Checklist of British and Irish Hymenoptera - Braconidae

Abstract Abstract

Background

The checklist of British and Irish Braconidae is revised, based in large part on the collections of the National Museums of Scotland, Edinburgh, and the Natural History Museum, London. Distribution records are provided at the country level together with extensive synonymy and bibliography.

New information

Of the 1,338 species regarded as valid, presumed native and certainly identified, 83 are here recorded for the first time from the British Isles. One new synonym is established (Dyscritus suffolciensis Morley, 1933 = Syntretus splendidus (Marshall, 1887) syn. nov.)

Introduction

The Braconidae is one of two families of the superfamily Ichneumonoidea, along with the Ichneumonidae. Given the size of each family in Britain (over 1,300 braconids and 2,500 ichneumonids) we are publishing the two checklists separately. This is one part of a series of papers revising the British and Irish list of Hymenoptera, that started with Broad and Livermore (2014a), Broad and Livermore (2014b) and Liston et al. (2014). For the background and rationale behind these British Hymenoptera checklists see Broad (2014). The bases for the taxonomy are Fauna Europaea and Taxapad (Yu et al. 2012) (braconid data for both compiled by Kees van Achterberg) but we have not relied on these sources for data on species occurrence in Britain. Rather, we have relied mostly on the primary literature and collections, especially NMS and BMNH. Indeed, large parts of the checklist rely on original work, identifying specimens in these collections.

Much of the synonymy adopted here is equivalent to the German list (Belokobylskij et al. 2003) but where there are taxonomic differences of opinion, usually van Achterberg’s interpretation has been followed, largely for consistency with Fauna Europaea, but also because of van Achterberg’s experience with many groups of braconids. The biggest departure from these sources is in the generic classification of Microgastrinae, for which we largely follow Mason (1981).

We reference all additions to and deletions from the British list since Huddleston (1978) and record country-level distribution within the British Isles, but regarding Ireland as one. We made much use of O'Connor et al. (1999) for Irish records. Because the braconid literature is extensive and scattered we also provide many taxonomic references. The braconid section of the 1978 British checklist (Huddleston 1978) repeats many of Shenefelt’s (e.g. Shenefelt 1973, Shenefelt 1974) mistakes and these are rectified here.

Figs 1, 2, 3, 4 illustrate a tiny part of the morphological and biological diversity of Braconidae. Shaw and Huddleston (1991) provide an introduction to the varied biology of this family of parasitoid wasps.

Representative British Braconidae: Alysiinae and Aphidiinae

Figure 1a.

Face of Alsysia manducator (Panzer) (Alysiinae), with exodont mandibles for emerging from Diptera puparia. BMNH specimen.

Figure 1b.

Pauesia sp. (Aphidiinae) (courtesy of P. Adams)

Figure 1c.

Aphidius ervi (Haliday) (Aphidiinae): mummified aphid host that the braconid pupates within. BMNH specimen.

Figure 1d.

Praon volucre (Haliday) (Aphidiinae): mummified aphid host with the braconid pupa beneath. BMNH specimen.

Representative British Braconidae: Doryctinae, Meteorinae and Microgastrinae

Figure 2a.

Dendrosoter protuberans (Nees) (Doryctinae) male (courtesy of P. Adams)

Figure 2b.

Zele albiditarsus Curtis (Meteorinae) female (courtesy of E. Brosens)

Figure 2c.

Cotesia affinis (Nees) (Microgastrinae) larvae emerging from their Laothoe populi (L.) (Lepidoptera: Sphingidae) host and then spinning cocoons (courtesy of M. Boddington)

Figure 2d.

Cotesia gonopterygis (Marshall) (Microgastrinae) overwintering cocoon mass ex Gonepteryx rhamni (L.) (Lepidoptera: Pieridae) (M.R. Shaw)

Representative British Braconidae

Figure 3a.

Cotesia spuria (Wesmael) (Microgastrinae) cocoons ex larval host Apocheima hispidaria (D. & S.) (Lepidoptera: Geometridae) (courtesy of J. Voogd)

Figure 3b.

Cotesia vanessae (Reinhard) (Microgastrinae) emerging from larval host Aglais urticae (Linnaeus) (Lepidoptera: Nymphalidae) (courtesy of N. Spring)

Figure 3c.

Microplitis ocellatae (Bouché) (Microgastrinae) cocoons ex larval host Smerinthus ocellatus (Linnaeus) (Lepidoptera: Sphingidae) (courtesy of M. Boddington)

Figure 3d.

Histeromerus mystacinus Wesmael (Rhyssalinae) eggs on pupal host Stictoleptura scutellata (L.) (Coleoptera: Cerambycidae) (M.R. Shaw)

Representative British Braconidae: Sigalphinae

Figure 4a.

Acampsis alternipes (Nees) ovipositing in larval host, Alsophila aescularia (D. & S.) (Lepidoptera: Geometridae) (M.R. Shaw)

Figure 4b.

Acampsis alternipes (Nees) larva emerging from its larval host Alsophila aescularia (D. & S.) (Lepidoptera: Geometridae) to complete its feeding externally (M.R. Shaw)

Materials and methods

For a more detailed description of the background and rationale to the Hymenoptera checklist, see Broad (2014). We have tried to account for every name on the 1978 (Huddleston 1978) checklist and have referenced all additions to and deletions from that list. We provide rather extensive synonymy and citations because if you do not have access to the Taxapad database (Yu et al. 2012) it can be very difficult to trace the fate of names in the voluminous and scattered literature. Conventions and abbreviations are listed below.

[species] taxon deleted from the British and Irish list

BMNH Natural History Museum, London

NMS National Museums of Scotland, Edinburgh

# known introductions occurring only under artificial conditions

? status (including uncertain synonymy) or identification in the British Isles uncertain

misident. has been misidentified as this name

nom. dub. nomen dubium, a name of doubtful status

nom. ob. nomen oblitum, ‘forgotten name’, does not have priority over a younger name

nom. nov. nomen novum, a replacement name

nom. nud. nomen nudum, an unavailable name, with no type specimen

preocc. name preoccupied (junior homonym)

stat. rev. status revocatus, revived status (e.g., raised from synonymy)

unavailable name unavailable under provisions of the ICZN code

var. variety, only available as a valid name under certain provisions of the ICZN code

Word document and spreadsheet versions of the checklist are available in the supplementary materials. Future updates to the British and Irish list will be incorporated in an online version of the checklist at Hymenoptera of the British Isles.

Checklists

Agathidinae

Agathidinae

Haliday, 1833

Notes

Except for Agathis and ‘Bassus’ species (i.e. including Lytopylus and Therophilus), distribution and synonymic data from Nixon (1986).

Agathidini

Nees, 1814

• BASSINI Nees, 1812 invalid

• EUMICRODINI Förster, 1863

• BASSINI Förster, 1869 preocc.

• MICRODINI Ashmead, 1900

• MESOCOELINI Viereck, 1918

• ANEUROBRACONINI Fahringer, 1936

• EARININI Sharkey, 1992

Agathis

Latreille, 1804

• AENIGMOSTOMUS Ashmead, 1900

• METRIOSOMA Szépligeti, 1902

• BAEOGNATHA Kokujev, 1903 synonymy by Achterberg (2011)

• LISSAGATHIS Cameron, 1911

• RHAMPHAGATHIS Tobias, 1962

Notes

Distribution and synonymic data from Nixon (1986) and Simbolotti and Achterberg (1999).

Species of Agathis excluded from the British and Irish list:

[malvacearum Latreille, 1805; syn. panzeri (Jurine, 1807, Ichneumon); metzneriae Muesebeck, 1967] Listed by Huddleston (1978); probably refers to varipes.

Agathis anglica

Marshall, 1885

• longicauda Kokujev, 1895 preocc.

• Agathis anglica ?marshalli Fahringer, 1937

• albanica Fischer, 1957

• syriaca Fischer, 1957

• caucasica Tobias, 1963

• taiwanensis Chou & Sharkey, 1989

Distribution

England, Wales

Agathis assimilis

Kokujev, 1895

• propinqua Kokujev, 1895

• jakowlewi Kokujev, 1895

• sibirica Telenga, 1933

• anchisiades Nixon, 1986

Distribution

Scotland

Notes

added by Nixon (1986)

Agathis breviseta

Nees, 1812

• achterbergi Nixon, 1986

Distribution

England, Scotland, Ireland

Notes

Simbolotti and Achterberg (1999) synonymised Agathis achterbergi Nixon, 1986 under breviseta but it is listed as a valid species in Taxapad (Yu et al. 2012).

Agathis fuscipennis

(Zetterstedt, 1838)

• Microgaster fuscipennis Zetterstedt, 1838

• breviseta misident.

• rostrata misident.

• glabricula Thomson, 1895

• schmiedeknechti Kokujev, 1895

• annulata Fahringer, 1937

• meridionellae Fischer, 1957

• albicostellae Fischer, 1966

• artemesiana Fischer, 1966

Distribution

England, Ireland

Agathis griseifrons

Thomson, 1895

• laticarpa Telenga, 1955

Distribution

England, Ireland

Agathis lugubris

(Förster, 1863)

• Cenostomus lugubris Förster, 1863

• minuta Niezabitowski, 1910

Distribution

England, Scotland, Wales, Ireland

Notes

added by Nixon (1986)

Agathis montana

Shestakov, 1932

• zaykovi Nixon, 1986

Distribution

England

Notes

added by Simbolotti and Achterberg (1999)

Agathis nigra

Nees, 1812

• testaceipes Fischer, 1957

• kasachstanica Tobias, 1963

• nixoni Belokobylskij & Jervis, 1998

Distribution

England

Agathis rufipalpis

Nees, 1812

Distribution

Ireland

Agathis tibialis

Nees, 1814

• genualis Marshall, 1898

Distribution

England

Notes

NMS, det. van Achterberg, added here

Agathis varipes

Thomson, 1895

• simulatrix Kokujev, 1895

• rufipes Ivanov, 1899

• dissimilis Shestakov, 1928

• rufilabialis Fahringer, 1937

• glabricollis Telenga, 1957

• serratulae Tobias, 1963

• lederi Fischer, 1968

• ariadne Nixon, 1986

Distribution

England

Notes

added by Nixon (1986)

Bassus

Fabricius, 1804

• MICRODUS Nees, 1814

• DIPLOZON Haliday, 1833

• EURYZONA Haliday, 1838 nom. nud.

• EUMICRODUS Förster, 1863

• HEMIOGASTER Enderlein, 1920 synonymy by Achterberg and Long (2010)

Notes

Following molecular phylogenetic analysis of Agathidinae (Sharkey et al. 2006), the genus Bassuss.l. was recognised as being a polyphyletic assemblage. Species are now being described in or reassigned to Lytopylus, Thermophilus and the non-British Camptothlipsis (Sharkey et al. 2009, Stevens et al. 2010, Stevens et al. 2011, Achterberg and Long 2010, Achterberg 2011) but very few European species have been formally transferred. The species on the British list have therefore been reassigned here on the bases of recent generic keys (e.g. Sharkey et al. 2009) in anticipation of future taxonomy. Whereas Bassuss. s. is now restricted to a small group of species (only one in Britain), Therophilus remains large and probably para- or polyphyletic. Distribution and synonymic data for Bassus, Lytopylus and Therophilus from Nixon (1986) and Simbolotti and Achterberg (1992).

Bassus calculator

(Fabricius, 1798)

• Ichneumon calculator Fabricius, 1798

• abscissus (Ratzeburg, 1844, Microdus)

Distribution

England

Earinus

Wesmael, 1837

• Earinus Förster, 1863

Earinus elator

(Fabricius, 1804)

• Banchus elator Fabricius, 1804

• nitidulus (Nees, 1814, Microdus)

• thoracicus (Nees, 1834, Microdus)

• major (Fonscolombe, 1846, Agathis)

• pilosus Tobias, 1960

Distribution

England, Scotland, Ireland

Earinus gloriatorius

(Panzer, 1809)

• Bassus gloriatorius Panzer, 1809

• gloriator (Nees, 1812, Microdus)

• ochropes (Curtis, 1829, Microdus) nom. nud.

• affinis (Wesmael, 1837, Microdus)

• delusor (Wesmael, 1837, Microdus)

• tuberculatus (Wesmael, 1837, Microdus)

• varicoxis (Wesmael, 1837, Microdus)

• niger (Zetterstedt, 1838, Microgaster)

• bicingulatus (Thomson, 1895, Agathis)

• ochropes Lyle, 1920

• ruficoxis Fahringer, 1937

Distribution

England, Scotland, Ireland, Isle of Man

Earinus transversus

Lyle, 1920

Distribution

England

Notes

distribution data from Shaw (2005)

Lytopylus

Förster, 1863

• AEROPHILUS Szépligeti, 1902 synonymy by Sharkey et al. (2006)

• NEOMICRODUS Szépligeti, 1908

• AEROPHILOPSIS Viereck, 1913

• AEROPHILINA Enderlein, 1920

• IOXIA Enderlein, 1920

• HORMAGATHIS Brues, 1926

• OBESOMICRODUS Papp, 1971

• FACILAGATHIS van Achterberg & Chen, 2004

Notes

See comments under Bassus; generic synonymy from Sharkey et al. (2009).

Lytopylus rufipes

(Nees, 1814)

• Microdus rufipes Nees, 1814

• germanicus (Enderlein, 1904, Braunsia)

• diversus Muesebeck, 1933

• amurensis (Shestakov, 1940, Microdus)

Distribution

England

Therophilus

Wesmael, 1837

• ORGILONEURA Ashmead, 1900

• AGATHIELLA Szépligeti, 1902

• AEROPHILIODES Strand, 1911

Notes

See comments under Bassus; generic synonymy from Sharkey et al. (2009).

Species excluded from the British and Irish list:

[brevicaudis (Reinhard, 1867, Microdus)] Listed as a ‘species inquirendae’ by Nixon (1986) but included as a German species by Simbolotti and Achterberg (1992). Transferred to Bassus by Simbolotti and Achterberg (1999) but not listed as a species of Therophilus by Sharkey and Stoelb (2012).

[nugax (Reinhard, 1867, Microdus); syn. rufiventris (Abdinbekova, 1975, Microdus)] Listed by Huddleston (1978) but no mention of British specimens by Nixon (1986) or Simbolotti and Achterberg (1992).

Therophilus arcuatus

(Reinhard, 1867)

• Microdus arcuatus Reinhard, 1867

Notes

Taken out of synonymy with conspicuus by Simbolotti and Achterberg (1992).

Therophilus cingulipes

(Nees, 1812)

• Microdus cingulipes Nees, 1812

• nantouensis Chou & Sharkey, 1989

Distribution

England, Ireland

Therophilus clausthalianus

(Ratzeburg, 1844)

• Ichneumon clausthalianus Ratzeburg, 1844

Distribution

England, Ireland

Therophilus conspicuus

(Wesmael, 1837)

• Microdus conspicuus Wesmael, 1837

• zonatus (Marshall, 1885, Earinus)

• carpocapsae Cushman, 1915

• angustatus (Telenga, 1955, Microdus)

• variabilis Chou & Sharkey, 1989

Distribution

England, Ireland

Therophilus tegularis

(Thomson, 1895)

• Agathis tegularis Thomson, 1895

Distribution

England

Notes

added by Simbolotti and Achterberg (1992)

Therophilus tumidulus

(Nees, 1812)

• Microdus tumidulus Nees, 1812

• intermedius (Ivanov, 1899, Eumicrodus)

• annae (Enderlein, 1908, Microdus)

• aino (Watanabe, 1937, Microdus)

• ruficoxis (Fahringer, 1937, Microdus)

• rufus (Fahringer, 1938, Microdus)

• bicolor (Shestakov, 1940, Microdus) preocc.

• victoris (Telenga, 1955, Microdus)

• shestakovi (Shenefelt, 1970, Agathis)

• anuphrievi (Tobias, 1986, Microdus)

Distribution

England, Scotland, Ireland

Therophilus

Wesmael, 1837

Notes

Species incertae sedis within Agathidinae; Sharkey and Stoelb (2012) defined Therophilus as a monophyletic genus and listed the included species, but several species currently classified in Therophilus are of uncertain generic placement.

Therophilus dimidiator

(Nees, 1834)

• Microdus dimidiator Nees, 1834

• cingulator (Ratzeburg, 1852, Microdus)

• laticinctus (Cresson, 1873, Microdus)

• ocellanae (Richardson, 1913, Microdus)

Distribution

England

Notes

Listed by Huddleston (1978) but no mention of British specimens by Nixon (1986), who states, however, that the traditional interpretation of the species is probably correct. No British or Irish specimens were seen by Simbolotti and Achterberg (1992) but van Achterberg has identified an English specimen in BMNH as Therophilus dimidiator.

Therophilus epinotiae

(van Achterberg, 1992)

• Bassus epinotiae van Achterberg, 1992

Distribution

England

Notes

added by Simbolotti and Achterberg (1992)

Therophilus linguarius

(Nees, 1812)

• Microdus linguarius Nees, 1812

• minor (Enderlein, 1908, Microdus)

• kaszabi (Papp, 1967, Vipio)

Distribution

England

Therophilus mediator

(Nees, 1814)

• Microdus mediator Nees, 1814

• lugubrator (Ratzeburg, 1852, Microdus)

Distribution

England

Notes

Transferred from Agathis by Simbolotti and Achterberg (1999).

Therophilus pumilus

(Ratzeburg, 1844)

• Microdus pumilus Ratzeburg, 1844

Distribution

England

Notes

NMS, det. Shaw, added here; listed by Huddleston (1978) but no mention of British specimens by Nixon (1986) or Simbolotti and Achterberg (1992). Transferred from Agathis by Simbolotti and Achterberg (1999) but listed as a species of Agathis in Taxapad (Yu et al. 2012).

Therophilus rugulosus

(Nees, 1834)

• Microdus rugulosus Nees, 1834

• compeditus (Vollenhoven, 1878, Microdus)

• punctatus (Abdinbekova, 1975, Microdus)

Distribution

England, Ireland

Alysiinae

Alysiinae

Leach, 1815

Alysiini

Leach, 1815

• ALLOEINI Ashmead, 1900

Adelurola

Strand, 1928

• ADELURA Förster, 1863 preocc.

• NEOCARPA Fischer, 1966

Adelurola florimela

(Haliday, 1838)

• Alysia florimela Haliday, 1838

• multiarticulata (Marshall, 1898, Phaenocarpa)

• pentapleuroides (Fischer, 1971, Dapsilarthra) unavailable

Distribution

England, Scotland, Ireland, Isle of Man

Alloea

Haliday, 1833

• DIASPASTA Förster, 1863

• LAMADATHA Cameron, 1900

Alloea contracta

Haliday, 1833

• testaceipes (Cameron, 1900, Lamadatha)

Distribution

England, Scotland, Wales, Ireland

Alloea lonchopterae

Fischer, 1966

Distribution

England, Isle of Man

Notes

Added by Godfray and Bland (2011) English record from specimen in BMNH.

Alysia

Latreille, 1804

Alysia

Latreille, 1804

• CECHENUS Illiger, 1807

• BASSUS Nees, 1812

• GONIARCHA Förster, 1863

• STROPHAEA Förster, 1863

Notes

Much of the taxonomy and distribution from Wharton (1986).

Species of Alysia (Alysia) excluded from the British and Irish list:

[cingulata Nees, 1834 nom. dub.]

