-1

• Dryopteris schippii Weath., Amer. Fern J. 25: 52. 1935.

• Thelypteris schippii (Weath.) A.R.Sm., Phytologia 34(3): 233. 1976.

Goniopteris schomburgkii (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151221-1

• Thelypteris schomburgkii A.R.Sm., Ann. Missouri Bot. Gard. 77(1): 202. f 1. 1990.

Goniopteris schunkei (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151224-1

• Thelypteris schunkei A.R.Sm., Fieldiana, Bot., n.s., 29: 63. 1992.

Goniopteris semihastata (Kunze)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151226-1

• Aspidium semihastatum Kunze, Linnaea 9: 91. 1834.

• Thelypteris semihastata (Kunze) Ching, Bull. Fan Mem. Inst. Biol. 10: 254. 1941.

Goniopteris semirii (Salino & L.C.N.Melo)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151332-1

• Thelypteris semirii Salino & L.C.N.Melo, Novon 10(1): 74. f 1A-e. 2000.

Goniopteris septemjuga (C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151228-1

• Dryopteris septemjuga C.Chr., Kongl. Svenska Vetenskapsakad. Handl., ser. 3, 16(2): 32, t. 7, f. 4-5. 1937.

Goniopteris skinneri (Hook.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151230-1

• Aspidium skinneri Hook., Icon. Pl. 10: t. 924. 1854.

• Thelypteris skinneri (Hook.) C.F.Reed, Phytologia 17(4): 314. 1968

Goniopteris stephanii (A.R.Sm. & M.Kessler)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151232-1

• Thelypteris stephanii A.R.Sm. & M.Kessler, Brittonia 60(1): 61. 2008.

Goniopteris stolzeana (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151333-1

• Thelypteris stolzeana A.R.Sm., Phytologia 34(3): 231. 1976.

Goniopteris subsagittata (Maxon & C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151253-1

• Dryopteris subsagittata Maxon & C.Chr., Kongl. Svenska Vetenskapsakad. Handl., ser. 3, 16(2): 28, t. 6, f. 5–6. 1937.

Goniopteris tenebrica (Jenman)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151234-1

• Nephrodium tenebricum Jenman, J. Bot. 20: 326. 1882.

• Thelypteris tenebrica (Jenman) Proctor, Bull. Inst. Jamaica, Sci. Ser. 5: 65. 1953.

Goniopteris tryoniorum (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151334-1

• Thelypteris tryoniorum A.R.Sm., Fieldiana, Bot., n.s., 29: 62. 1992.

Goniopteris tuxtlensis (T.Krömer, Acebey & A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151236-1

• Thelypteris tuxtlensis T.Krömer, Acebey & A.R.Sm., Amer. Fern J. 97(3): 137–139. 2007.

Goniopteris urbani (Sodiro)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151335-1

• Polypodium urbani Sodiro, Crypt. Vasc. Quit. 301. 1893.

• Thelypteris urbani (Sodiro) A.R.Sm., Fl. Ecuador 18: 116. 1983

Goniopteris verecunda (Proctor)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151237-1

• Thelypteris verecunda Proctor, Amer. Fern J. 75(2): 65. f 5. 1985.

Goniopteris yaucoensis (Proctor)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151336-1

• Thelypteris yaucoensis Proctor, Mem. New York Bot. Gard. 53: 208. 1989.

New combinations in Steiropteris

Steiropteris subg. Glaphyropteris (Fée)

A.R.Sm. comb. et stat. nov.

urn:lsid:ipni.org:names:77151337-1

• Glaphyropteris C.Presl ex Fée, Crypt. Vasc. Brésil, 2. 40. 1873.

• Dryopteris subg. Glaphyropteris (C.Presl. ex Fée) C.Chr., Biol. Arb. Til. Eug. Warming 80. 1911.

• Thelypteris subg. Glaphyropteris (C.Presl. ex Fée), Alston, J. Wash. Acad. Sci. 48: 234. 1958.

Steiropteris buchtienii (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151338-1

• Thelypteris buchtienii A.R.Sm., Univ. Calif. Publ. Bot. 76: 10. 1980.

Steiropteris clypeolutata (Desv.) Pic.Serm. var. holmei (Baker)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151240-1

• Nephrodium holmei Baker, Ann. Bot. 5: 317. 1891.

• Thelypteris clypeolutata (Desv.) Proctor var. holmei (Baker) A.R.Sm., Univ. Calif. Publ. Bot. 76: 11. 1980.

Steiropteris comosa (C.V.Morton)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151242-1

• Dryopteris comosa C.V.Morton, J. Wash. Acad. Sci. 28: 528. 1938.

• Thelypteris comosa (C.V.Morton) C.V.Morton, Amer. Fern J. 51(1): 38. 1961.

Steiropteris decussata (L.)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77151339-1

• Polypodium decussatum L., Species Plantarum 2: 1093. 1753.

• Thelypteris decussata (L.) Proctor, Bull. Inst. Jamaica, Sci. Ser. 5: 59. 1953.

Steiropteris decussata (L.) A.R.Sm. var. brasiliensis (C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151340-1

• Dryopteris decussata var. brasiliensis C.Chr., Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10: 161. 1913.

• Thelypteris decussata var. brasiliensis (C.Chr.) A.R.Sm., Univ. Calif. Publ. Bot. 76: 15. 1980.

Steiropteris decussata var. costaricensis (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151341-1

• Thelypteris decussata var. costaricensis A.R.Sm., Univ. Calif. Publ. Bot. 76: 16. 1980.

