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. 2016 Apr 21;4:e1897. doi: 10.7717/peerj.1897

Table 4. Eocene fossil record of Quercus.

Locality, site, age Reference Taxon, organ Affinity
Clarno Fm., Oregon, western U.S.A.; ∼48 Ma Manchester, 1994 Quercuspaleocarpa Manchester; cupules and acorns Group Cyclobalanopsis (? Lithocarpus)
Axel-Heiberg Island, Canadian Arctic; ∼45 Ma McIntyre, 1991; McIver & Basinger, 1999 Pollen and leaves New World Clade; Quercus Group Quercus/Lobatae
Hareøn, western Greenland; ∼42 Ma Grímsson et al., 2015 Quercus sp. 4–5; pollen Extinct/ancestral type
Quercus sp. 6–7; pollen New World Clade (aff. Group Protobalanus)
Quercus sp. 1–3; pollen Quercus Group Quercus and/or Lobatae
Baltic amber, northern Europe; ∼45 Ma Crepet & Nixon, 1989; Weitschat & Wichard, 2003 Flower and in situ pollen Quercus Group Quercus
Königsaue, near Aschersleben, Germany; middle Eocene (48–38 Ma) Kvaček & Walther, 1989 Quercus subhercynica H Walther & Kvaček; leaf Quercus Group Lobatae
Ube coal-field, southwestern Honshu, Japan; middle Eocene (48–38 Ma) Huzioka & Takahasi, 1970 Cyclobalanopsis nathoi Huzioka & Takahashi; leaf, acorn (?), cupule (?) Quercus, affinity unclear
Changchang, Hainan; middle (?) Eocene (50–35 Ma) Hofmann, 2010 Quercus pollen types 2–8, 10; pollen Quercus, affinity unclear (extinct, Group Quercus/Lobatae?, Group Protobalanus?)
Quercus pollen type 1; pollen Quercus Group Ilex
Quercus pollen type 9; pollen Quercus Group Cyclobalanopsis