Alysia (Alysia) alticola

(Ashmead, 1890)

• Pentapleura alticola Ashmead, 1890

• soror Marshall, 1894

Distribution

England, Scotland, Wales, Isle of Man

Alysia (Alysia) frigida

Haliday, 1838

Distribution

England

Notes

added by Godfray and Achterberg (2015)

Alysia (Alysia) incongrua

Nees, 1834

Distribution

England, Scotland, Ireland

Alysia (Alysia) lucia

Haliday, 1838

• diversiceps Fischer, 1967

Distribution

Scotland

Alysia (Alysia) lucicola

Haliday, 1838

Distribution

England, Scotland, Ireland

Alysia (Alysia) luciella

Stelfox, 1941

Distribution

Ireland

Alysia (Alysia) manducator

(Panzer, 1799)

• Ichneumon manducator Panzer, 1799

• haematopa (Gmelin, 1790, Ichneumon) preocc.

• stercoraria Latreille, 1805

• apicalis Curtis, 1826

• similis Curtis, 1826

• stercorator Lamarck, 1835

• curtungula Thomson, 1895

• bucephala Marshall, 1898

• manducatrix Schulz, 1906

Distribution

England, Scotland, Wales, Ireland, Isle of Man

Alysia (Alysia) truncator

(Nees, 1812)

• Bassus truncator Nees, 1812

Distribution

England, Scotland, Ireland

Anarcha

Förster, 1863

Notes

Much of the taxonomy and distribution from Wharton (1988b).

Species of Alysia (Anarcha) excluded from the British and Irish list:

[similis (Nees, 1812, Bassus) nom. dub.]

Alysia (Anarcha) atra

Haliday, 1838

Distribution

Ireland

Alysia (Anarcha) fuscipennis

Haliday, 1838

• obscuripes Thomson, 1895

Distribution

England, Scotland, Ireland

Alysia (Anarcha) mandibulator

(Nees, 1812)

• Bassus mandibulator Nees, 1812

• loripes Haliday, 1838

• mandibulatrix Schulz, 1906

Distribution

England, Ireland

Alysia (Anarcha) rufidens

Nees, 1834

• puncticollis Thomson, 1895

Distribution

England, Ireland

Alysia (Anarcha) sophia

Haliday, 1835

Distribution

Ireland

Alysia (Anarcha) subaperta

Thomson, 1895

• similis misident.

Distribution

England

Notes

added by Wharton (1988b)

Alysia (Anarcha) thapsina

Wharton, 1988

Distribution

Ireland

Notes

added by Wharton (1988b)

Alysia (Anarcha) tipulae

(Scopoli, 1763)

• Ichneumon tipulae Scopoli, 1763

• abdominator (Nees, 1814, Bassus)

• notabilis (Förster, 1863, Anarcha)

Distribution

England, Scotland, Ireland

Alysia (Anarcha) umbrata

Stelfox, 1941

Distribution

Ireland

Anisocyrta

Förster, 1863

Anisocyrta perdita

(Haliday, 1838)

• Alysia perdita Haliday, 1838

Distribution

Scotland

Aphaereta

Förster, 1863

• TRICHESIA Provancher, 1880

• TRINARIA Provancher, 1886

• APHAERETE Dalla Torre, 1898

• ATOPANDRIUM Graham, 1952

• TRISYNALDIS Fischer, 1958

Notes

Atopandrium was synonymised under Aphaereta by Achterberg (1995) but treated as a valid genus again in Taxapad (Yu et al. 2012).

Aphaereta debilitata

Morley, 1933

• loripenne (Graham, 1952, Atopandrium)

• confluctum (Fischer, 1958, Trisynaldis)

Distribution

England, Wales

Aphaereta falcigera

Graham, 1960

Distribution

England, Ireland

Aphaereta major

(Thomson, 1895)

• Alysia major Thomson, 1895

• major Marshall, 1898 preocc.

Distribution

England, Scotland, Ireland

Aphaereta minuta

(Nees, 1811)

• Stephanus minutus Nees, 1811

• cephalotes (Haliday, 1833, Alysia)

• fuscipes (Nees, 1834, Alysia)

• confluens (Ratzeburg, 1844, Alysia)

• stigmaticalis (Thomson, 1895, Alysia)

• inepta Morley, 1933

Distribution

England, Scotland, Ireland

Aphaereta pallipes

(Say, 1829)

• Alysia pallipes Say, 1829

• auripes (Provancher, 1881, Trichesia)

• pilicornis (Provancher, 1886, Trinaria)

• californica Ashmead, 1889

• muscae Ashmead, 1889

• oscinidis Ashmead, 1889

• pallidipes (Dalla Torre, 1898, Aphaerete)

• delosa Viereck, 1905

• subtricarinata Viereck, 1905

• pegomyiae Brues, 1907

• sarcophagae Gahan, 1914

Distribution

England

Notes

added by Shaw (1983)

Aphaereta tenuicornis

Nixon, 1939

Distribution

England, Wales, Ireland

Asobara

Förster, 1863

• SPANISTA Förster, 1863

Asobara tabida

(Nees, 1834)

• Alysia tabida Nees, 1834

• anomala (Thomson, 1895, Alysia)

• crenulata (Fahringer, 1935, Phaenocarpa)

Distribution

England, Ireland

Aspilota

Förster, 1863

• DIPIESTA Förster, 1863

• EUSYNALDIS Zaykov & Fischer, 1982

• SYNALDIS misident.

Notes

Some distribution data from Stelfox and Graham (1951a). Some species treated as belonging to Dinotrema by other authors have been included in Aspilota in Taxapad (Yu et al. 2012) and there is clearly much work to be done in allocating species to the current generic concepts (see Wharton 1985). The late T. Munk was preparing a revision of the European species of Aspilotas. l. (see note under Dinotrema).

Aspilota acutidentata

(Fischer, 1970)

• Synaldis acutidentata Fischer, 1970

Distribution

England, Scotland

Notes

NMS, BMNH, det. Munk, added here; identified by T. Munk as a species of Dinotrema.

Aspilota anaphoretica

Fischer, 1973

Distribution

England, Scotland, Ireland

Notes

NMS, BMNH, det. Munk, added here

Aspilota blasii

Fischer, 1973

Distribution

England

Notes

BMNH, det. Munk, added here

Aspilota compressiventris

Stelfox & Graham, 1951

Distribution

England

Aspilota curta

Marshall, 1895

Distribution

England

Aspilota daemon

Stelfox and Graham, 1948

Distribution

England, Scotland, Ireland

Aspilota delicata

Fischer, 1973

Distribution

England, Scotland, Ireland

Notes

BMNH, det. Munk, added here

Aspilota efoveolata

Thomson, 1895

• pneumatica Fischer, 1973

Distribution

England, Scotland, Wales

Notes

NMS, BMNH, det. Munk, added here

Aspilota flagellaris

Fischer, 1973

Distribution

England, Scotland, Ireland

Notes

BMNH, det. Munk, added here

Aspilota fuscicornis

(Haliday, 1838)

• Alysia fuscicornis Haliday, 1838

• minuta (Nees, 1812, Bassus)

• exile (Ruthe, 1859, Orthostigma)

• dilatata (Thomson, 1895, Alysia)

Distribution

England, Scotland, Ireland

Aspilota globipes

(Fischer, 1962)

• Synaldis globipes Fischer, 1962

Distribution

England, Scotland

Notes

NMS, BMNH, det. Munk, added here

Aspilota imparidens

Fischer, 1974

Distribution

England, Scotland, Wales

Notes

NMS, BMNH, det. Munk, added here

Aspilota insolita

(Tobias, 1962)

• Orthostigma insolita Tobias, 1962

Distribution

England, Ireland

Notes

BMNH, det. Munk, added here

Aspilota intermediana

Fischer, 1975

Distribution

England

Notes

added by Notton (1991)

Aspilota iocosipecta

Fischer, 1974

Distribution

England, Scotland, Ireland

Notes

NMS, BMNH, det. Munk, added here

Aspilota macrops

Stelfox & Graham, 1951

Distribution

England, Ireland

Aspilota nidicola

Hedqvist, 1972

Distribution

England

Notes

BMNH, det. Munk, added here

Aspilota pillerensis

Fischer, 1973

Distribution

England

Notes

BMNH, det. Munk, added here

Aspilota ruficornis

(Nees, 1834)

• Alysia ruficornis Nees, 1834

Distribution

England, Scotland, Wales, Ireland

Aspilota stenogaster

Stelfox & Graham, 1951

Distribution

England

Aspilota tetragona

Fischer, 1976

Distribution

England, Scotland, Ireland

Notes

NMS, BMNH, det. Munk, added here

Aspilota vernalis

Stelfox & Graham, 1951

Distribution

England, Scotland, Ireland

Chasmodon

Haliday, 1838

Chasmodon apterus

(Nees, 1812)

• Bassus apterus Nees, 1812

Distribution

England, Scotland, Wales, Ireland, Isle of Man

Cratospila

Förster, 1863

• HEDYLUS Marshall, 1891; synonymy by Papp (2009b)

Cratospila circe

(Haliday, 1838)

• Alysia circe Haliday, 1838

• habilis (Marshall, 1891, Hedylus); synonymy by Papp (2009b)

• annellata (Thomson, 1895, Alysia)

Distribution

England, Ireland

Dapsilarthra

Förster, 1863

Notes

Dapsilarthra has been used in a very broad sense to include Adelurola, Mesocrina, Heterolexis and Grammospila, and in a more narrow sense to include the last two genera. Achterberg (2014) is followed here in according each generic rank.

Dapsilarthra apii

(Curtis, 1826)

• Alysia apii Curtis, 1826

• laevipectus (Thomson, 1895, Alysia)

• americana (Brues, 1907, Orthostigma)

Distribution

England, Scotland, Ireland

Dapsilarthra sylvia

(Haliday, 1839)

• Alysia sylvia Haliday, 1839

• carpathica van Achterberg, 1983; synonymy by Achterberg (1997)

Distribution

England, Scotland, Ireland

Dinotrema

Förster, 1863

Notes

Wharton (1980) summarises the arguments against recognition of the genus Synaldis; Achterberg (1988b) likewise notes that the genus is defined only by the absence of fore wing vein 2-SR, which is known to be intraspecifically variable (Wharton 1980). Despite this, various authors attach great taxonomic weight to this venational character and maintain Synaldis as a valid genus (as reflected by the classification in Yu et al. 2012). Species that would be classified in Synaldis (acutidentata, concolor, distracta and globipes) are listed here in Aspilota and Dinotrema, according to Fauna Europaea. Some distribution data from Stelfox and Graham (1951b), Stelfox and Graham (1951a) and Achterberg (1988a). The late T. Munk was revising the European species of Aspilotas.l. and gave much helpful advice on the generic placements of the species occurring in Britain and Ireland, some of which is unpublished and will therefore differ from the generic combinations found in, e.g. Belokobylskij et al. (2003) and Taxapad (Yu et al. 2012). Munk also advised that although Aspilota is a well-defined genus, Dinotrema is not defined by any apomorphies and will be split up.

Species excluded from the British and Irish list:

[pusillum (Nees, 1812, Bassus)] T. Munk (pers. comm.) regarded this as an unidentified species; the type has been destroyed.

Dinotrema

Förster, 1863

• ASPILOTA misident.

• SYNALDIS Förster, 1863

• COLOBOMA Förster, 1863

• SPANOMERIS Förster, 1863

• SCOTIONEURUS Provancher, 1886

• PTERUSA Fischer, 1958; synonymy by Achterberg and Vikberg (2014)

• EUDINOSTIGMA Tobias, 1986; synonymy by Wharton (2002)

Dinotrema (Dinotrema) aluum

(Stelfox & Graham, 1950)

• Aspilota alua Stelfox & Graham, 1950

• alva misspelling

Distribution

Ireland

Dinotrema (Dinotrema) alysiae

Munk & Peris-Felipo, 2013

Distribution

England

Notes

added by Munk et al. (2013)

Dinotrema (Dinotrema) areolatum

(Stelfox & Graham, 1950)

• Aspilota areolata Stelfox & Graham, 1950

Distribution

England, Scotland

Dinotrema (Dinotrema) brevicorne

(Nees, 1814)

• Bassus brevicornis Nees, 1814

Distribution

England

Dinotrema (Dinotrema) brevissimicorne

(Stelfox & Graham, 1948)

• Aspilota brevissimicornis Stelfox & Graham, 1948

Distribution

Scotland, Ireland

Dinotrema (Dinotrema) compressum

(Haliday, 1838)

• Alysia compressa Haliday, 1838

Distribution

England

Dinotrema (Dinotrema) concinnum

(Haliday, 1838)

• Alysia concinna Haliday, 1838

• maximum (Fischer, 1962, Synaldis)

• tyrrhena (Masi, 1933, Aspilota)

Distribution

England, Ireland

Notes

Synaldis maximum Fischer, 1962 has been treated as a junior synonym by König (1972) and subsequent authors but T. Munk (pers. comm.) regarded this as a valid species of Dinotrema that may be found to occur in Britain.

Dinotrema (Dinotrema) concolor

(Nees, 1812)

• Bassus concolor Nees, 1812

• distractum (Ruthe, 1859, Orthostigma) unavailable

Distribution

England, Ireland

Dinotrema (Dinotrema) crassicosta

(Thomson, 1895)

• Alysia crassicosta Thomson, 1895

Distribution

Scotland

Notes

NMS, det. Munk, added here

Dinotrema (Dinotrema) cratocera

(Thomson, 1895)

• Alysia cratocera Thomson, 1895

Distribution

Scotland

Notes

BMNH, det. Munk, added here

Dinotrema (Dinotrema) denticulatum

(Stelfox & Graham, 1951)

• Aspilota denticulata Stelfox & Graham, 1951

Distribution

Ireland

Dinotrema (Dinotrema) dimidiatum

(Thomson, 1895)

• Alysia dimidiata Thomson, 1895

Distribution

England

Notes

BMNH, det. Munk, added here

Dinotrema (Dinotrema) distractum

(Nees, 1834)

• Alysia distracta Nees, 1834

Dinotrema (Dinotrema) divisum

(Stelfox & Graham, 1950)

• Aspilota divisa Stelfox & Graham, 1950

• aureliae (Fischer, 1973, Aspilota); tentative synonymy by Papp (2004b)

Distribution

England, Scotland, Ireland

Dinotrema (Dinotrema) erythropum

Förster, 1863

• praecipuum (Marshall, 1895, Aspilota)

Distribution

England, Wales, Ireland

Dinotrema (Dinotrema) falsificum

(Stelfox & Graham, 1950)

• Aspilota falsifica Stelfox & Graham, 1950

Distribution

Scotland, Ireland

Dinotrema (Dinotrema) glabrum

(Stelfox & Graham, 1951)

• Aspilota glabra Stelfox & Graham, 1951

• venustum (Tobias, 1962, Aspilota)

Distribution

England, Scotland, Ireland

Dinotrema (Dinotrema) insidiatrix

(Marshall, 1895)

• Aspilota insidiatrix Marshall, 1895

Distribution

England

Dinotrema (Dinotrema) insignis

(Stelfox & Graham, 1950)

• Aspilota insignis Stelfox & Graham, 1950

Distribution

England

Dinotrema (Dinotrema) iuxtanaeviam

(Fischer, 1980)

• Aspilota iuxtanaeviam Fischer, 1978

Distribution

England

Notes

added by Fischer (1980)

Dinotrema (Dinotrema) jaculans

(Haliday, 1838)

• Alysia jaculans Haliday, 1838

Distribution

England, Scotland, Ireland

Dinotrema (Dinotrema) latistigma

(Fischer, 1962)

• Synaldis latistigma Fischer, 1962

Distribution

England

Notes

NMS, det. Munk, added here; treated as a species of Eudinostigma in Fauna Europaea and Taxapad (Yu et al. 2012).

Dinotrema (Dinotrema) lineola

(Thomson, 1895)

• Alysia lineola Thomson, 1895

Distribution

England

Dinotrema (Dinotrema) liosoma

(Stelfox & Graham,1951)

• Aspilota liosoma Stelfox & Graham,1951

• caudatum (Thomson, 1895, Aspilota) preocc.

Dinotrema (Dinotrema) mesocaudatum

van Achterberg, 1988

Distribution

England

Notes

added by Achterberg (1988b)

Dinotrema (Dinotrema) microcera

(Thomson, 1895)

• Alysia microcera Thomson, 1895

Distribution

England, Scotland, Wales, Ireland

Notes

BMNH, det. Munk, added here

Dinotrema (Dinotrema) necrophilum

(Hedqvist, 1972)

• Aspilota necrophila Hedqvist, 1972

Distribution

England

Notes

Added by Disney and Munk (2005) and transferred from Aspilota.

Dinotrema (Dinotrema) nervosum

(Haliday, 1833)

• Alysia nervosa Haliday, 1833

Distribution

England, Scotland, Ireland

Dinotrema (Dinotrema) pulvinatum

(Stelfox & Graham, 1949)

• Aspilota pulvinata Stelfox & Graham, 1949

Distribution

England, Scotland, Wales, Ireland

Notes

Treated as a species of Eudinostigma in Fauna Europaea and Taxapad (Yu et al. 2012).

Dinotrema (Dinotrema) ruficollis

(Stelfox & Graham, 1950)

• Aspilota ruficollis Stelfox & Graham, 1950

Distribution

England, Ireland

Notes

Distribution data from Achterberg (1988b).

Dinotrema (Dinotrema) semicompressum

(Stelfox & Graham, 1949)

• Aspilota semicompressa Stelfox & Graham, 1949

• parapunctatum (Fischer, 1976, Aspilota); synonymy by Papp (2004b)

Distribution

England, Scotland, Ireland

Dinotrema (Dinotrema) sphaerimembre

(Fischer, 1973)

• Aspilota sphaerimembris Fischer, 1973

Distribution

Scotland

Notes

NMS, det. Munk, added here

Dinotrema (Dinotrema) sternaulicum

(Fischer, 1973)

• Aspilota sternaulica Fischer, 1973

Distribution

England, Scotland

Notes

NMS, det. Munk, added here

Dinotrema (Dinotrema) tauricum

(Telenga, 1935)

• Aspilota taurica Telenga, 1935

Distribution

England

Notes

added by Achterberg (1988b)

Dinotrema (Dinotrema) vesparum

(Stelfox, 1943)

• Aspilota vesparum Stelfox, 1943

Distribution

England, Scotland

Dinotrema (Dinotrema) vituperatum

(Fischer, 1974)

• Aspilota vituperata Fischer, 1974

Distribution

England, Scotland

Notes

NMS, det. Munk, added here

Leptotrema

van Achterberg, 1988

Notes

Described as a separate genus by Achterberg (1988b), who gives distribution data for the one included species; treated as a subgenus of Dinotrema by Wharton (2002).

Dinotrema (Leptotrema) dentifemur

(Stelfox, 1943)

• Aspilota dentifemur Stelfox, 1943

Distribution

England, Scotland, Ireland

Prosapha

Förster, 1863

Dinotrema (Prosapha) speculum

(Haliday, 1838)

• Alysia speculum Haliday, 1838

• venustum (Haliday, 1838, Alysia)

Distribution

England, Ireland

Grammospila

Förster, 1863

• PARAORTHOSTIGMA Königsmann, 1972

Grammospila isabella

(Haliday, 1838)

• Alysia isabella Haliday, 1838

Distribution

England, Scotland

Grammospila rufiventris

(Nees, 1812)

• Bassus rufiventris Nees, 1812

• flaviventris (Haliday, 1838, Alysia)

• ochrogaster (Szépligeti, 1898, Phaenocarpa)

• fuscula Griffiths, 1968

Distribution

England, Scotland, Ireland, Isle of Man

Notes

Grammospila fuscula (Griffiths, 1968, Dapsilarthra) removed from synonymy by Achterberg (2014)​.