Steiropteris decussata var. mapiriensis (Rosenst.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151342-1

• Dryopteris mapiriensis Rosenst., Repert. Spec. Nov. Regni Veg. 6: 313. 1909.

• Thelypteris decussata var. mapiriensis (Rosenst.) A.R.Sm., Univ. Calif. Publ. Bot. 76: 16. 1980.

Steiropteris decussata var. velutina (Sodiro)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151343-1

• Polypodium velutinum Sodiro, Recens. Crypt. Vasc. Quit. 59. 1883.

• Thelypteris decussata var. velutina (Sodiro) A.R.Sm., Univ. Calif. Publ. Bot. 76: 17. 1980.

Steiropteris fraseri (Mett. ex Kuhn)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151344-1

• Aspidium fraseri Mett. ex Kuhn, Linnaea 36: 109. 1869.

• Thelypteris fraseri (Mett. ex Kuhn) A.R.Sm., Univ. Calif. Publ. Bot. 76: 32. 1980.

Steiropteris glandulosa (Desv.) Pic.Serm. var. brachyodus (Kunze)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151256-1

• Polypodium brachyodus Kunze, Linnaea 9: 48. 1834.

• Thelypteris glandulosa var. brachyodus (Kunze) A.R.Sm., Phytologia 34: 233. 1976.

Steiropteris glandulosa (Desv.) Pic.Serm. var. longipilosa (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151345-1

• Thelypteris glandulosa (Desv.) Proctor var. longipilosa A.R.Sm., Univ. Calif. Publ. Bot. 76: 22. 1980.

Steiropteris hatschbachii (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151346-1

• Thelypteris hatschbachii A.R.Sm., Univ. Calif. Publ. Bot. 76: 22. 1980.

Steiropteris hottensis (C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151244-1

• Dryopteris hottensis C.Chr., Kongl. Svenska Vetenskapsacad. Handl., ser. 3, 16(2): 26. t 2(4-6). 1937.

• Thelypteris hottensis (C.Chr.) A.R.Sm., Univ. Calif. Publ. Bot. 76: 23. 1980.

Steiropteris leprieurii (Hook.) Pic.Serm. var. costalis (Baker)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77151347-1

• Nephrodium costale Baker, Syn Fil, 2. 495.1874.

• Thelypteris leprieurii (Hook.) R.M.Tryon var. costalis (Baker) A.R.Sm., Univ. Calif. Publ. Bot. 76: 25. 1980.

Steiropteris leprieurii (Hook.) Pic.Serm. var. glandifera (A.R.Sm)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77151348-1

• Thelypteris leprieurii (Hook.) R.M.Tryon var. glandifera A.R.Sm., Univ. Calif. Publ. Bot. 76: 25. 1980.

Steiropteris leprieurii (Hook.) Pic.Serm. var. incana (Christ)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77151258-1

• Aspidium incanum Christ, Hedwigia 44: 367. 1905.

• Thelypteris leprieurii (Hook.) R.M.Tryon var. incana (Christ) A.R.Sm., Univ. Calif. Publ. Bot. 76: 26. 1980.

Steiropteris leprieurii (Hook.) Pic.Serm. var. subcostalis (A.R.Sm.)

A.R.Sm. comb. nov.

urn:lsid:ipni.org:names:77151349-1

• Thelypteris leprieurii (Hook.) R.M.Tryon var. subcostalis A.R.Sm., Univ. Calif. Publ. Bot. 76: 26. 1980.

Steiropteris mexiae (C.Chr. ex Copel.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151246-1

• Dryopteris mexiae C.Chr. ex Copel., Univ. Calif. Publ. Bot., 17: 32. 1932.

• Thelypteris mexiae (C.Chr. ex Copel.) Ching, Bull. Fan Mem. Inst. Biol. 10: 252. 1941.

Steiropteris parva (A.R.Sm. & M.Kessler)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151350-1

• Thelypteris parva A.R.Sm. & M.Kessler, Brittonia 60(1): 57. 2008.

Steiropteris pennellii (A.R.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151351-1

• Thelypteris pennellii A.R.Sm., Univ. Calif. Publ. Bot. 76: 28. 1980.

Steiropteris polyphlebia (C.Chr.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151248-1

• Dryopteris polyphlebia C.Chr., Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 7, 10: 161. 1913.

• Thelypteris polyphlebia (C.Chr.) C.V.Morton, Amer. Fern J. 51(1): 38. 1961.

Steiropteris polypodioides (Raddi)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151250-1

• Ceterach polypodioides Raddi, Opusc. Sci. 3: 284. 1819.

• Thelypteris polypodioides (Raddi) C.F.Reed, Phytologia 7: 305. 1968.

Steiropteris seemannii (J.Sm.)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151353-1

• Phegopteris seemannii J.Sm., in Seem., Bot. Voy. Herald. 228. 1854.

• Thelypteris seemannii (J.Sm.) A.R.Sm., Univ. Calif. Publ. Bot. 76: 30. 1980.

Steiropteris villosa (Link)

Salino & T.E.Almeida comb. nov.

urn:lsid:ipni.org:names:77151252-1

• Gymnogramma villosa Link, Hort. Berol. 2: 51. 1833.

• Thelypteris villosa (Link) C.F.Reed, Phytologia 7: 323. 1968.

Citation

Salino A, Almeida TE, Smith AR (2015) New combinations in Neotropical Thelypteridaceae. PhytoKeys 57: 11–50. doi: 10.3897/phytokeys.57.5641