Heterolexis

Förster, 1863

Heterolexis balteata

(Thomson, 1895)

• Alysia balteata Thomson, 1895

Distribution

England, Scotland, Wales, Ireland

Heterolexis dictynna

(Marshall, 1895)

• Adelura dictynna Marshall, 1895

Distribution

England, Scotland, Ireland

Idiasta

Förster, 1863

• EUPHAENOCARPA Tobias, 1975

Idiasta dichrocera

Königsmann, 1960

Distribution

England

Notes

NMS, det. Shaw, added here

Idiasta maritima

(Haliday, 1838)

• Alysia maritima Haliday, 1838

Distribution

England, Scotland, Ireland

Idiasta nephele

(Haliday, 1838)

• Alysia nephele Haliday, 1838

Distribution

England, Scotland

Mesocrina

Förster, 1863

• PSEUDOMESOCRINA Königsmann, 1959

Mesocrina indagatrix

Förster, 1863

• venatrix Marshall, 1895

Distribution

England

Orthostigma

Ratzeburg, 1844

• DELOCARPA Förster, 1863

• ISCHNOCARPA Förster, 1863

Orthostigma cratospilum

(Thompson, 1895)

• Alysia cratospila Thompson, 1895

Distribution

England

Notes

added by Godfray and Achterberg (2015)

Orthostigma longicorne

Königsmann, 1969

Distribution

Ireland

Orthostigma maculipes

(Haliday, 1838)

• Alysia maculipes Haliday, 1838

Distribution

England, Ireland

Orthostigma pumilum

(Nees, 1834)

• Alysia pumila Nees, 1834

• flavipes (Ratzeburg, 1844, Aphidius)

• fulvipes Rondani, 1876 nom. nud.

• brunnipes Ratzeburg, 1852

• bruneipes Dalla Torre, 1898

Distribution

England, Ireland

Panerema

Förster, 1863

Notes

Distribution data from Achterberg (1988b).

Panerema fulvicornis

(Haliday, 1838)

• Alysia fulvicornis Haliday, 1838

Distribution

Wales, Ireland

Panerema inops

Förster, 1863

Distribution

England, Wales, Ireland

Pentapleura

Förster, 1863

• OPISENDEA Förster, 1863

• GNATHOSPILA Fischer, 1966

Pentapleura angustula

(Haliday, 1838)

• Alysia angustula Haliday, 1838

• laevipleuris (Tobias, 1962, Aspilota)

Distribution

England, Scotland, Ireland

Pentapleura fuliginosa

(Haliday, 1838)

• Alysia fuliginosa Haliday, 1838

• carinata (Thomson, 1895, Alysia)

Distribution

England, Wales, Ireland

Pentapleura pumilio

(Nees, 1812)

• Bassus pumilio Nees, 1812

• triticaphis (Fitch, 1861, Toxares)

• mesocrinoides Goidanich, 1936

Distribution

England, Scotland, Wales, Ireland

Phaenocarpa

Förster, 1863

Notes

Some taxonomic and distribution data from Achterberg (1997).

Homophyla

Förster, 1863

Notes

species of Phaenocarpa (Homophyla) excluded from the British and Irish list:

[pegomyiae Marshall, 1898] Listed as British by Lyle (1933) but we have not seen any British or Irish specimens.

Phaenocarpa (Homophyla) pullata

(Haliday, 1838)

• Alysia pullata Haliday, 1838

Distribution

England, Ireland

Phaenocarpa

Förster, 1863

• IDIOLEXIS Förster, 1863; synonymy by Wharton (2002)

• MESOTHESIS Förster, 1863

• SATHRA Förster, 1863

• ASYNAPHES Provancher, 1886

• KAHLIA Ashmead, 1900

• HOLCALYSIA Cameron, 1905

• STIRALYSIA Cameron, 1910

• RHOPALONEURA Stelfox, 1941

Phaenocarpa (Phaenocarpa) canaliculata

Stelfox, 1941

Distribution

England, Ireland

Phaenocarpa (Phaenocarpa) conspurcator

(Haliday, 1838)

• Alysia conspurcator Haliday, 1838

• arctica (Thomson, 1895, Alysia)

• tatrica Niezabitowski, 1910

• remota Papp, 1981

Distribution

England, Scotland, Wales, Ireland

Phaenocarpa (Phaenocarpa) eugenia

(Haliday, 1838)

• Alysia eugenia Haliday, 1838

• pectoralis (Zetterstedt, 1838, Alysia)

• orbicularis Gurasashvilli, 1983

Distribution

England, Scotland, Ireland

Phaenocarpa (Phaenocarpa) eunice

(Haliday, 1838)

• Alysia eunice Haliday, 1838

• nimia Stelfox, 1941

Distribution

England, Wales, Ireland

Phaenocarpa (Phaenocarpa) flavipes

(Haliday, 1838)

• Alysia flavipes Haliday, 1838

Distribution

England, Ireland

Phaenocarpa (Phaenocarpa) frequentator

(Zetterstedt, 1838)

• Alysia frequentator Zetterstedt, 1838

• frequentatrix (Schulz, 1906, Alysia)

Distribution

Wales

Notes

added by Godfray and Achterberg (2015)

Phaenocarpa (Phaenocarpa) galatea

(Haliday, 1838)

• Alysia galatea Haliday, 1838

Distribution

Scotland, Ireland

Phaenocarpa (Phaenocarpa) helophilae

van Achterberg, 1998

Distribution

England

Notes

added by Achterberg (1998)

Phaenocarpa (Phaenocarpa) livida

(Haliday, 1838)

• Alysia livida Haliday, 1838

• debilis (Förster, 1863, Sathra)

Distribution

England, Ireland

Phaenocarpa (Phaenocarpa) luteipes

Stelfox, 1950

Distribution

Ireland

Phaenocarpa (Phaenocarpa) maria

(Haliday, 1838)

• Alysia maria Haliday, 1838

Distribution

England, Ireland

Phaenocarpa (Phaenocarpa) nina

(Haliday, 1838)

• Alysia nina Haliday, 1838

Distribution

Scotland

Notes

Not listed in Huddleston (1978), although described from Scottish material (Achterberg 1998).

Phaenocarpa (Phaenocarpa) notabilis

Stelfox, 1944

Distribution

Scotland, Ireland

Phaenocarpa (Phaenocarpa) picinervis

(Haliday, 1838)

• Alysia picinervis Haliday, 1838

• americana Ashmead, 1889

Distribution

England, Scotland, Ireland

Phaenocarpa (Phaenocarpa) pratellae

(Curtis, 1826)

• Alysia pratellae Curtis, 1826

• piceator (Zetterstedt, 1838, Alysia)

• psalliotae Telenga, 1935; synonymy by Achterberg (2014)

Distribution

England, Scotland

Phaenocarpa (Phaenocarpa) punctigera

(Haliday, 1838)

• Alysia punctigera Haliday, 1838

Distribution

England, Ireland

Notes

Listed as a species of Idiolexis in Fauna Europaea.

Phaenocarpa (Phaenocarpa) ruficeps

(Nees, 1812)

• Bassus ruficeps Nees, 1812

• testacea (Nees, 1812, Bassus)

• gracilis (Curtis, 1826, Alysia)

• pallida (Curtis, 1826, Alysia)

• agricolator (Zetterstedt, 1838, Alysia)

• oculator (Ratzeburg, 1848, Alysia)

• rubriceps (Provancher, 1883, Alysia) preocc.

• rubricepes Provancher, 1888

• testaceipes (Cameron, 1905, Holcalysia)

• divergens Fischer, 1975

• fervida Fischer, 1975

• incerta Fischer, 1975

• meritoria Papp, 1981

• ferga Papp, 1982

Distribution

England, Scotland, Wales, Ireland, Isle of Man

Phaenocarpa (Phaenocarpa) tacita

Stelfox, 1941

• caucasica Gurasashvili, 1983 preocc.

• caucasicola Tobias, 1986

Distribution

Ireland

Phaenocarpa (Phaenocarpa) trisulcata

Stelfox, 1950

Distribution

Ireland

Syncrasis

Förster, 1863

• PHAENOLYTA Förster, 1863

Syncrasis fucicola

(Haliday, 1838)

• Alysia fucicola Haliday, 1838

Distribution

Ireland

Syncrasis halidayi

(Förster, 1863)

• Phaenolyta halidayi Förster, 1863

• fuscipes preocc., unavailable

Distribution

England, Ireland

Tanycarpa

Förster, 1863

• ACROBELA Förster, 1863; synonymy by Wharton (2002)

• EPICLISTA Förster, 1863

• HYPOSTROPHA Förster, 1863

Tanycarpa bicolor

(Nees, 1812)

• Bassus bicolor Nees, 1812

• ancilla (Haliday, 1838, Alysia)

Distribution

England, Ireland

Tanycarpa gracilicornis

(Nees, 1812)

• Bassus gracilicornis Nees, 1812

Distribution

Ireland

Tanycarpa mitis

Stelfox, 1941

Distribution

England, Ireland

Tanycarpa punctata

van Achterberg, 1976

Tanycarpa rufinotata

(Haliday, 1838)

• Alysia rufinotata Haliday, 1838

• carinata (Förster, 1863, Acrobela)

• erythrogaster (Förster, 1863, Epiclista)

• foersteri (Shenefelt, 1974, Alysia)

Distribution

England, Ireland

Trachyusa

Ruthe, 1854

• COSMIOCARPA Förster, 1863

Notes

Distribution data from Achterberg and O'Connor (1990) and the collections of NMS.

Trachyusa aurora

(Haliday, 1838)

• Alysia aurora Haliday, 1838

• nigriceps Ruthe, 1854

Distribution

England, Ireland

Trachyusa nigrothoracica

van Achterberg & O’Connor, 1990

Distribution

England, Ireland

Notes

added by Achterberg and O'Connor (1990)

Dacnusini

Förster, 1863

Notes

The classification of the Dacnusini was revised by Griffiths (Griffiths 1964, Griffiths 1967a, Griffiths 1967b, Griffiths 1967c, Griffiths 1968a, Griffiths 1968b) in a pioneering treatment that was one of the first applications of explicit phylogenetic methods in taxonomy. It was also unusual in being based on extensive reared material. Griffiths’ classification is followed here although one of the consequences of his strict adherence to phylogenetic principles is that Chorebus and Dacnusa are very large genera. Also, as Griffiths realised, the definition of Exotela is unsatisfactory as it lacks clear apomorphic characters. This group of insects had previously been revised by Nixon (Nixon 1943, Nixon 1944, Nixon 1945, Nixon 1946, Nixon 1948, Nixon 1949, Nixon 1954), who recognised a greater number of genera, and some authors (Tobias 1986, Tobias 1998, Perepechayenko 2000, Fischer et al. 2004) have resurrected Nixon’s genera or used them as subgenera. This seems a retrograde step given Griffiths’ clear statements of phylogenetic hypotheses and is not used here. In both Chorebus and Dacnusa, Griffiths defined a series of species groups that with further research should form the basis of better classifications of these genera. Griffiths (1964) gives a key to genera and Wharton (1997) is also very helpful. The Coelinius genus group (Coelinius, Trachionus, Epimicta, Aristelix, Laotris, Sarops and Synelix) is generally agreed to be monophyletic and was not revised at species level by Nixon or Griffiths. Distribution data from Griffiths’ and Nixon’s revisions (loc. cit.) and NMS.

Amyras

Nixon, 1943

Amyras clandestina

(Haliday, 1839)

• Alysia clandestina Haliday, 1839

• quadridentata (Thomson, 1895, Dacnusa)

Distribution

Ireland

Aristelix

Nixon, 1943

Aristelix phaenicura

(Haliday, 1839)

• Alysia phaenicura Haliday, 1839

• phoenicura misspelling

Distribution

England, Ireland

Chaenusa

Haliday, 1839

• CHOREBIDEA Viereck, 1914

• CHOREBIDEA Nixon, 1943 preocc.

• CHOREBIDELLA Riegel, 1947

Notes

No recent revision of this genus.

Chaenusa conjungens

(Nees, 1811)

• Bracon conjungens Nees, 1811

• conjugens misspelling

Distribution

England, Scotland, Ireland

Chaenusa elongata

Stelfox, 1957

Distribution

Ireland

Chaenusa limoniadum

(Marshall, 1896)

• Chorebus limoniadum Marshall, 1896

Distribution

England

Notes

Transferred from Chorebus by Perepechayenko (2000).

Chaenusa lymphata

(Haliday, 1839)

• Alysia lymphata Haliday, 1839

Distribution

England, Ireland

Notes

Transferred from Chorebus by Achterberg (1997).

Chaenusa naiadum

(Haliday, 1839)

• Alysia naiadum Haliday, 1839

• naiadum (Curtis, 1837, Chorebus) nom. nud.

Distribution

England, Scotland, Ireland

Chaenusa nereidum

(Haliday, 1839)

• Alysia nereidum Haliday, 1839

• nereidum (Curtis, 1837, Chorebus) nom. nud.

Distribution

England, Ireland

Chaenusa opaca

Stelfox, 1957

Distribution

Ireland

Chorebus

Haliday, 1833

• AMETRIA Förster, 1863

• GYROCAMPA Förster, 1863

• PHAENOLEXIS Förster, 1863

• STIPHROCERA Förster, 1863

• DIPLUSIA Ruthe, 1882

• ETRIPTES Nixon, 1943

• PARAGYROCAMPA Tobias, 1962

Chorebus abaris

(Nixon, 1943)

• Dacnusa abaris Nixon, 1943

Distribution

England, Scotland

Chorebus abnormiceps

(Nixon, 1943)

• Dacnusa abnormiceps Nixon, 1943

• quadriceps (Nixon, 1941, Dacnusa) preocc.

Distribution

England

Chorebus agraules

(Nixon, 1945)

• Dacnusa agraules Nixon, 1945

Distribution

England

Chorebus albipes

(Haliday, 1839)

• Alysia albipes Haliday, 1839

Distribution

England, Scotland, Ireland

Chorebus alecto

(Morley, 1924)

• Rhizarcha alecto Morley, 1924

• turissa (Nixon, 1937, Dacnusa)

Distribution

England, Scotland, Ireland

Chorebus alua

(Nixon, 1944)

• Dacnusa alua Nixon, 1944

Distribution

Ireland

Chorebus amasis

(Nixon, 1945)

• Dacnusa amasis Nixon, 1945

Distribution

England

Chorebus ampliator

(Nees, 1834)

• Alysia ampliator Nees, 1834

• nigricornis (Förster, 1863, Stiphrocera)

Distribution

England, Scotland, Ireland

Chorebus anasellus

(Stelfox, 1952)

• Dacnusa anasella Stelfox, 1952

Distribution

Scotland, Ireland

Chorebus angelicae

(Nixon, 1945)

• Dacnusa angelicae Nixon, 1945

Distribution

England

Chorebus anita

(Nixon, 1943)

• Dacnusa anita Nixon, 1943

Distribution

England, Scotland

Chorebus aphantus

(Marshall, 1895)

• Dacnusa aphanta Marshall, 1895

Distribution

England, Scotland, Wales, Ireland

Chorebus apollyon

(Morley, 1924)

• Dacnusa apollyon Morley, 1924

Distribution

England, Ireland

Chorebus ares

(Nixon, 1944)

• Dacnusa ares Nixon, 1944

Distribution

England

Chorebus armida

(Nixon, 1945)

• Dacnusa armida Nixon, 1945

Distribution

England, Scotland, Wales

Chorebus artemisiellus

Griffiths, 1968

Distribution

England

Chorebus asramenes

(Nixon, 1943)

• Dacnusa asramenes Nixon, 1943

Distribution

England, Ireland

Chorebus avesta

(Nixon, 1944)

• Dacnusa avesta Nixon, 1944

Distribution

England, Scotland

Chorebus bathyzonus

(Marshall, 1895)

• Dacnusa bathyzona Marshall, 1895

• ornatus (Telenga, 1935, Dacnusa)

Distribution

England, Ireland

Chorebus bensoni

(Nixon, 1943)

• Dacnusa bensoni Nixon, 1943

Distribution

England, Scotland

Chorebus bres

(Nixon, 1944)

• Dacnusa bres Nixon, 1944

Distribution

England, Ireland

Chorebus brevicornis

(Thomson, 1895)

• Dacnusa brevicornis Thomson, 1895

• chrysippe (Nixon, 1944, Dacnusa)

• ea (Nixon, 1944, Dacnusa)

Distribution

England, Scotland, Ireland

Chorebus caelebs

(Nixon, 1944)

• Dacnusa caelebs Nixon, 1944

Distribution

England

Chorebus calthae

Griffiths, 1967

Distribution

England

Chorebus cambricus

Griffiths, 1968

Distribution

Wales

Chorebus cinctus

(Haliday, 1839)

• Alysia cincta Haliday, 1839

• castaneiventris (Thomson, 1895, Dacnusa)

Distribution

England, Ireland

Chorebus coxator

(Thomson, 1895)

• Dacnusa coxator Thomson, 1895

Distribution

England, Scotland, Wales

Chorebus crassipes

(Stelfox, 1954)

• Dacnusa crassipes Stelfox, 1954

Distribution

Scotland, Ireland

Chorebus credne

(Nixon, 1944)

• Dacnusa credne Nixon, 1944

Distribution

England, Scotland, Ireland

Chorebus crenulatus

(Thomson, 1895)

• Dacnusa crenulata Thomson, 1895

• elegantulus (Nixon, 1937, Dacnusa)

Distribution

England, Scotland, Ireland

Chorebus crocale

(Nixon, 1945)

• Dacnusa crocale Nixon, 1945

Distribution

England

Chorebus cubocephalus

(Telenga, 1934)

• Rhizarcha cubocephala Telenga, 1934

• cyclops (Nixon, 1937, Dacnusa); synonymy by Tobias (1986)

Distribution

Ireland

Chorebus cylindricus

(Telenga, 1934)

• Dacnusa cylindrica Telenga, 1934

• cybele (Nixon, 1937, Dacnusa); synonymy by Tobias (1986)

Distribution

England, Scotland

Chorebus cytherea

(Nixon, 1937)

• Dacnusa cytherea Nixon, 1937

• calliope (Nixon, 1944, Dacnusa)

• tesmia (Nixon, 1944, Dacnusa)

Distribution

England, Scotland, Ireland

Chorebus dagda

(Nixon, 1943)

• Dacnusa dagda Nixon, 1943

Distribution

England

Chorebus daimenes

(Nixon, 1945)

• Dacnusa daimenes Nixon, 1945

Distribution

England, Scotland

Chorebus deione

(Nixon, 1944)

• Dacnusa deione Nixon, 1944

Distribution

England, Scotland

Chorebus didas

(Nixon, 1944)

• Dacnusa didas Nixon, 1944

Distribution

England, Scotland

Chorebus difficilis

Griffiths, 1968

Distribution

England

Chorebus diremtus

(Nees, 1834)

• Alysia diremta Nees, 1834

• diremptus (Haliday, 1839, Alysia)

Distribution

England, Ireland

Chorebus dirona

(Nixon, 1945)

• Dacnusa dirona Nixon, 1945

Distribution

England, Ireland

Chorebus enephes

(Nixon, 1945)

• Dacnusa enephes Nixon, 1945

Distribution

England

Chorebus eros

(Nixon, 1937)

• Dacnusa eros Nixon, 1937

Distribution

England, Scotland, Ireland

Chorebus esbelta

(Nixon, 1937)

• Dacnusa esbelta Nixon, 1937

Distribution

England, Scotland, Ireland

Chorebus euryale

(Nixon, 1944)

• Dacnusa euryale Nixon, 1944

Distribution

England

Chorebus fallaciosae

Griffiths, 1967

Distribution

England

Chorebus fallax

(Nixon, 1937)

• Dacnusa fallax Nixon, 1937

Distribution

England, Scotland, Ireland

Chorebus flavipes

(Goureau, 1851)

• Dacnusa flavipes Goureau, 1851

• raissa (Nixon, 1937, Dacnusa)

Distribution

England, Scotland, Ireland

Chorebus fordi

(Nixon, 1954)

• Dacnusa fordi Nixon, 1954

Distribution

England

Chorebus foveolus

(Haliday, 1839)

• Alysia foveola Haliday, 1839

Distribution

England, Scotland, Wales, Ireland

Chorebus fuscipennis

(Nixon, 1937)

• Dacnusa fuscipennis Nixon, 1937

Distribution

England

Chorebus gedanensis

(Ratzeburg, 1852)

• Alysia gedanensis Ratzeburg, 1852

• anguligena (Nixon, 1937, Dacnusa)

Distribution

England

Chorebus glaber

(Nixon, 1944)

• Dacnusa glabra Nixon, 1944

Distribution

England, Ireland

Chorebus glabriculus

(Thomson, 1895)

• Dacnusa glabricula Thomson, 1895

• cortipalpis (Nixon, 1937, Dacnusa)

Distribution

England, Scotland, Ireland

Chorebus hilaris

Griffiths, 1967

Distribution

England

Chorebus hirtigena

Stelfox, 1957

Distribution

Ireland

Chorebus humeralis

Griffiths, 1968

Distribution

Ireland

Chorebus incertus

(Goureau, 1851)

• Dacnusa incerta Goureau, 1851

Distribution

England

Chorebus iphias

(Nixon, 1943)

• Dacnusa iphias Nixon, 1943

Distribution

England

Chorebus kama

(Nixon, 1945)

• Dacnusa kama Nixon, 1945

Distribution

England

Chorebus lanigerus

(Stelfox, 1957)

• Gyrocampa lanigera Stelfox, 1957

Distribution

Ireland

Chorebus lar

(Morley, 1924)

• Dacnusa lar Morley, 1924

• innanus (Nixon, 1943, Dacnusa)

Distribution

England

Chorebus larides

(Nixon, 1944)

• Dacnusa larides Nixon, 1944

Distribution

England, Ireland

Chorebus lateralis

(Haliday, 1839)

• Alysia lateralis Haliday, 1839

• fuscula (Haliday, 1839, Alysia)

• albicoxa (Thomson, 1895, Dacnusa)

Distribution

England, Scotland, Wales, Ireland

Chorebus leptogaster

(Haliday, 1839)

• Alysia leptogaster Haliday, 1839

• naenia (Morley, 1924, Dacnusa)

• dinae (Burghele, 1960, Dacnusa)

Distribution

England, Ireland

Chorebus longicornis

(Nees, 1811)

• Bracon longicornis Nees, 1811

• affinis (Nees, 1812, Bassus)

Distribution

England, Scotland, Ireland

Notes

Synonymy first established in Shenefelt (1974) but overlooked until O'Connor et al. (1999).

Chorebus lugubris

(Nixon, 1937)

• Dacnusa lugubris Nixon, 1937

Distribution

England, Ireland

Chorebus luzulae

Griffiths, 1966

Distribution

Scotland

Notes

added by Godfray and Bland (2011)

Chorebus lychnidis

Griffiths, 1967

Distribution

England

Chorebus maculigastra

Shenefelt, 1974

• maculata (Nixon, 1944, Dacnusa) preocc.

Distribution

England, Ireland

Chorebus merellus

(Nixon, 1937)

• Dacnusa merella Nixon, 1937

Distribution

England, Scotland, Ireland

Chorebus merion

(Nixon, 1945)

• Dacnusa merion Nixon, 1945

Distribution

England

Chorebus miodes

(Nixon, 1949)

• Gyrocampa miodes Nixon, 1949

Distribution

England, Ireland

Chorebus misellus

(Marshall, 1895)

• Dacnusa misella Marshall, 1895

Distribution

England

Chorebus mitrus

(Nixon, 1945)

• Dacnusa mitra Nixon, 1945

Distribution

England, Scotland, Ireland

Chorebus nanus

(Nixon, 1943)

• Dacnusa nana Nixon, 1943

Distribution

England

Chorebus navicularis

(Nees, 1812)

• Bassus navicularis Nees, 1812

Distribution

England

Chorebus nerissus

(Nixon, 1937)

• Dacnusa nerissa Nixon, 1937

Distribution

England

Chorebus nigriscaposus

(Nixon, 1949)

• Gyrocampa nigriscaposa Nixon, 1949

• propodealis (Nixon, 1949, Gyrocampa)

Distribution

Ireland

Chorebus ninella

(Nixon, 1945)

• Dacnusa ninella Nixon, 1945

Distribution

England, Scotland, Ireland

Chorebus nobilis

Griffiths, 1968

Distribution

England, Ireland

Chorebus nomia

(Nixon, 1937)

• Dacnusa nomia Nixon, 1937

Distribution

England, Ireland

Chorebus nydia

(Nixon, 1937)

• Dacnusa nydia Nixon, 1937

Distribution

England, Scotland, Ireland

Chorebus orbiculatae

Griffiths, 1967

Distribution

England, Ireland

Chorebus ovalis

(Marshall, 1896)

• Dacnusa ovalis Marshall, 1896

Distribution

England, Ireland

Chorebus parvungula

(Thomson, 1895)

• Dacnusa parvungula Thomson, 1895

• acco (Nixon, 1943, Dacnusa)

Distribution

England, Scotland, Ireland

Chorebus perkinsi

(Nixon, 1944)

• Dacnusa perkinsi Nixon, 1944

Distribution

England

Chorebus petiolatus

(Nees, 1834)

• Alysia petiolata Nees, 1834

Distribution

England, Scotland, Ireland

Chorebus phaedra

(Nixon, 1937)

• Dacnusa phaedra Nixon, 1937

Distribution

England, Scotland

Chorebus pimpinellae

Griffiths, 1967

Distribution

England

Chorebus pione

(Nixon, 1944)

• Dacnusa pione Nixon, 1944

Distribution

England

Chorebus poemyzae

Griffiths, 1968

Distribution

England, Scotland

Chorebus posticus

(Haliday, 1839)

• Alysia postica Haliday, 1839

• gracilis (Nees, 1834, Alysia); synonymy by Achterberg (1997)

• egregia (Marshall, 1895, Dacnusa)

• dentatus (Tobias, 1962, Dacnusa); synonymy by Tobias (1986)

Distribution

England, Scotland, Ireland

Chorebus pulverosus

(Haliday, 1839)

• Alysia pulverosa Haliday, 1839

• marsyas (Nixon, 1937, Dacnusa)

Distribution

England, Ireland

Chorebus punctum

(Goureau, 1851)

• Dacnusa punctum Goureau, 1851

Distribution

England, Scotland, Ireland

Chorebus resa

(Nixon, 1937)

• Dacnusa resa Nixon, 1937

Distribution

England

Chorebus rhanis

(Nixon, 1943)

• Dacnusa rhanis Nixon, 1943

Distribution

Scotland

Chorebus risilis

(Nixon, 1949)

• Gyrocampa risilis Nixon, 1949

Distribution

Ireland

Chorebus rondanii

(Giard, 1904)

• Dacnusa rondanii Giard, 1904

• galbus (Nixon, 1944, Dacnusa)

Distribution

England

Chorebus rotundiventris

(Thomson, 1895)

• Dacnusa rotundiventris Thomson, 1895

Distribution

Ireland

Chorebus rousseaui

(Schulz, 1907)

• Dacnusa rousseaui Schulz, 1907

Notes

The status of this taxon needs further research.

Chorebus ruficollis

(Stelfox, 1957)

• Gyrocampa ruficollis Stelfox, 1957

Distribution

Ireland

Chorebus rufimarginatus

(Stelfox, 1954)

• Dacnusa rufimarginata Stelfox, 1954

Distribution

Ireland

Chorebus scabiosae

Griffiths, 1967

Distribution

England

Chorebus scabrifossa

Stelfox, 1957

Distribution

Ireland

Chorebus selene

(Nixon, 1937)

• Dacnusa selene Nixon, 1937

Distribution

England, Scotland

Chorebus senilis

(Nees, 1812)

• Bassus senilis Nees, 1812

• tomentosus (Thomson, 1895, Dacnusa)

• nemesis (Morley, 1924, Dacnusa)

Distribution

England, Scotland, Wales, Ireland

Chorebus serus

(Nixon, 1937)

• Dacnusa sera Nixon, 1937

Distribution

England

Chorebus siniffa

(Nixon, 1937)

• Dacnusa siniffa Nixon, 1937

Distribution

England, Ireland

Chorebus solstitialis

(Stelfox, 1952)

• Dacnusa solstitialis Stelfox, 1952

Distribution

England

Chorebus spenceri

Griffiths, 1964

Distribution

England, Scotland

Chorebus striola

Stelfox, 1957

Distribution

Ireland

Chorebus sylvestris

Griffiths, 1967

Distribution

England, Scotland, Ireland

Chorebus talaris

(Haliday, 1839)

• Alysia talaris Haliday, 1839

Distribution

England, Scotland, Ireland

Chorebus tamiris

(Nixon, 1943)

• Dacnusa tamiris Nixon, 1943

Distribution

England

Chorebus tamsi

(Nixon, 1944)

• Dacnusa tamsi Nixon, 1944

Distribution

England

Chorebus tanis

(Nixon, 1945)

• Dacnusa tanis Nixon, 1945

Distribution

England

Chorebus tenellae

Griffiths, 1967

Distribution

Scotland, Ireland

Notes

added by Griffiths (1984)

Chorebus thecla

(Nixon, 1943)

• Dacnusa thecla Nixon, 1943

Distribution

England

Chorebus thisbe

(Nixon, 1937)

• Dacnusa thisbe Nixon, 1937

Distribution

England

Chorebus thusa

(Nixon, 1937)

• Dacnusa thusa Nixon, 1937

Distribution

England, Ireland

Chorebus transversus

(Nixon, 1954)

• Dacnusa transversa Nixon, 1954

Distribution

England, Scotland, Ireland

Chorebus trilobomyzae

Griffiths, 1968

Distribution

England, Scotland

Chorebus uliginosus

(Haliday, 1839)

• Alysia uliginosa Haliday, 1839

• thienemanni (Ruschka, 1913, Gyrocampa)

Distribution

England, Ireland

Chorebus uma

(Nixon, 1944)

• Dacnusa uma Nixon, 1944

Distribution

England

Chorebus varuna

(Nixon, 1945)

• Dacnusa varuna Nixon, 1945

Distribution

England, Scotland

Chorebus vernalis

Griffiths, 1968

Distribution

England

Chorebus vitripennis

Griffiths, 1968

Distribution

Scotland, Wales, Ireland

Coelinidea

Viereck, 1913

• LEPTON Zetterstedt, 1838 preocc.

• ERICOELINIUS Viereck, 1913

• FISCHERASTRIOLUS Perepechayenko, 1999

Notes

There is no recent treatment of this genus; generic synonymy follows Achterberg (2014).

Coelinidea elegans

(Curtis, 1829)

• Chaenon elegans Curtis, 1829

• brevicornis (Curtis, 1829, Chaenon)

• cingulatus (Curtis, 1829, Chaenon)

• rufinotatus (Curtis, 1829, Chaenon)

• similis (Curtis, 1829, Chaenon)

Distribution

England, Ireland

Coelinidea fuliginosus

(Curtis, 1829)

• Chaenon fuliginosus Curtis, 1829

Distribution

England

Coelinidea gracilis

(Curtis, 1829)

• Chaenon gracilis Curtis, 1829

• attenuator (Zetterstedt, 1838, Lepton)

Distribution

England, Scotland, Ireland

Coelinidea niger

(Nees, 1811)

• Stephanus niger Nees, 1811

• affinis (Curtis, 1829, Chaenon)

• nigricans (Westwood, 1835, Chaenon)

• olivieri (Guérin-Ménéville, 1842, Alysia)

Distribution

England, Ireland

Coelinidea obscurus

(Curtis, 1829)

• Chaenon obscurus Curtis, 1829

Distribution

England

Coelinidea podagricus

(Haliday, 1839)

• Alysia podagrica Haliday, 1839

Distribution

Ireland

Coelinidea ruficollis

(Herrich-Schäffer, 1838)

• Coelinius ruficollis Herrich-Schäffer, 1838

• procerus (Haliday, 1839, Alysia)

Distribution

England

Coelinidea viduus

(Curtis, 1829)

• Chaenon viduus Curtis, 1829

• ater (Curtis, 1837, Chaenon) nom. nud.

Distribution

England, Ireland

Coelinius

Nees, 1818

• CHAENON Curtis, 1829

• COPISURA Schiødte,1837

• COPIDURA Förster,1862

Notes

Griffiths (1964) proposed that Polemochartus and Coelinidea should be included as subgenera of Coelinius. Wharton and Austin (1991), Wharton (1994) and Kula (2008) supported the concept of an enlarged Coelinius but, pointing to intermediate taxa in the Oriental Region, argued against the retention of subgenera. Achterberg (2014) retains the three genera which is current European usage and is followed here pending a modern review of the group.

Coelinius parvulus

(Nees, 1811)

• Chaenon parvulus Nees, 1811

• Coelinius parvulus ?cultriformis (Latreille, 1802, Ichneumon)

• circulator (Gravenhorst, 1807, Ichneumon) preocc.

• anceps (Curtis, 1829, Chaenon)

• rimator (Schiødte, 1837, Copisura)

• bicarinatus Herrich-Schäffer, 1838

• flexuosus Herrich-Schäffer, 1838

• bicolor Maréchal, 1938

Distribution

England, Scotland, Ireland, Isle of Man

Coloneura

Förster, 1863

• ISOMERISTA Förster, 1863

• TRISISA Förster, 1863

• MERITES Nixon, 1943

• PRIAPSIS Nixon, 1943

Coloneura dice

(Nixon, 1943)

• Priapsis dice Nixon, 1943

Distribution

England

Coloneura stylata

Förster, 1863

• exilis (Förster, 1863, Trisisa)

• oligomera (Förster, 1863, Isomerista)

• taras (Nixon, 1943, Merites)

Distribution

England, Ireland

Dacnusa

Haliday, 1833

• AGONIA Förster, 1863

• APHANTA Förster, 1863

• BRACHYSTROPHA Förster, 1863

• LIPOSCIA Förster, 1863

• PACHYSEMA Förster, 1863

• RHIZARCHA Förster, 1863

• TANYSTROPHA Förster, 1863

• RADIOLARIA Provancher, 1886

• COLONEURELLA van Achterberg, 1976

Dacnusa abdita

Haliday, 1838

• incidens Thomson, 1895

• lepida Marshall, 1896

Distribution

England, Scotland, Wales, Ireland

Dacnusa adducta

(Haliday, 1839)

• Alysia adducta Haliday, 1839

• abducta misspelling

Distribution

England, Scotland, Ireland

Notes

Based on its unusual venation, adducta is often placed in the monotypic genus Agonia; Griffiths’ (Griffiths 1964) argument that it is a derived species of Dacnusa is followed here.

Dacnusa alticeps

Nixon, 1937

Distribution

England, Scotland

Dacnusa aquilegiae

Marshall, 1896

Distribution

England, Scotland

Dacnusa areolaris

(Nees, 1811)

• Bracon areolaris Nees, 1811

• lysias Goureau, 1851

Distribution

England, Scotland, Wales, Ireland

Dacnusa astarte

(Nixon, 1948)

• Rhizarcha astarte Nixon, 1948

Distribution

England

Dacnusa aterrima

Thomson, 1895

Distribution

England

Dacnusa confinis

Ruthe, 1859

• minuta (Curtis, 1826, Alysia)

Distribution

England, Wales, Ireland

Notes

Alysia minuta was listed as a species of Alysia by Shenefelt (1974) and Huddleston (1978) although a junior homonym of Alysia minuta Nees, 1812; Wharton (1986) showed it to belong to Dacnusa and van Achterberg (in litt.) concluded that it is conspecific with confinis.

Dacnusa delphinii

Griffiths, 1967

Distribution

England

Dacnusa discolor

(Förster, 1863)

• Liposcia discolor Förster, 1863

• cercides (Nixon, 1954, Pachysema)

Distribution

England, Scotland, Ireland

Dacnusa dryas

(Nixon, 1948)

• Rhizarcha dryas Nixon, 1948

Distribution

England

Dacnusa ergeteles

(Nixon, 1954)

• Pachysema ergeteles Nixon, 1954

Distribution

Ireland

Dacnusa euphrasiella

Griffiths, 1984

Distribution

Ireland

Notes

added by Griffiths (1984)

Dacnusa evadne

Nixon, 1937

Distribution

England, Scotland, Ireland

Dacnusa faeroeensis

(Roman, 1917)

• Rhizarcha faeroeensis Roman, 1917

• lestes Nixon, 1937

Distribution

England, Scotland, Wales, Ireland

Dacnusa fasciata

Stelfox, 1954

Distribution

Ireland

Dacnusa hospita

(Förster, 1863)

• Aphanta hospita Förster, 1863

Distribution

England, Scotland, Ireland

Dacnusa laeta

(Nixon, 1954)

• Pachysema laeta Nixon, 1954

Distribution

England, Scotland, Ireland

Dacnusa laevipectus

Thomson, 1895

• nox (Morley, 1924, Rhizarcha)

Distribution

England, Scotland, Wales, Ireland

Dacnusa lissos

(Nixon, 1954)

• Pachysema lissos Nixon, 1954

Distribution

England

Dacnusa longiradialis

Nixon, 1937

Distribution

England, Scotland, Ireland

Dacnusa lugens

(Haliday, 1839)

• Alysia lugens Haliday, 1839

Distribution

England, Ireland

Dacnusa macrospila

(Haliday, 1839)

• Alysia macrospila Haliday, 1839

Distribution

England, Scotland, Ireland

Dacnusa maculipes

Thomson, 1895

Distribution

England, Scotland, Wales, Ireland

Dacnusa maxima

(Fischer, 1961)

• Pachysema maxima Fischer, 1961

Distribution

Wales

Dacnusa melicerta

(Nixon, 1954)

• Pachysema melicerta Nixon, 1954

• fumipes Tobias, 1998

Distribution

England, Scotland, Ireland

Dacnusa merope

(Nixon, 1948)

• Rhizarcha merope Nixon, 1948

Distribution

England

Dacnusa metula

(Nixon, 1954)

• Pachysema metula Nixon, 1954

Distribution

England, Scotland, Ireland

Dacnusa monticola

(Förster, 1863)

• Brachystropha monticola Förster, 1863

• mutia (Nixon, 1948, Rhizarcha)

• coracina Stelfox, 1957

Distribution

Ireland

Dacnusa nigrella

Griffiths, 1967

Distribution

Scotland

Notes

added by Godfray and Bland (2011)

Dacnusa nigropygmaea

Stelfox, 1954

Distribution

Ireland

Dacnusa obesa

Stelfox, 1954

Distribution

Scotland, Ireland

Dacnusa ocyroe

Nixon, 1937

Distribution

England, Scotland, Wales, Ireland

Dacnusa plantaginis

Griffiths, 1967

• discolor misident.

Distribution

England, Scotland, Wales, Ireland

Dacnusa pubescens

(Curtis, 1826)

• Alysia pubescens Curtis, 1826

• exserens (Nees, 1834, Alysia)

Distribution

England, Scotland, Ireland

Dacnusa sibirica

Telenga, 1934

Distribution

England, Scotland, Wales, Ireland

Dacnusa comis

(Nixon, 1954)

• Pachysema comis Nixon, 1954

Dacnusa soma

(Nixon, 1948)

• Rhizarcha soma Nixon, 1948

Distribution

Scotland, Ireland

Dacnusa stramineipes

(Haliday, 1839)

• Alysia stramineipes Haliday, 1839

• haemorrhoa (Förster, 1863, Tanystropha)

• longicauda Thomson, 1895

Distribution

England, Scotland, Ireland

Notes

Dacnusa longicauda was synonymised by Nixon (1937) although this was overlooked by Huddleston (1978).

Dacnusa tarsalis

Thomson, 1895

• nitetis (Nixon, 1948, Rhizarcha)

Distribution

England, Scotland, Wales, Ireland

Dacnusa temula

(Haliday, 1839)

• Alysia temula Haliday, 1839

Distribution

England, Scotland, Ireland

Dacnusa veronicae

Griffiths, 1967

Distribution

England, Scotland

Epimicta

Förster, 1863

Epimicta marginalis

(Haliday, 1839)

• Alysia marginalis Haliday, 1839

Distribution

England, Ireland

Exotela

Förster, 1863

• MESORA Förster, 1863

• TOXELEA Nixon, 1943

• ANTRUSA Nixon, 1943

Notes

Griffiths’ (Griffiths 1964, Griffiths 1967a, Griffiths 1967b, Griffiths 1967c, Griffiths 1968a, Griffiths 1968b) concept of Exotela is followed here; the most plesiomorphic species (flavicoxa, interstitialis, melanocera and vaenia) are often placed in Antrusa (e.g. Achterberg 2014).

Exotela cyclogaster

Förster, 1863

• bellina (Nixon, 1937, Dacnusa)

• umbellina (Nixon, 1954, Toxelea)

• sonchina Griffiths, 1967

Distribution

England, Scotland, Wales, Ireland

Notes

Griffiths (1967c) divided cyclogaster into three subspecies, nominate cyclogaster, umbellina and sonchina. Tobias (1986) later treated these as separate species although Godfray (1984) argued that umbellina and cyclogaster are host race variants. Pending further research they are treated here as a single taxon.

Exotela dives

(Nixon, 1954)

• Toxelea dives Nixon, 1954

Distribution

Ireland

Exotela flavicoxa

(Thomson, 1895)

• Dacnusa flavicoxa Thomson, 1895

Distribution

England, Scotland, Ireland

Exotela gilvipes

(Haliday, 1839)

• Alysia gilvipes Haliday, 1839

• albilabris (Thomson, 1895, Dacnusa)

Distribution

England, Scotland, Ireland

Exotela hera

(Nixon, 1937)

• Dacnusa hera Nixon, 1937

Distribution

England, Scotland, Isle of Man

Exotela interstitialis

(Thomson, 1895)

• Dacnusa interstitialis Thomson, 1895

• mamertes (Nixon, 1943, Dacnusa)

Notes

Placed in Chorebus by Tobias and Jakimavicius (1973) and in Antrusa by Papp (2007); needs further research.

Exotela lonicerae

Griffiths, 1967

Distribution

England, Scotland

Exotela melanocera

(Thomson, 1895)

• Dacnusa melanocera Thomson, 1895

• persimilis (Nixon, 1954, Antrusa)

Distribution

England, Ireland

Exotela phryne

(Nixon, 1954)

• Toxelea phryne Nixon, 1954

Distribution

England, Scotland

Exotela spinifer

(Nixon, 1954)

• Toxelea spinifer Nixon, 1954

Distribution

England, Scotland, Ireland

Exotela sulcata

(Tobias, 1962)

• Pachysema sulcata Tobias, 1962

Distribution

England, Scotland, Ireland

Exotela vaenia

(Nixon, 1954)

• Antrusa vaenia Nixon, 1954

Distribution

England

Exotela viciae

Griffiths, 1984

Distribution

England

Notes

added by Griffiths (1984)

Laotris

Nixon, 1943

Laotris striatula

(Haliday, 1839)

• Alysia striatula Haliday, 1839

Distribution

England, Scotland, Ireland

Polemochartus

Nees, 1818

Polemochartus liparae

(Giraud, 1863)

• Polemon liparae Giraud, 1863

Distribution

England

Polemochartus melas

(Giraud, 1863)

• Polemon melas Giraud, 1863

Distribution

England

Notes

added by Shaw and Jennings (2008)

Protodacnusa

Griffiths, 1964

Protodacnusa litoralis

Griffiths, 1964

Distribution

Ireland

Protodacnusa tristis

(Nees, 1834)

• Alysia tristis Nees, 1834

• ampliator (Haliday, 1839, Alysia) preocc.

• longistigma (Telenga, 1935, Dacnusa)

Distribution

England, Ireland

Sarops

Nixon, 1942

Sarops rea

Nixon, 1942

Distribution

England

Synelix

Förster, 1863

• ECTILIS Nixon, 1943

Synelix semirugosa

(Haliday, 1839)

• Alysia semirugosa Haliday, 1839

• agnata Förster, 1863

• amaurosomae (Telenga, 1935, Dacnusa)

Distribution

England, Scotland, Wales, Ireland

Tates

Nixon, 1943

Tates heterocera

(Thomson, 1895)

• Dacnusa heterocera Thomson, 1895

Distribution

England, Scotland

Trachionus

Haliday, 1833

Trachionus

Haliday, 1833

• AENONE Curtis, 1837 preocc.

• AENONE Haliday, 1838 preocc., nom. nud.

• OENONE Haliday, 1839 preocc.

• SYMPHYA Förster, 1863

• ANARMUS Ruthe, 1882

Notes

Generic synonymy follows Achterberg (1997).

Trachionus (Trachionus) mandibularis

(Nees, 1816)

• Sigalphus mandibularis Nees, 1816

Distribution

England, Wales, Ireland

Planiricus

Perepechayenko, 2000

Trachionus (Planiricus) hians

(Nees, 1816)

• Sigalphus hians Nees, 1816

Distribution

England, Scotland, Ireland

Trachionus (Planiricus) ringens

(Haliday, 1839)

• Alysia ringens Haliday, 1839

Distribution

England, Scotland, Ireland

Aphidiinae

Aphidiinae

Haliday, 1833

Notes

In the older literature (and some very recent literature) often treated as a separate family (i.e. Aphidiidae) within Ichneumonoidea. Some distribution data from Mackauer (1961), Starý (1978) and Baker (2013).

Aphidiini

Haliday, 1833

Adialytus

Förster, 1863

Adialytus ambiguus

(Haliday, 1834)

• Aphidius ambiguus Haliday, 1834

• diminuens (Nees, 1834, Aphidius)

• Adialytus ambiguus ?exiguus (Haliday, 1834, Aphidius)

• delhiensis (Subba Rao & Sharma, 1960, Aphidius)

• arvicola (Starý, 1961, Lysiphlebus)

• mackaueri (Starý, 1961, Lysiphlebus)

• crocinus (Mackauer, 1962, Lysiphlebus)

Distribution

Scotland, Wales

Notes

The type of exiguus is lost (Achterberg 1997).

Adialytus salicaphis

(Fitch, 1855)

• Trioxys salicaphis Fitch, 1855

• populaphis (Fitch, 1855, Trioxys)

• tenuis Förster, 1863

• salicaphidis (Ashmead, 1889, Lipolexis)

• laticephalus (Telenga, 1953, Aphidius)

Distribution

England, Wales

Notes

added by Baker and Broad (2009)

Aphidius

Nees, 1818

• INCUBUS Schrank, 1802 nom. ob.

• THERACMION Holmgren, 1872

• EUAPHIDIUS Mackauer, 1961

Notes

doubtfully placed species of Aphidius:

[constrictus (Nees, 1811, Bracon)] Included as a British species by Lyle (1933) on the basis of Szépligeti’s (Szépligeti 1904) Palaearctic catalogue; the type is lost and the species has not been interpreted by recent authors.

[dimidiatus Curtis, 1831]

[pallidinotus Haliday, 1834 nom. nud.]

[pseudoplatanus Curtis, 1837; constrictus misident.] There have been no references to this species other than catalogue listings (Smith 1853, Marshall 1872). Although proposed (Curtis 1837) as a name for the species called Aphidius constrictus by Haliday, it has not been interpreted by recent authors.

[viminalis Haliday, 1834 nom. nud.]

Aphidius absinthii

Marshall, 1896

Distribution

England

Notes

Treated as a valid species of Aphidius by Rakhshani et al. (2011) rather than a synonym of asteris.

Aphidius aquilus

Mackauer, 1961

• sicarius Mackauer, 1961

• callipterinellae (Takada, 1966, Lysaphidus)

Distribution

England, Wales

Aphidius asteris

Haliday, 1834

• melanocephalus (Nees, 1811, Bracon)

• lutescens Haliday, 1834

• commodus Gahan, 1926

• artemisiae Ivanov, 1927

Distribution

England, Wales

Aphidius avenae

Haliday, 1834

• picipes (Nees, 1811, Bracon) suppressed

• crithmi Marshall, 1896

• granarius Marshall, 1896

• pascuorum Marshall, 1896

• hungaricus (Györfi, 1958, Lysiphlebus)

• caraganae Starý, 1963

Distribution

England, Wales, Ireland

Aphidius cingulatus

Ruthe, 1859

• arcticus (Holmgren, 1872, Theracmion)

• gregarius Marshall, 1872

• lachni Ashmead, 1889

• pterocommae Ashmead, 1889

• pterocommae Marshall, 1896 preocc.

• luzhetzki Telenga, 1958

Distribution

England, Wales

Aphidius colemani

Viereck, 1912

• huebrichi Brèthes, 1913

• platensis Brèthes, 1913

• porteri Brèthes, 1915

• aphidiphilus Benoit, 1955

• leroyi Benoit, 1955

• transcaspicus Telenga, 1958

Notes

#Introduced into greenhouses for biocontrol (Starý 1975).

Aphidius eadyi

Starý, Gonzalez & Hall, 1980

• urticae misident.

Distribution

England

Notes

added by Müller et al. (1999)

Aphidius eglanteriae

Haliday, 1834

Aphidius ervi

Haliday, 1834

• infirmus (Nees, 1811, Bracon)

• ulmi Marshall, 1896

• medicaginis Marshall, 1898

• fumipennis Györfi, 1958

• nigrescens Mackauer, 1962

• mirotarsi Starý, 1963

Distribution

England, Wales, Ireland

Aphidius fumatus

Haliday, 1834

Aphidius funebris

Mackauer, 1961

• cirsii Ivanov, 1925 preocc.

• bispinosus Telenga, 1958

• eriophori Mackauer, 1967

Distribution

England, Wales

Notes

added by Pennacchio (1989)

Aphidius hortensis

Marshall, 1896

• berberidis Smith, 1944

Distribution

England, Wales

Aphidius matricariae

Haliday, 1834

• arundinis Haliday, 1834

• cirsii Haliday, 1834 preocc.

• phorodontis Ashmead, 1889

• chrysanthemi Marshall, 1896

• lychnidis Marshall, 1896

• polygoni Marshall, 1896

• affinis Quilis, 1931

• baudysi Quilis, 1931

• discrytus Quilis, 1931

• merceti Quilis, 1931

• obscuriformis Quilis, 1931

• valentinus Quilis, 1931

• renominatus Hincks, 1943

• nigriteleus Smith, 1944

Distribution

England, Wales, Ireland

Aphidius microlophii

Pennacchio & Tremblay, 1987

Distribution

England, Wales

Notes

Added by Pennacchio (1989); also recorded by Müller et al. (1999).

Aphidius rhopalosiphi

de Stefani-Perez, 1902

• equiseticola Starý, 1963

• poacearum Starý, 1963

Distribution

England

Aphidius ribis

Haliday, 1834

• ribaphidis (Ashmead, 1889, Lysiphlebus)

• scabiosae Marshall, 1896

• ribis Ashmead, 1898 preocc.

Distribution

England

Aphidius rosae

Haliday, 1833

• aphidum (Linnaeus, 1758, Ichneumon) nom. ob.

• aphidator (Thunberg, 1824, Ichneumon) nom. ob.

• rosae Curtis, 1831 nom. nud.

• protaeus Wesmael, 1835

• rosarum Nees, 1834

• xanthostoma Bouché, 1834

• protaeus Wesmael, 1835

• cancellatus Buckton, 1876

Distribution

England, Wales, Ireland

Aphidius rubi

Starý, 1962

Distribution

England

Notes

Added by Müller et al. (1999); Belshaw (appendix in Müller et al. 1999) demonstrates that this species should be treated as separate from urticae.

Aphidius salicis

Haliday, 1834

• restrictus Nees, 1834

• duodecimarticulatus Ratzeburg, 1852

• dauci Marshall, 1896

Distribution

England, Wales

Aphidius setiger

(Mackauer, 1961)

• Euaphidius setiger Mackauer, 1961

• aceri Ivanov, 1925

Distribution

Wales

Notes

added by Baker and Broad (2009)

Aphidius sonchi

Marshall, 1896

Distribution

England, Wales

Aphidius tanacetarius

Mackauer, 1962

• tanaceti Curtis, 1837 nom. nud.

• tanaceticola Starý, 1963

Notes

Belokobylskij et al. (2003) treat tanacetarius as the valid name for tanaceti, which is a nomen nudum. This species has not been listed as British since its original description (Curtis 1837) and Smith’s (Smith 1853) catalogue. Pungerl (1986) treated tanacetarius as a valid species but did not see any British or Irish material.

Aphidius urticae

Haliday, 1834

• euphorbiae Marshall, 1896

• longulus Marshall, 1896

• lonicerae Marshall, 1896

• silenes Marshall, 1896

• goidanichi Quilis, 1932

• ivanovae Telenga, 1958

• rubi Starý, 1962

• silvaticus Starý, 1962

• aulacorthi Starý, 1963

Distribution

England

Aphidius uzbekistanicus

Luzhetski, 1960

• urticae misident.; Powell (1982)

• Aphidius uzbekistanicus ?beltrani Quilis, 1931

• Aphidius uzbekistanicus ?indivisus Quilis, 1931

• Aphidius uzbekistanicus ?macropterus Quilis, 1931

• Aphidius uzbekistanicus ?pailloti Quilis, 1931

• impressus Mackauer, 1965

Distribution

England

Notes

Aphidius beltrani is a possible senior synonym of uzbekistanicus (Starý 1973).

Betuloxys

Mackauer, 1960

Betuloxys compressicornis

(Ruthe, 1859)

• Trioxys compressicornis Ruthe, 1859

• testaceus (Stelfox, 1948, Trioxys)

Distribution

England, Wales, Ireland

Binodoxys

Mackauer, 1960

• MISAPHIDUS Rondani, 1848 nom. ob.

Notes

In Belokobylskij et al. (2003), van Achterberg listed these species under the genus name Misaphidus but, according to van Achterberg (pers. comm.), Misaphidus Rondani, 1848 s.l. (type-species: Misaphidus crudelis Rondani, 1848 (= Aphidius centaureae Haliday, 1833)) is a nomen oblitum according to Article 23.9 of ICZN.

Binodoxys acalephae

(Marshall, 1896)

• Aphidius acalephae Marshall, 1896

• rietscheli (Mackauer, 1959, Trioxys)

• urticae (Mackauer, 1959, Trioxys)

Distribution

England

Binodoxys angelicae

(Haliday, 1833)

• Aphidius angelicae Haliday, 1833

• placidus (Gautier, 1922, Trioxys)

• boscai (Quilis, 1931, Trioxys)

• fumariae (Quilis, 1931, Trioxys)

• granatensis (Quilis, 1931, Trioxys)

• obscuriformis (Quilis, 1931, Trioxys)

• amoplanus (Quilis, 1934, Trioxys)

• mediterraneus (Mackauer, 1960, Trioxys)

• wollastonii (Cabrera, 1962, Trioxys) nom. nud.

• sikkimensis (Raychaudhuri, Samanta, Pramanik, Tamili & Sarkar, 1990, Trioxys)

Distribution

Wales

Binodoxys brevicornis

(Haliday, 1833)

• Aphidius brevicornis Haliday, 1833

• minutus (Haliday, 1833, Aphidius)

Distribution

Wales, Ireland

Binodoxys centaureae

(Haliday, 1833)

• Aphidius centaureae Haliday, 1833

• crudelis (Rondani, 1848, Misaphidus)

• orientalis (Starý & Schlinger, 1967, Trioxys)

• uroleucon Takada & Rishi, 1980

Distribution

England, Scotland, Wales, Ireland

Binodoxys heraclei

(Haliday, 1833)

• Aphidius heraclei Haliday, 1833

• obsoletus (Wesmael, 1835, Aphidius)

• variegator (Szépligeti, 1898, Trioxys)

Distribution

Wales

Binodoxys letifer

(Haliday, 1833)

• Aphidius letifer Haliday, 1833

Distribution

Ireland

Diaeretellus

Starý, 1960

Diaeretellus ephippium

(Haliday, 1834)

• Aphidius ephippium Haliday, 1834

Distribution

England, Ireland

Diaeretiella

Starý, 1960

Diaeretiella rapae

(McIntosh, 1855)

• Aphidius rapae McIntosh, 1855

• vulgaris (Bouché, 1834, Aphidius)

• rapae (Curtis, 1860, Aphidius) preocc.

• chenopodii (Förster, 1867, Diaeretus) nom. nud.

• halticae (Rondani, 1877, Misaphidus)

• piceus (Cresson, 1879, Trioxys)

• chenopodiaphidis (Ashmead, 1889, Lipolexis)

• ferruginipes (Ashmead, 1890, Diaeretus) nom. nud.

• brassicae (Marshall, 1896, Aphidius)

Distribution

England, Wales

Diaeretus

Förster, 1863

Diaeretus leucopterus

(Haliday, 1834)

• Aphidius leucopterus Haliday, 1834

• exspectatus (Gautier & Bonnamour, 1936, Aphidius)

Distribution

England, Wales

Harkeria

Cameron, 1900

• PARAMONOCTONUS Starý, 1959

Harkeria rufa

Cameron, 1900

Distribution

England

Notes

Distribution from Achterberg (1989), who elevated Harkeria from a subgenus of Monoctonus.

Lipolexis

Förster, 1863

• GYNOCRYPTUS Quilis, 1931

Lipolexis gracilis

Förster, 1863

• palpator (Gautier & Bonnamour, 1931, Aphidius)

• pieltaini (Quilis, 1931, Gynocryptus)

• chinensis Chen, 1980

Distribution

England

Notes

BMNH, det. Torrance & Broad, added here

Lysaphidus

Smith, 1944

Lysaphidus schimitscheki

Starý, 1960

Distribution

England

Lysaphidus

Förster, 1863

Lysiphlebus

Förster, 1863

• PLATYCYPHUS Mackauer, 1960

Lysiphlebus (Lysiphlebus) dissolutus

(Nees, 1811)

• Bracon dissolutus Nees, 1811

• macrocornis Mackauer, 1960

Distribution

England

Phlebus

Starý, 1975

• APHIDARIA Provancher, 1888 preocc.

Lysiphlebus (Phlebus) confusus

Tremblay & Eady, 1978

Distribution

England, Wales

Notes

Added here on the basis of specimens in BMNH, identified by R.D. Eady as ambiguus sensu Mackauer and Starý nec Haliday, later described by Tremblay and Eady (1978) as a separate species, confusus, the true ambiguus belonging in Adialytus.

Lysiphlebus (Phlebus) fabarum

(Marshall, 1896)

• Aphidius fabarum Marshall, 1896

• aphidiperda (Rondani, 1877, Misaphidus)

• monilicornis (Thomson, 1895, Aphidius)

• cardui (Marshall, 1896, Aphidius)

• aurantii (Pierantoni, 1907, Aphidius)

• polygoni (Ivanov, 1927, Aphidius) preocc.

• gomezi (Quilis, 1930, Aphidius)

• janinii (Quilis, 1930, Aphidius)

• inermis Quilis, 1931

• innovatus Quilis, 1931

• moroderi Quilis, 1931

• ivanovi Mackauer, 1967

Distribution

England, Wales

Monoctonia

Starý, 1962

Monoctonia vesicarii

Tremblay, 1991

Distribution

England

Notes

added by Rakhshani et al. (2015)

Monoctonus

Haliday, 1833

Falciconus

Mackauer, 1961

Monoctonus (Falciconus) pseudoplatani

(Marshall, 1896)

• Aphidius pseudoplatani Marshall, 1896

Distribution

England, Wales

Monoctonus

Haliday, 1833

Monoctonus (Monoctonus) caricis

(Haliday, 1833)

• Aphidius caricis Haliday, 1833

Distribution

England, Wales

Monoctonus (Monoctonus) cerasi

(Marshall, 1896)

• Aphidius cerasi Marshall, 1896

Distribution

England

Monoctonus (Monoctonus) crepidis

(Haliday, 1834)

• Aphidius crepidis Haliday, 1834

• tuberculatus (Wesmael, 1835, Aphidius)

• paludum Marshall, 1896

Distribution

England, Wales

Monoctonus (Monoctonus) ligustri

van Achterberg, 1989

Distribution

England, Wales

Notes

added by Achterberg (1989)

Monoctonus (Monoctonus) nervosus

(Haliday, 1833)

• Aphidius nervosus Haliday, 1833

• paulensis (Ashmead, 1902, Aphidius)

• secundus Viereck, 1915

• biroi Györfi, 1958

• breviantennalis Starý, 1959

Paralipsis

Förster, 1863

• Myrmecobosca Maneval, 1940

Paralipsis enervis

(Nees, 1834)

• Aphidius enervis Nees, 1834

• mandibularis (Maneval, 1940, Myrmecobosca)

• linnei (Hincks, 1949, Myrmecobosca)

Distribution

England, Wales

Pauesia

Quilis Pérez, 1931

Paraphidius

Starý,1958

Notes

species of Pauesia (Paraphidius) excluded from the British and Irish list:

[silvestris (Starý, 1960, Paraphidius)] added by Enobakhare (2001) but we do not know where Enobokhare’s material is deposited and the identification needs to be checked.

Pauesia (Paraphidius) abietis

(Marshall, 1896)

• Aphidius abietis Marshall, 1896

Distribution

England

Pauesia (Paraphidius) cupressobii

(Starý, 1960)

• Paraphidius cupressobii Starý, 1960

Distribution

Scotland

Notes

BMNH, det. Gärdenfors, added here

Pauesia (Paraphidius) juniperorum

(Starý, 1960)

• Paraphidius juniperorum Starý, 1960

Distribution

England, Scotland

Pauesia (Paraphidius) pini

(Haliday, 1834)

• Aphidius pini Haliday, 1834

• planistipes (Nees,1834, Aphidius)

• varia (Nees, 1834, Aphidius)

• panzerii (Rondani, 1848, Aphidius)

• lachnivorus (Ashmead, 1906, Aphidius)

Distribution

England, Scotland, Wales, Ireland

Pauesia

Quilis Pérez, 1931

Pauesia (Pauesia) infulata

(Haliday, 1834)

• Aphidius infulatus Haliday, 1834

• albiflagellaris (Starý, 1960, Paraphidius)

Distribution

Scotland

Pauesia (Pauesia) laricis

(Haliday, 1834)

• Aphidius laricis Haliday, 1834

Distribution

Wales

Pauesia (Pauesia) picta

(Haliday, 1834)

• Aphidius pictus Haliday, 1834

Distribution

Wales

Pauesia (Pauesia) unilachni

(Gahan, 1926)

• Aphidius unilachni Gahan, 1926

• albuferensis Quilis, 1931

• praevisus (Gautier & Bonnamour, 1936, Aphidius)

• basilewskyi (Benoit, 1955, Trioxys)

Distribution

England, Wales

Notes

added by Belshaw and Quicke (1997)

Trioxys

Haliday, 1833

Pectoxys

Mackauer, 1960

Trioxys (Pectoxys) macroceratus

Mackauer, 1960

Trioxys

Haliday, 1833

• APHIDILEO Rondani, 1877

• NEVROPENES Provancher, 1886

• BIOXYS Starý & Schlinger, 1967

Trioxys (Trioxys) auctus

(Haliday, 1833)

• Aphidius auctus Haliday, 1833

• flaviceps Szépligeti, 1898

Distribution

Ireland

Trioxys (Trioxys) betulae

Marshall, 1896

• solani Ivanov, 1925

• hincksi Mackauer, 1960

Distribution

England, Wales

Trioxys (Trioxys) cirsii

(Curtis, 1831)

• Aphidius cirsii Curtis, 1831

• aceris (Haliday, 1833, Aphidius)

Distribution

England, Wales, Ireland

Trioxys (Trioxys) curvicaudus

Mackauer, 1967

Distribution

England, Wales

Notes

added by Starý (1978)

Trioxys (Trioxys) falcatus

Mackauer, 1959

Distribution

Wales

Notes

added by Baker and Broad (2009)

Trioxys (Trioxys) ibis

Mackauer, 1961

Trioxys (Trioxys) pallidus

(Haliday, 1833)

• Aphidius pallidus Haliday, 1833

• resolutus (Nees, 1834, Aphidius)

• callipteri (Marshall, 1896, Aphidius)

• pulcher Gautier & Bonnamour, 1924

Distribution

England, Wales

Trioxys (Trioxys) tenuicaudus

Starý, 1978

Distribution

Wales

Notes

added by Baker and Broad (2013)

Ephedrini

Mackauer, 1961

Ephedrus

Haliday, 1833

Notes

Synonymy and distribution data from Gärdenfors (1986).

Breviephedrus

Gärdenfors,1986

Ephedrus (Breviephedrus) brevis

Stelfox, 1941

• picticornis Stelfox, 1941

• niger Stelfox, 1941 nom. nud.

Distribution

England, Scotland, Ireland

Ephedrus

Haliday, 1833

• ELASSUS Wesmael, 1835

Ephedrus (Ephedrus) cerasicola

Starý, 1962

Distribution

England

Notes

added by Van Veen et al. (2008)

Ephedrus (Ephedrus) helleni

Mackauer, 1968

• salicicola Takada, 1968

Distribution

Scotland

Notes

added by Baker and Broad (2013)

Ephedrus (Ephedrus) lacertosus

(Haliday, 1833)

• Aphidius lacertosus Haliday, 1833

• muesebecki Smith, 1944

Distribution

England, Scotland, Ireland

Ephedrus (Ephedrus) laevicollis

(Thomson, 1895)

• Aphidius laevicollis Thomson, 1895

• brevicornis (Nees, 1834, Aphidius) preocc.

• minor Stelfox, 1941

• salicicola Takada, 1968

Distribution

Wales, Ireland

Ephedrus (Ephedrus) niger

Gautier, Bonnamour & Gaumont, 1929

• Ephedrus (Ephedrus) niger ?aphidivora (Rondani, 1848, Alysia); synonymy by Papp (1996a)

• campestris Starý, 1962

Distribution

England

Notes

added by Van Veen et al. (2008)

Ephedrus (Ephedrus) persicae

Froggat, 1904

• nevadensis Baker, 1909

• nitidus Gahan, 1917

• vidali Quilis, 1931

• interstitialis Watanabe, 1941

• pulchellus Stelfox, 1941

• impressus Granger, 1949

• holmani Starý, 1958

• palaestinensis Mackauer, 1959

Distribution

England, Ireland

Ephedrus (Ephedrus) plagiator

(Nees, 1811)

• Bracon plagiator Nees, 1811

• parcicornis (Nees, 1834, Aphidius)

• japonicus Ashmead, 1906

• homostigma Fahringer, 1934

Distribution

England, Scotland, Wales, Ireland

Lysephedrus

Starý, 1958

Ephedrus (Lysephedrus) validus

(Haliday, 1833)

• Aphidius validus Haliday, 1833

Distribution

England, Ireland

Toxares

Haliday, 1840

• TRIONYX Haliday, 1833 preocc.

• TERONYX Haldeman, 1842

Toxares deltiger

(Haliday, 1833)

• Aphidius deltiger Haliday, 1833

• flaveolus (Györfi, 1958, Ephedrus)

Distribution

England, Ireland

Praini

Mackauer, 1961

Areopraon

Mackauer, 1959

• MESOPRAON Starý, 1981; synonymy by Tomanovic et al. (2006)

Areopraon lepelleyi

(Waterston, 1926)

• Praon lepelleyi Waterston, 1926

Distribution

England

Areopraon silvestre

(Starý, 1971)

• Praon silvestre Starý, 1971

Distribution

Wales

Notes

Added by Baker and Broad (2013); transferred to Areopraon by Tomanovic et al. (2006).

Dyscritulus

Hincks, 1943

• DYSCRITUS Marshall, 1896 preocc.

Notes

Dyscritus suffolciensis Morley, 1933 is listed by default as a species of Dyscritulus in Taxapad (Yu et al. 2012) as, although Starý (1959) recognised it as belonging to Euphorinae, its identity had not yet been established. Here it is synonymised under Syntretus splendidus (Marshall) q.v.

Dyscritulus planiceps

(Marshall, 1896)

• Dyscritus planiceps Marshall, 1896

Distribution

England, Wales

Dyscritulus pygmaeus

Mackauer, 1961

Praon

Haliday, 1833

• ACHORISTUS Ratzeburg, 1852

• APHIDARIA Provancher, 1886

• PARAPRAON Starý, 1983

Notes

species of Praon excluded from the British and Irish list:

[simulans (Provancher, 1886, Aphidaria); syn. aguti Smith, 1944] A Nearctic species, listed by Kloet and Hincks (1945) in error.

Praon abjectum

(Haliday, 1833)

• Aphidius abjectum Haliday, 1833

• aphidiiforme (Ratzeburg, 1852, Bracon)

• peregrinum Ruthe, 1859

Distribution

England, Ireland

Praon absinthii

Bignell, 1894

Notes

Erroneously listed in Taxapad as a junior synonym of flavinode; see, for example, Kavallieratos et al. (2005).

Praon barbatum

Mackauer, 1967

Distribution

England

Notes

BMNH, det. Torrance & Broad, added here

Praon bicolor

Mackauer, 1959

Distribution

Wales

Notes

added by Baker and Broad (2009)

Praon cavariellae

Starý, 1971

Distribution

Scotland, Wales

Notes

added by Baker and Broad (2013)

Praon dorsale

(Haliday, 1833)

• Aphidius dorsale Haliday, 1833

• discolor (Nees, 1834, Blacus)

• collare Förster, 1867 nom. nud.

Distribution

England, Wales, Ireland

Praon exsoletum

(Nees, 1811)

• Bracon exsoletum Nees, 1811

• palitans Muesebeck, 1956

Praon flavinode

(Haliday, 1833)

• Aphidius flavinode Haliday, 1833

• emacerator (Nees, 1834, Blacus)

• glabrum Starý & Schlinger, 1967

Distribution

England, Wales

Praon gallicum

Starý, 1971

Distribution

England

Notes

added by Traugott et al. (2008)

Praon longicorne

(Marshall, 1896)

• Aphidius longicorne Marshall, 1896

• grossum Starý, 1971

Distribution

Wales

Praon necans

Mackauer, 1959

• nympheae Subba Rao, Sarup & Sharma, 1963

Distribution

Wales

Notes

added by Baker and Broad (2013)

Praon spinosum

Mackauer, 1959

Distribution

Wales

Notes

added by Baker and Broad (2009)

Praon volucre

(Haliday, 1833)

• Aphidius volucre Haliday, 1833

• angulator (Nees, 1834, Blacus)

• aphidivorum (Ratzeburg, 1844, Aphidius)

• pruni Ivanov, 1925

• breve Fahringer, 1935

• mongolicum Watanabe, 1949

• myzophagum Mackauer, 1959

Distribution

England, Wales, Ireland

Praon yomenae

Takada, 1968

Distribution

Wales

Notes

added by Baker and Broad (2013)

Brachistinae

Brachistinae

Förster, 1863

• CALYPTINAE Marshall, 1872

Notes

Following Belshaw and Quicke (2002), Achterberg (2003a) and Belokobylskij et al. (2003) we treat the Brachistinae as a subfamily separate from Helconinae. Sharanowski et al. (2011) expanded the limits of the Brachistinae by including the tribe Diospilini and the former subfamily Blacinae (as well as the extralimital Brulleiini). The tribes employed here are those that have been used in previous classifications under the former subfamilies, but the tribal classification remains essentially untested.

Blacini

Förster, 1863

Notes

Generally considered to be a distinct subfamily but Sharanowski et al. (2011) found that the blacines nested within the Brachistinae in their molecular phylogenetic analyses. We think it likely that these results will be upheld with further phylogenetic work and follow Sharanowski et al.’s recommended changes to the subfamily classification. Distribution and taxonomic data from Achterberg (1988a), with additional references given. Note that Tobias (1986) records B. (Neoblacus) koenigi Fischer, 1967 from England but there is no evidence that this species occurs here.

Blacometeorus

Tobias, 1976

Blacometeorus brevicauda

(Hellén, 1958)

• Diospilus brevicauda Hellén, 1958

Distribution

England

Notes

added by Achterberg (1988a); Shaw (1996b)

Blacometeorus pusillus

(Hellén, 1958)

• Diospilus pusillus Hellén, 1958

Distribution

England

Notes

added by Shaw (1996b)

Blacus

Nees, 1818

Blacus

Nees, 1818

• GONIOCORMUS Förster, 1863

• MIOCOLUS Förster, 1863

Blacus (Blacus) errans

(Nees, 1811)

• Bracon errans Nees, 1811

• vagans Ruthe, 1861

Distribution

England

Blacus (Blacus) exilis

(Nees, 1811)

• Bracon exilis Nees, 1811

• lactucaphis (Fitch, 1855, Aphidius)

• pallipes (Förster, 1863, Miocolus)

• pallidipes (Dalla Torre, 1898, Miocolus)

• nanus Ashmead, 1905 nom. nud.

• propallipes Shenefelt, 1969

• intermedius Janzon, 1975

Distribution

England, Ireland

Blacus (Blacus) filicornis

Haeselbarth, 1973

Distribution

England, Ireland

Blacus (Blacus) forticornis

Haeselbarth, 1973

Distribution

England

Blacus (Blacus) hastatus

Haliday, 1835

• terebrator Ruthe, 1861

Distribution

England, Scotland, Ireland

Blacus (Blacus) humilis

(Nees, 1811)

• Bracon humilis Nees, 1811

• trivialis Haliday, 1835

• wesmaeli Ruthe, 1861

Distribution

England, Wales, Ireland

Blacus (Blacus) instabilis

Ruthe, 1861

• petiolatus Haeselbarth, 1973 nom. nud.

Distribution

England, Scotland, Ireland

Blacus (Blacus) leptostigma

Ruthe, 1861

Distribution

Ireland

Blacus (Blacus) longipennis

(Gravenhorst, 1809)

• Ophion longipenne Gravenhorst, 1809

• dubius Ruthe, 1861

Distribution

England, Scotland, Ireland

Blacus (Blacus) nigricornis

Haeselbarth, 1973

Distribution

England, Scotland, Ireland

Blacus (Blacus) paganus

Haliday, 1835

• brevicornis Ruthe, 1861

Distribution

Ireland

Blacus (Blacus) stelfoxi

Haeselbarth, 1973

Distribution

England

Notes

The holotype was collected in Sandhurst, Kent (Haeselbarth 1973) but not listed for Britain or Ireland by Achterberg (1988a).

Ganychorus

Haliday, 1835

Blacus (Ganychorus) ambulans

Haliday, 1835

Distribution

Ireland

Blacus (Ganychorus) armatulus

Ruthe, 1861

Distribution

England

Blacus (Ganychorus) diversicornis

Nees, 1834

• compar Ruthe, 1861

Distribution

England, Ireland

Blacus (Ganychorus) macropterus

Haeselbarth, 1973

Distribution

England, Scotland

Notes

Described as a subspecies of ambulans by Haeselbarth (1973), but not included by Huddleston (1978), where it should have been listed as a synonym. Treated as a separate species by Achterberg (1997).

Blacus (Ganychorus) maculipes

Wesmael, 1835

Distribution

England, Scotland, Ireland

Blacus (Ganychorus) nitidus

Haeselbarth, 1973

• petiolatus Tobias, 1976

Distribution

England

Blacus (Ganychorus) pallipes

Haliday, 1835

• barynoti misident.

• tuberculatus Wesmael, 1835

• florus Goureau, 1851

Distribution

England, Ireland

Notes

Included, as barynoti (Boudier, 1834) (a junior synonym of Pygostolus sticticus), as a doubtfully placed species of Blacus by Huddleston (1978).

Blacus (Ganychorus) ruficornis

(Nees, 1811)

• Bracon ruficornis Nees, 1811

• bisstigmatus (Say, 1836, Microgaster)

• tipulator (Zetterstedt, 1838, Bracon)

• cerealis (Curtis, 1860, Dacnusa)

• pallidipes (Costa, 1885, Dinocampus)

• dentatus Hellén, 1958

Distribution

England, Scotland, Wales, Ireland

Blacus (Ganychorus) strictus

Stelfox, 1941

• strictus (Curtis, 1837, Ganychorus) nom. nud.

Distribution

Scotland, Ireland

Blacus (Ganychorus) tripudians

Haliday, 1835

Distribution

England, Ireland

Hysterobolus

Viereck, 1913

Blacus (Hysterobolus) mamillanus

Ruthe, 1861

• aptenodytes Marshall, 1889

Distribution

England

Brachistini

Förster, 1863

Eubazus

Nees, 1814

Aliolus

Say, 1836

Eubazus (Aliolus) lepidus

(Haliday, 1835)

• Helcon lepidus Haliday, 1835

• hofferi (Šnoflák, 1953, Triaspis); synonymy by Achterberg (2003a)

• kusarensis (Abdinbekova, 1969, Allodorus); synonymy by Achterberg (2003a)

Distribution

England

Allodorus

Förster, 1863

Eubazus (Allodorus) convexope

van Achterberg, 2000

Distribution

Scotland

Notes

added by Achterberg (2000)

Eubazus (Allodorus) semirugosus

(Nees, 1816)

• Sigalphus semirugosus Nees, 1816

• rufipes (Herrich-Schäffer, 1838, Eubadizon)

• tuberculator (Zetterstedt, 1838, Bracon)

• curculionum (Hartig, 1847, Sigalphus)

• atricornis (Ratzeburg, 1848, Brachistes)

• mucronatus (Thomson, 1892, Calyptus)

• truncatus (Thomson, 1892, Calyptus)

• glabratus (Fahringer, 1941, Calyptus)

• arete (Fahringer, 1944, Calyptus)

Distribution

England, Scotland

Eubazus (Allodorus) tricoloripes

van Achterberg, 2000

Distribution

England

Notes

added by Achterberg (2000)

Brachistes

Wesmael, 1835

Eubazus (Brachistes) fasciatus

(Nees, 1816)

• Sigalphus fasciatus Nees, 1816

• fuscipalpis (Wesmael, 1835, Brachistes)

Distribution

England, Ireland

Eubazus (Brachistes) minutus

(Ratzeburg, 1848)

• Brachistes minutus Ratzeburg, 1848

Distribution

England

Eubazus (Brachistes) ruficoxis

(Wesmael, 1835)

• Brachistes ruficoxis Wesmael, 1835

• politus (Ratzeburg, 1852, Brachistes); synonymy by van Achterberg in Belokobylskij et al. (2003)

• byctisci (Watanabe, 1933, Calyptus)

• ruficornis misspelling

Distribution

England

Eubazus (Brachistes) segmentatus

(Marshall, 1889)

• Calyptus segmentatus Marshall, 1889

Distribution

England

Eubazus (Brachistes) semicastaneus

(Marshall, 1893)

• Calyptus semicastaneus Marshall, 1893

Distribution

England

Eubazus (Brachistes) tibialis

(Haliday, 1835)

• Helcon tibialis Haliday, 1835

• uncigenus (Wesmael, 1835, Brachistes)

Distribution

England, Scotland, Ireland

Calyptus

Haliday, 1835

Eubazus (Calyptus) macrocephalus

Nees, 1812

• synchitae (Hedqvist, 1956, Eubadizon)

• ratzeburgi (Fischer, 1962, Eubadizon); synonymy by van Achterberg in Belokobylskij et al. (2003)

• xiphydriae Tobias, 1986; synonymy by van Achterberg in Belokobylskij et al. (2003)

Distribution

England

Eubazus (Calyptus) sigalphoides

(Marshall, 1889)

• Calyptus sigalphoides Marshall, 1889

Distribution

England

Eubazus

Nees, 1814

• Eubadizon Nees, 1834

• Eubadizus Nees, 1834

Eubazus (Eubazus) flavipes

(Haliday, 1835)

• Helcon flavipes Haliday, 1835

• laevis (Herrich-Schäffer, 1838, Eubadizon)

Distribution

England, Ireland

Eubazus (Eubazus) longicauda

(Curtis, 1832)

• Zele longicauda Curtis, 1832

Distribution

England

Notes

Listed as a species of Zele (=Homolobus) in Huddleston (1978). Will be synonymised under another species of Eubazus (van Achterberg, pers. comm.).

Eubazus (Eubazus) pallipes

Nees, 1814

• coxalis (Nees, 1834, Eubadizon)

• semistriatus (Haliday, 1835, Helcon) unavailable

• americanus (Cresson, 1872, Eubadizon)

• pallipede misspelling

• pallidipes misspelling

• pollipes misspelling

Distribution

England

Foersteria

Szépligeti, 1896

Foersteria (Foersteria) laeviuscula

Szépligeti, 1896

Distribution

England, Scotland

Notes

NMS, det. van Achterberg, added here

Foersteria (Foersteria) puber

(Haliday, 1835)

• Helcon puber Haliday, 1835

• opaca (Reinhard, 1867, Calyptus)

• flavipes Szépligeti, 1896

• talitzkii Tobias, 1961

Distribution

England, Ireland

Schizoprymnus

Förster, 1863

Notes

species of Schizoprymnus excluded from the British and Irish list:

[nigripes (Thomson, 1892, Sigalphus)] Seems to have been recorded in error by Kloet and Hincks (1945), perpetuated by Huddleston (1978).

Schizoprymnus ambiguus

(Nees, 1816)

• Sigalphus ambiguus Nees, 1816

Distribution

England, Ireland

Schizoprymnus collaris

(Thomson, 1874)

• Sigalphus collaris Thomson, 1874

Distribution

England

Notes

Added by Notton et al. (2014); listed as a species of Triaspis by Yu et al. (2012) but transferred back to Schizoprymnus by Notton et al. (2014).

Schizoprymnus obscurus

(Nees, 1816)

• Sigalphus obscurus Nees, 1816

Distribution

England

Triaspis

Haliday, 1838

• MUIRIELLA Fullaway, 1919

Notes

species of Triaspis excluded from the British and Irish list:

[striola (Thomson, 1874, Sigalphus)] Listed as a doubtfully British species in Huddleston (1978); the only record is of a tentative identification by Nixon (Hussey 1952).

Triaspis aciculata

(Ratzeburg, 1848)

• Sigalphus aciculatus Ratzeburg, 1848

Distribution

England

Notes

BMNH, NMS, det. van Achterberg, added here; although synonymised with obscurella by van Achterberg (in Belokobylskij et al. 2003) this is listed as a separate species in Fauna Europaea. There are English specimens identified by van Achterberg as aciculata in NMS and BMNH.

Triaspis caledonica

(Marshall, 1888)

• Sigalphus caledonicus Marshall, 1888

Distribution

Scotland

Triaspis caudata

(Nees, 1816)

• Sigalphus caudatus Nees, 1816

• gracilis (Herrich-Schäffer, 1840, Sigalphus)

• australis (Szépligeti, 1901, Sigalphus)

• arctica Hellén, 1958

Distribution

England, Ireland

Triaspis flavipalpis

(Wesmael, 1835)

• Sigalphus flavipalpis Wesmael, 1835

Distribution

England

Triaspis floricola

(Wesmael, 1835)

• Sigalphus floricola Wesmael, 1835

• minima Snoflák, 1953

Distribution

England, Ireland

Triaspis luteipes

(Thomson, 1874)

• Sigalphus luteipes Thomson, 1874

Distribution

England

Triaspis obscurella

(Nees, 1816)

• Sigalphus obscurellus Nees, 1816

• simulator (Szépligeti, 1901, Sigalphus)

Distribution

England

Triaspis pallipes

(Nees, 1816)

• Sigalphus pallipes Nees, 1816

• fulvipes (Haliday, 1835, Helcon)

• fagi (Ratzeburg, 1852, Brachistes)

• similis (Szépligeti, 1901, Sigalphus) preocc.

• pallidipes misspelling

Distribution

England, Ireland, Isle of Man

Notes

some distribution data from Shaw and Askew (1976)

Triaspis podlussanyi

Papp, 1998

Distribution

England

Notes

added by Shaw & Mendel (in prep.)

Triaspis striatula

(Nees, 1816)

• Sigalphus striatulus Nees, 1816

Distribution

England, Ireland

Triaspis thoracica

(Curtis, 1860)

• Sigalphus thoracicus Curtis, 1860

• gibberosa (Szépligeti, 1901, Sigalphus); synonymy by Papp (2004b)

• rugosa (Szépligeti, 1901, Sigalphus); synonymy by Papp (2004b)

Distribution

England

Diospilini

Förster, 1863

Aspicolpus

Wesmael, 1838

• ASPIDOCOLPUS Agassiz, 1846

Aspicolpus clipealis

(Tobias, 1967)

• Aspidocolpus clipealis Tobias, 1967

Distribution

England

Notes

NMS, det. van Achterberg, added here; tentative identification; included here as a certain generic record of an Aspicolpus species in Britain.

Aspigonus

Wesmael, 1835

• ASPIDOGONUS Agassiz, 1846

Aspigonus flavicornis

(Nees, 1834)

• Bracon flavicornis Nees, 1834

• diversicornis Wesmael, 1835; synonymy by van Achterberg in Belokobylskij et al. (2003)

Distribution

England, Wales

Diospilus

Haliday, 1833

• BAEACIS Förster, 1878; synonymy by Achterberg (2014)

• ALLOCHROMUS Marshall, 190

Diospilus abietis

(Ratzeburg, 1844)

• Aspigonus abietis Ratzeburg, 1844

Diospilus capito

(Nees, 1834)

• Bracon capito Nees, 1834

• filator (Nees, 1834, Bracon) preocc.

• fuscipes (Wesmael, 1835, Taphaeus)

Distribution

England, Ireland

Diospilus dispar

(Nees, 1811)

• Bracon dispar Nees, 1811

• ephippium (Nees, 1834, Bracon)

Distribution

England

Diospilus inflexus

Reinhard, 1862

• ovatus Marshall, 1889

Distribution

England

Diospilus intermedius

(Förster, 1878)

• Baeacis intermedia Förster, 1878

Distribution

England

Notes

BMNH, det. van Achterberg, added here

Diospilus morosus

Reinhard, 1862

Distribution

England

Diospilus nigricornis

(Wesmael, 1835)

• Taphaeus nigricornis Wesmael, 1835

• affinis (Wesmael, 1835, Taphaeus)

• rufipes Reinhard, 1862

Distribution

England

Notes

Listed as Taphaeus affinis and T. nigricornis in Huddleston (1978).

Diospilus oleraceus

Haliday, 1833

• conformis (Wesmael, 1835, Taphaeus)

• ruficornis Szépligeti, 1896

Distribution

England, Ireland

Diospilus productus

Marshall, 1894

Distribution

England

Taphaeus

Wesmael, 1835

• Anostenus Förster, 1863

• Anastenus Dalla Torre, 1898

Notes

The systematic position of Taphaeus is uncertain. Treated as a genus of Blacinae in Fauna Europaea.

Taphaeus hiator

(Thunberg, 1824)

• Ichneumon hiator Thunberg, 1824

• irregularis Wesmael, 1835

• speculator (Haliday, 1835, Helcon)

• polydrusi (Gahan, 1916, Diospilus)

Distribution

England, Scotland, Ireland

Vadumasonium

Kammerer, 2006

• VADUM Mason, 1987 preocc.

Vadumasonium vardyorum

van Achterberg & Broad, 2013

Distribution

England

Notes

added by Achterberg and Broad (2013)

Dyscoletini

van Achterberg, 1984

Notes

This is generally considered to be a tribe of Blacinae; we list the Dyscoletini as a tribe of Brachistinae because we follow Sharanowski et al. (2011) in including the Blacinae within Brachistinae.

Dyscoletes

Haliday, 1840

• DYSCOLUS Haliday, 1836 preocc.

• DISCOLUS misspelling

• MICROCENTRUS Szépligeti, 1904 preocc.

• ELACHISTOCENTRUM Schulz, 1911

Dyscoletes lancifer

(Haliday, 1836)

• Dyscolus lancifer Haliday, 1836

• similis (Szépligeti, 1896, Discoletes)

Distribution

England, Scotland

Braconinae

Braconinae

Nees, 1812

Notes

Tribal classification follows Belshaw et al. (2001) and Quicke (pers. comm.). For genera other than Bracon and Coeloides, Shaw (1994) and Shaw and Quicke (1999) give distribution data.

Aphrastobraconini

Ashmead, 1900

Atanycolus

Förster, 1862

Notes

species excluded from the British and Irish list by Shaw and Quicke (1999):

[denigrator (Linnaeus, 1758, Ichneumon); syn. incertus (Sulzer, 1776, Ichneumon); heteropus (Thomson, 1892, Bracon) albiscutis Telenga, 1936]

Cyanopterus

Haliday, 1835

Notes

species excluded from the British and Irish list by Shaw and Quicke (1999):

[flavator (Fabricius, 1793, Ichneumon); syn. flavulator (Ratzeburg, 1844, Bracon); longipalpis (Thomson, 1892, Bracon); barcinonensis (Marshall, 1897, Coeloides)]

Pseudovipio

Szépligeti, 1896

Pseudovipio guttiventris

(Thomson, 1892)

• Bracon guttiventris Thomson, 1892

• variegatus (Boheman, 1853, Agathis) preocc.

• biroi Szépligeti, 1896

Distribution

England

Notes

Listed as Glyptomorpha variegata in Huddleston (1978).

Vipio

Latreille, 1804

Notes

species excluded from the British and Irish list by Shaw and Quicke (1999):

[terrefactor (Villers, 1789, Ichneumon); syn. improvisus Kokujev, 1898; interpellator Kokujev, 1898; neesii Kokujev, 1898] The only record was due to a misidentification of Pseudovipio guttiventris (Shaw and Quicke 1999).

Braconini

Nees, 1812

Baryproctus

Ashmead, 1900

• Barycryptus Hoffmeyer, 1932

Baryproctus barypus

(Marshall, 1885)

• Bracon barypus Marshall, 1885

• hungaricus Szépligeti, 1901

• caucasicus Telenga, 1936

• apti Györfi, 1953

Distribution

England

Notes

See Shaw (1994) and Jennings (2008) for recent records.

Bracon

Fabricius, 1804

Notes

Comprehensive taxonomic revisions of Bracon are lacking for all but a few species groups. Published works differ substantially in their treatments of valid species, synonyms and subgeneric placements (compare, for example, the differing treatments of Tobias (1986), Papp (1974), Papp (1996a), and Shenefelt (1978)) so the taxonomy in Taxapad (Yu et al. 2012) has, for the most part, been used as a default classification.

Bracon

Fabricius, 1804

• BRACO Wesmael, 1838

• BRACHON Agassiz, 1846

• TROPIDOBRACON Ashmead, 1900

• MICROBRACON Ashmead, 1890

• AMICOPLIDEA Ashmead, 1900

• MACRODYCTIUM Ashmead, 1900

• LIOBRACON Nason, 1905 preocc.

• LORENZOA de Stefani-Perez, 1909

• SELIODUS Brèthes, 1909

• KULCZYNSKIA Niezabitowski, 1910

• BRAZON Schulz, 1911

• STRIOBRACON Fahringer, 1927

• CHIVINIA Shestakov, 1932

• EUTROPOBRACON Ayyar, 1928

Bracon (Bracon) alutaceus

Szépligeti, 1901

• pygmaeus Niezabitowski, 1910; synonymy by Papp (2008a)

• polonicus Fahringer, 1927; synonymy by Papp (2008a)

• pallidalatus Tobias, 1957

Distribution

England, Scotland

Notes

NMS, det. Papp, added here

Bracon (Bracon) flavipes

Nees, 1834

Distribution

England

Bracon (Bracon) fulvipes

Nees, 1834

• carinatus Szépligeti, 1901

• marshalli Vayssière, 1902 preocc.

• glabratus Fahringer, 1927

• apionis Strand, 1928

• sylvanus Greese, 1928

• kiritshenkoi Telenga, 1936

Distribution

England, Scotland, Wales, Ireland, Isle of Man

Bracon (Bracon) intercessor

Nees, 1834

• laetus (Wesmael, 1838, Braco); synonymy by Papp (2012), as a 'var' of intercessor

• lativentris Thomson, 1892 preocc.

• fulvus Szépligeti, 1896

• universitatis Dalla Torre, 1898

• adjectus Szépligeti, 1901

• bisinuatus Szépligeti, 1901

• dubiosus Szépligeti, 1901

• duplicatus Szépligeti, 1901

• fallaciosus Szépligeti, 1901

• elegans Szépligeti, 1901

• mixtus Szépligeti, 1901

• mundus Szépligeti, 1901

• nigropictus Szépligeti, 1901

• nitidiusculus Szépligeti, 1901

• rufiscapus Szépligeti, 1901

• subtilis Szépligeti, 1901

• suspectus Szépligeti, 1901

• vigilax Kokujev, 1912

• maidli Fahringer, 1925

• asiaticus Fahringer, 1927 preocc.

• concolor Fahringer, 1927 preocc.

• major Fahringer, 1927

• concorellus Strand, 1928

• megasomides Strand, 1928

• rhynchiti Greese, 1928

• kachetinus Telenga, 1933

• kansensis Fahringer, 1934 preocc.

• maslovskii Telenga, 1936

• segregatus Telenga, 1936

Distribution

England, Scotland

Bracon (Bracon) leptus

Marshall, 1897

• centaureae Szépligeti, 1901

• rufipedator Szépligeti, 1901

• rufipalpis Szépligeti, 1901]

Distribution

England

Notes

Listed as British in Fauna Europaea, probably because it was described by Marshall, but the first published British record was by Jennings (2012).

Bracon (Bracon) longicollis

(Wesmael, 1838)

• Braco longicollis Wesmael, 1838

• fraudator Marshall, 1885

• brevicauda Thomson, 1892 preocc.

• crassicauda Thomson, 1892

• pseudowesmaeli Strand, 1928

• wesmaeli Fahringer, 1927 preocc.

Distribution

England, Scotland, Wales, Isle of Man

Notes

spme distribution data from Papp (1999c)

Bracon (Bracon) luteator

Spinola, 1808

• nigripedator Nees, 1834

• filicauda Costa, 1888

• hypopygialis Szépligeti, 1901

• intermedius Szépligeti, 1901

• pilosulus Szépligeti, 1901

Distribution

England

Notes

added by Fulmek (1968) (omitted by Huddleston 1978)

Bracon (Bracon) nigratus

(Wesmael, 1838)

• Braco nigratus Wesmael, 1838

Distribution

England, Ireland

Bracon (Bracon) pectoralis

(Wesmael, 1838)

• Braco pectoralis Wesmael, 1838

• ochrosus Szépligeti, 1896

• sulphurator Szépligeti, 1896

• unicolor Szépligeti, 1896

• fumigatus Szépligeti, 1901; synonymy by Papp (2008a)

Distribution

England

Bracon (Bracon) rugulosus

Szépligeti, 1901

• depressiusculus Szépligeti, 1901

• neglectus Szépligeti, 1904

• spurnensis Hincks, 1951

Distribution

England, Scotland, Wales

Notes

According to Taxapad (Yu et al. 2012), rugulosus Szépligeti is preoccupied by Bracon rugulosus Nees, 1811 (now classified as a species of Aleiodes, in Rogadinae). Omitted by Huddleston (1978).

Bracon (Bracon) scutellaris

(Wesmael, 1838)

• Braco scutellaris Wesmael, 1838

Distribution

England, Scotland, Ireland

Bracon (Bracon) speerschneideri

Schmiedeknecht, 1897

Distribution

England

Notes

added by Papp (1999a)

Bracon (Bracon) subrugosus

Szépligeti, 1901

• sulcatulus Szépligeti, 1896

• subglaber Szépligeti, 1901

• quinquemaculatus Szépligeti, 1901

• trypetanus Fahringer, 1927; synonymy by Papp (2008a)

• tauricus Telenga, 1936

Distribution

England

Notes

added by Papp (2008a)

Bracon (Bracon) trucidator

Marshall, 1888

• minutator misident.

• bilineatus Thomson, 1892

• hilaris Marshall, 1897

• pannonicus Szépligeti, 1901

• marshalli Telenga, 1936 preocc.

Distribution

England

Notes

BMNH, NMS, added here; previously confused under minutator (van Achterberg, pers. comm.). Both species are present in the BMNH and NMS collections, det. Papp and van Achterberg.

Bracon (Bracon) variegator

Spinola, 1808

• melanosoma Szépligeti, 1901

• micros Szépligeti, 1901

• nanulus Szépligeti, 1901

• lineatellae (Fischer, 1968, Habrobracon); synonymy by Papp (2008b)

Distribution

England, Scotland, Ireland

Glabrobracon

Fahringer, 1927

Notes

Some taxonomic and distribution data from Papp (1999a), Papp (1999c), Papp (2000).

Species of Bracon (Glabrobracon) excluded from the British and Irish list:

[coniferarum Fahringer, 1928] Should not have been listed by Huddleston (1978) as it was recorded as British only on the basis of Carr’s Staffordshire lists (see Perkins 1953, Fitton et al. 1978, Shaw 2003a).

Bracon (Glabrobracon) abbreviator

Nees, 1834

• abscissor Nees, 1834; synonymy by Papp (2008a)

• oestmaeli (Wesmael, 1838, Braco); synonymy by Papp (2008a)

• regularis (Wesmael, 1838, Braco); synonymy by Papp (2008a)

• eutrephes Marshall, 1897

• rufigaster Szépligeti, 1901; synonymy by Papp (2008a)

• rufiventris Telenga, 1936 preocc.

• minimula Strand, 1928

• minimus Fahringer, 1927 preocc.

Distribution

England

Bracon (Glabrobracon) admotus

Papp, 2000

Distribution

England

Notes

BMNH, det. Papp, added here

Bracon (Glabrobracon) albion

Papp, 1999

Distribution

England, Scotland

Notes

added by Papp (1999c)

Bracon (Glabrobracon) arcuatus

Thomson, 1892

Distribution

England, Scotland

Notes

Added by Papp (2000); listed by Kloet and Hincks (1945) (but not by Huddleston 1978), although it is not clear on what basis.

Bracon (Glabrobracon) atrator

Nees, 1834

• longicauda Thomson, 1892

Distribution

England, Scotland, Ireland

Bracon (Glabrobracon) caudatus

Ratzeburg, 1848

Distribution

England

Bracon (Glabrobracon) claripennis

Thomson, 1892

Distribution

England

Notes

added by Papp (2000)

Bracon (Glabrobracon) colpophorus

(Wesmael, 1838)

• Braco colpophorus Wesmael, 1838

• mokrzeckii Niezabitowski, 1927; synonymy by Papp (1997)

Distribution

England

Bracon (Glabrobracon) conjugellae

Bengtsson, 1924

• bengtssoni Fahringer, 1928 preocc.

• minor Fahringer, 1928 preocc.

• nanana Strand, 1928

Distribution

England, Scotland

Notes

NMS, det. Papp, added here

Bracon (Glabrobracon) curticaudis

Szépligeti, 1901

Distribution

Scotland

Notes

Added by Papp (2008a) and removed from synonymy with terebella.

Bracon (Glabrobracon) delibator

Haliday, 1833

• anthracinus Nees, 1834; synonymy by Achterberg (1997)

• breviseta Fahringer, 1935

Distribution

England, Scotland, Ireland

Bracon (Glabrobracon) fuscicoxis

(Wesmael, 1838)

• Braco fuscicoxis Wesmael, 1838

• levicarinatus Niezabitowski, 1910

Distribution

England

Bracon (Glabrobracon) glaphyrus

Marshall, 1897

Distribution

England

Notes

Faure (1924) mentions English material but glaphyrus was omitted by Huddleston (1978).

Bracon (Glabrobracon) guttator

Panzer, 1804

Distribution

England

Notes

Omitted by Huddleston (1978); recorded as British by Fulmek (1968), although we are not sure how reliably; English material in NMS.

Bracon (Glabrobracon) immutator

Nees, 1834

• efoveolatus Thomson, 1892

• hemirugosus Szépligeti, 1901

• flavicoxanus Strand, 1928

• marshalli Fahringer, 1927 preocc.

• romani Fahringer, 1927 preocc.

• uplandiae Strand, 1928

• austriacus Fahringer, 1936

• nigripalpis Telenga, 1936

Distribution

England, Scotland

Bracon (Glabrobracon) instabilis

Marshall, 1897

Distribution

England

Bracon (Glabrobracon) kopelkei

Papp, 2000

Distribution

England

Notes

BMNH, det. Papp, added here

Bracon (Glabrobracon) longulus

Thomson, 1892

Distribution

England, Scotland

Notes

NMS, det. Papp, added here

Bracon (Glabrobracon) marshalli

Szépligeti, 1901

• obscurator misident.

Distribution

England, Scotland

Notes

Added by Papp (2000); listed by Kloet and Hincks (1945) (but not by Huddleston 1978), although it is not clear on what basis.

Bracon (Glabrobracon) minutator

(Fabricius, 1798)

• Ichneumon minutator Fabricius, 1798

• thalassinus Schmiedeknecht, 1897; synonymy by Papp (1999a)

• tener Szépligeti, 1904

• abscissoris Strand, 1928

• flavipalpula Strand, 1928

• flavipalpis Fahringer, 1928 preocc.

• rufiventris Fahringer, 1928 preocc.

• notatus Telenga, 1936

• unicolor Telenga, 1936 preocc.

Distribution

England, Ireland

Notes

Braconminutator auct. is trucidator Marshall (van Achterberg, pers. comm.).

Bracon (Glabrobracon) momphae

Papp, 1999

Distribution

England, Scotland

Notes

added by Papp (1999c)

Bracon (Glabrobracon) nigricollis

(Wesmael, 1838)

• Braco nigricollis Wesmael, 1838

• brunneomaculatus Schütze & Roman, 1931

Distribution

England

Notes

NMS, det. Papp, added here

Bracon (Glabrobracon) obscurator

Nees, 1811

• kotulai Niezabitowski, 1910

• nigripes Fahringer, 1928 preocc.

• rytrensis Fahringer, 1928 preocc.

• rytronis Strand, 1928

• zaleszczykiensis Strand, 1928

Distribution

England, Scotland, Wales

Bracon (Glabrobracon) otiosus

Marshall, 1885

• macrurus Thomson, 1892; synonymy by Papp (1999c)

• explorator Szépligeti, 1904 preocc.; synonymy by Papp (1999c)

• pumilionis Roman, 1928; synonymy by Papp (1999c)

Distribution

England

Notes

Treated as a synonym of bipartitus (=variator) by Belokobylskij et al. (2003).

Bracon (Glabrobracon) pallicarpus

Thomson, 1892

Distribution

England

Notes

added by Papp (2000)

Bracon (Glabrobracon) parvicornis

Thomson, 1892

• carbonarius Szépligeti, 1901 preocc.

• aterrimus Telenga, 1936 preocc.

Distribution

England

Notes

NMS, det. Papp, added here

Bracon (Glabrobracon) parvulus

Wesmael, 1838

• fumipennis Thomson, 1892

• fuscipennis Thomson, 1892 preocc.

• thomsoni Marshall, 1897

Distribution

England, Scotland, Wales

Notes

NMS, det. Papp, added here

Bracon (Glabrobracon) pineti

Thomson, 1892

Distribution

England

Notes

NMS, det. Papp, added here

Bracon (Glabrobracon) praecox

(Wesmael, 1838)

• Braco praecox Wesmael, 1838

• biorrhizae Fahringer, 1928

• elongatus Dutu-Lacatusu, 1956 preocc.

Distribution

Scotland

Notes

NMS, det. Papp, added here

Bracon (Glabrobracon) ratzeburgii

Dalla Torre, 1898

• longicaudis Ratzeburg, 1852 preocc.

Distribution

England

Bracon (Glabrobracon) terebella

(Wesmael, 1838)

• Braco terebella Wesmael, 1838

• breviterebris Fahringer, 1928

• miaricola Strand, 1928

• wesmaeli Fahringer, 1928 preocc.

• unicolor Telenga, 1936 preocc.

Distribution

England

Bracon (Glabrobracon) variator

Nees, 1811

• bipartitus (Wesmael, 1838, Braco)

• maculiger (Wesmael, 1838, Braco)

• collinus Szépligeti, 1901

• breviventris Szépligeti, 1901

• rytrensis Niezabitowski, 1910

• sueciensis Fahringer, 1927

• dimidiatus Fahringer, 1928 preocc.

• nigerrimus Fahringer, 1928

• rytrensis Fahringer, 1928 preocc.

• rytrocola Strand, 1928

• chinensis Fahringer, 1929 preocc.

• caucasicus Telenga, 1936

• collaris Telenga, 1936 preocc.

• meridionalis Telenga, 1936

• ornatulus Telenga, 1936

• rytrensis Telenga, 1936 preocc.

• turcmenus Telenga, 1936

• asiaticus Telenga, 1949 preocc.

Distribution

England, Scotland, Wales, Ireland

Notes

Braco bipartitus is listed as a subspecies of variator in Taxapad (Yu et al. 2012). Listed as a British species by Huddleston (1978) but this actually refers to otiosus.

Bracon (Glabrobracon) xanthogaster

Nees, 1834

• breviseta Hedwig, 1961 preocc.

Distribution

England

Notes

NMS, det. Papp, added here

Habrobracon

Ashmead, 1895

Bracon (Habrobracon) concolorans

Marshall, 1900

• concolor Thomson, 1892 preocc.

• nigricans (Szépligeti, 1901, Habrobracon); synonymy by Papp (2008b)

• mongolicus (Telenga, 1936, Habrobracon)

Distribution

England, Scotland, Ireland

Notes

Added by Papp (2008b) and removed from synonymy with stabilis.

Bracon (Habrobracon) crassicornis

Thomson, 1894

• flavosignatus (Tobias, 1957, Habrobracon); synonymy by Papp (2008b)

Distribution

England, Scotland

Notes

added by Papp (2008b)

Bracon (Habrobracon) hebetor

Say, 1836

• dorsator Say, 1836

• brevicornis (Wesmael, 1838, Braco)

• juglandis Ashmead, 1889

• brunneus (Szépligeti, 1901, Habrobracon)

• vernalis (Szépligeti, 1901, Habrobracon)

• beneficientior (Viereck, 1911, Habrobracon)

• plotnicovi (Bogoljubov, 1914, Habrobracon)

• breviantennatus de Stefani, 1919

• serinopae (Cherian, 1929, Microbracon)

• tortricidarum (Goidanich, 1934, Habrobracon)

• pectinophorae (Watanabe, 1935, Habrobracon)

• asiaticus (Telenga, 1936, Habrobracon)

• flavus (Telenga, 1936, Habrobracon)

• turkestanicus (Telenga, 1936, Habrobracon)

• lozinskii (Bogacev, 1939, Habrobracon)

Distribution

England

Notes

Mainly an indoors species in Britain, attacking pests of stored products; brevicornis been variously treated as a valid species or as a synonym of hebetor, being listed under the latter in Taxapad (Yu et al. 2012) and by Papp (2008b). Both brevicornis and hebetor have been recorded as British.

Bracon (Habrobracon) stabilis

(Wesmael, 1838)

• Braco stabilis Wesmael, 1838

• opacus Stelfox, 1953 preocc.

Distribution

England, Ireland

Lucobracon

Fahringer, 1927

Notes

species of Bracon (Lucobracon) excluded from the British and Irish list:

[strobilorum Ratzeburg, 1848] England and Ireland are listed under the distribution in Shenefelt (1978), and Britain in Fauna Europaea, but we have been unable to trace any British or Irish literature records or specimens. In Belokobylskij et al. (2003) and in Taxapad (Yu et al. 2012) this is listed as a species of Coeloides but Papp (1992) had established that this species had been mis-interpreted and is in fact a species of Bracon (Lucobracon). Swiss specimens identified as strobilorum by Papp in NMS and BMNH belong in Bracon.

Bracon (Lucobracon) brachycerus

Thomson, 1892

• kudsiricus Papp, 1965

Distribution

England

Notes

NMS, det. Papp, added here

Bracon (Lucobracon) crassungula

Thomson, 1892

Distribution

England

Notes

NMS, BMNH, det. Papp, added here

Bracon (Lucobracon) erraticus

(Wesmael, 1838)

• Braco erraticus Wesmael, 1838

• superciliosus (Wesmael, 1838, Braco)

• erythrostictus Marshall, 1885

• exarator Marshall, 1885

• praetermissus Marshall, 1885

• vectensis Marshall, 1885; synonymy by Papp (1999c)

• foveola Thomson, 1892

• aestivalis Szépligeti, 1901; synonymy by Papp (2005b)

• confinis Szépligeti, 1901

• congruus Szépligeti, 1901 preocc.

• similis Szépligeti, 1901

• ventricosus Szépligeti, 1901

• secernendus Schulz, 1906

• lagodechianus Telenga, 1936

• maculatus Telenga, 1936

• planiceps Telenga, 1936

• talitzkii Telenga, 1936

• transcaspicus Telenga, 1936

• hades Papp, 1965

Distribution

England, Scotland, Ireland, Isle of Man

Notes

Although pratermissus was synonymised with Bracon erraticus by Papp (1999c), this has not been followed by Belokobylskij et al. (2003).

Bracon (Lucobracon) flagellaris

Thomson, 1892

• facialis Thomson, 1892 preocc.

• thomsonii Dalla Torre, 1898 preocc.

• dallatorrei Szépligeti, 1901

Distribution

England

Notes

BMNH, det. Papp, added here

Bracon (Lucobracon) grandiceps

Thomson, 1892

• gallicus Thomson, 1892

Distribution

Scotland

Notes

NMS, BMNH, det. Papp, added here

Bracon (Lucobracon) guttiger

(Wesmael, 1838)

• Braco guttiger Wesmael, 1838

• fasciatus Fahringer, 1927

Distribution

England, Scotland

Bracon (Lucobracon) hylobii

Ratzeburg, 1848

• bruchorum Fahringer, 1934

Distribution

England, Scotland

Bracon (Lucobracon) larvicida

(Wesmael, 1838)

• Braco larvicida Wesmael, 1838

• crassiusculus Szépligeti, 1901

• romani Fahringer, 1927 preocc.

• szepligetii Fahringer, 1927 preocc.

• fahringeriensis Strand, 1928

• pseudoromani Strand, 1928

Distribution

England

Bracon (Lucobracon) nigriventris

Wesmael, 1838

• Braco nigriventris Wesmael, 1838

• subornatus Szépligeti, 1901; synonymy by Papp (2008a)

• biroi Fahringer, 1927

• minor Fahringer, 1927

• albanicus Telenga, 1936

• laticeps Telenga, 1936

• lencoranus Telenga, 1936

• persimilis Telenga, 1936

• turolus Papp, 1984

Distribution

England

Notes

NMS, det. Papp, added here

Bracon (Lucobracon) sphaerocephalus

Szépligeti, 1901

• globiceps Szépligeti, 1901; synonymy by Papp (2005b)

Distribution

England, Scotland

Notes

NMS, det. Papp, added here

Bracon (Lucobracon) thuringiacus

Schmiedeknecht, 1897

• schmiedeknechti Fahringer, 1927 preocc.

• blankenburgiae Strand, 1928

Distribution

England

Notes

added by Papp (1999a)

Bracon (Lucobracon) tornator

Marshall, 1885

• aequalis Thomson, 1892

Distribution

England, Scotland, Wales

Notes

some distribution data from Papp (1997)

Bracon (Lucobracon) triangularis

Nees, 1834

Distribution

England, Ireland

Orthobracon

Fahringer, 1927

Notes

species of Bracon (Orthobracon) excluded from the British and Irish list:

[tenuicornis (Wesmael, 1838, Braco)] Probably not a valid British or Irish species now that B. (Lucobracon) erythrostictus has been taken out of synonymy (Papp 1999c).

Bracon (Orthobracon) discoideus

(Wesmael, 1838)

• Braco discoideus Wesmael, 1838

• opionus Fahringer, 1928

• sculpturatus Fahringer, 1928

• sculpturifera Strand, 1928

Distribution

England, Scotland, Ireland

Bracon (Orthobracon) epitriptus

Marshall, 1885

• pallidipes Szépligeti, 1896; synonymy by Papp (2008a)

• melanogaster Szépligeti, 1901

Distribution

England, Scotland, Wales, Ireland

Notes

some distribution data from Papp (1999c)

Bracon (Orthobracon) exhilarator

Nees, 1834

• satanas (Wesmael, 1838, Braco)

• tibialis Zetterstedt, 1838

• striolatus Thomson, 1892

• marshalli Fahringer, 1927 preocc,

• polonicus Fahringer, 1927 preocc.

• polonicella Strand, 1928

• varicoloris Strand, 1928

Distribution

England, Scotland, Wales, Isle of Man

Bracon (Orthobracon) filicornis

Thomson, 1892

Distribution

Scotland

Notes

NMS, det. Papp, added here

Bracon (Orthobracon) laevigatissimus

Dalla Torre, 1898

• laevigatus Ratzeburg, 1852 preocc.

Distribution

England

Notes

Omitted by Huddleston (1978); listed as British by various authors dating back to Marshall (1885).

Bracon (Orthobracon) mediator

Nees, 1834

Distribution

England, Scotland, Ireland

Bracon (Orthobracon) ochropus

Nees, 1834

• marshalli Fahringer, 1927 preocc.

• neesi Fahringer, 1927

• ochropodis Strand, 1928

Distribution

England, Scotland

Notes

NMS, det. Papp, added here

Bracon (Orthobracon) orbus

Papp, 1981

Distribution

England

Notes

NMS, det. Papp, added here

Bracon (Orthobracon) picticornis

Wesmael, 1838

• gallarum Ratzeburg, 1852

• versicolor Szépligeti, 1901

• ratzeburgensis Strand, 1928

• laevigatus Fahringer, 1927 preocc.

Distribution

England, Ireland

Bracon (Orthobracon) procerus

Papp, 1965

Distribution

England

Notes

NMS, det. Papp, added here

Bracon (Orthobracon) pulcher

Bengtsson, 1924

• bengtssoni Fahringer, 1927

Distribution

England

Notes

NMS, det. Papp, added here

Bracon (Orthobracon) roberti

(Wesmael, 1838)

• Braco roberti Wesmael, 1838

Distribution

England, Ireland

Bracon (Orthobracon) romani

Fahringer, 1927

Distribution

England, Scotland, Wales, Ireland

Notes

NMS, det. Papp, added here

Bracon (Orthobracon) subcylindricus

(Wesmael, 1838)

• Braco subcylindricus Wesmael, 1838

• niger (Vojnovskaja-Krieger, 1929, Baryproctus)

Distribution

England

Bracon (Orthobracon) subsinuatus

Szépligeti, 1901

Distribution

England, Scotland, Ireland

Notes

Added by Papp (2008a). Removed from synonymy with epitriptus by Papp (2008a), who treats both species as belonging to the subgenus Glabrobracon, contrary to their placement in Taxapad (Yu et al. 2012).

Bracon (Orthobracon) titubans

(Wesmael, 1838)

• Braco titubans Wesmael, 1838

• tarsator Thomson, 1892

• terebrator Szépligeti, 1901

Distribution

England

Notes

NMS, BMNH, det. Papp, added here

Bracon (Orthobracon) virgatus

Marshall, 1897

• lineifer van Achterberg, 1988; synonymy by Papp (1999c)

Distribution

England, Scotland

Notes

some distribution data from Shaw and Bailey (1991)

Osculobracon

Papp, 2008

Bracon (Osculobracon) osculator

Nees, 1811

• bisignatus (Wesmael, 1838, Braco)

• degenerator Marshall, 1885

• minutus Szépligeti, 1901

• temporalis Telenga, 1936

• venustus Telenga, 1936

Distribution

England, Scotland, Wales, Ireland

Pigeria

van Achterberg, 1985

Notes

Described as a separate genus but Quicke and Sharkey (1989) suggested that Pigeria could be treated as a subgenus of Bracon, which was followed by Papp (1998).

Bracon (Pigeria) piger

(Wesmael, 1838)

• Braco piger Wesmael, 1838

• rotundatus Szépligeti, 1901; synonymy by Papp (2008a)

• rotundulus Szépligeti, 1904

Distribution

England, Ireland

Notes

Bracon semiluteus Walker, 1874 is not a synonym of piger (contra Morley (1913) and subsequent authors) but is a junior synonym of another species of Bracon (Papp, in prep.).

Bracon (Pigeria) wolschrijni

(van Achterberg, 1985)

• Pigeria wolschrijni van Achterberg, 1985

Distribution

Wales

Notes

NMS, det. van Achterberg, added here

Rostrobracon

Tobias, 1957

Notes

species of Bracon (Rostrobracon) excluded from the British and Irish list:

[urinator (Fabricius, 1798, Ichneumon); syn. cuspidator (Rossi, 1792, Ichneumon); comptus Marshall, 1897] Marshall (1900) mentions an English specimen but this seems unlikely in view of its present southern European range.

Coeloidini

Tobias, 1957

Coeloides

Wesmael, 1838

• SYNTOMOMELUS Kokujev, 1902

• HABROBRACONIDEA Viereck, 1912

• COELOIDINA Viereck, 1921

• CEROBRACON Viereck, 1926

Notes

Distribution data from Shaw (2000b).

Coeloides abdominalis

(Zetterstedt, 1838)

• Bracon abdominalis Zetterstedt, 1838

Distribution

England

Coeloides filiformis

Ratzeburg, 1852

• melanurus Ivanov, 1896

Distribution

England

Notes

added by Shaw (2000b)

Coeloides melanostigma

Strand, 1918

• sordidator misident.

• stigmaticus Hellén, 1928

Distribution

England

Notes

Added by Shaw (2000b); according to Haeselbarth (in Belokobylskij et al. 2003), the name sordidator (Ratzeburg, 1844, Bracon) probably does not belong in Coeloides, so the species usually referred to as sordidator takes the name melanostigma.

Coeloides melanotus

Wesmael, 1838

• flavus Ivanov, 1896

• maculatus Ivanov, 1896

Distribution

England, Wales

Coeloides scolyticida

Wesmael, 1838

• initiatellus (Ratzeburg, 1848, Bracon)

Distribution

England

Cardiochilinae

Cardiochilinae

Ashmead, 1900

Notes

no British or Irish species

Cardiochiles

Nees, 1819

• Ditherus Cameron, 1902

Notes

species of Cardiochiles excluded from the British and Irish list:

[saltator (Fabricius, 1781, Ichneumon); syn. brachialis Rondani, 1877; katkowi Kokujev, 1895; fumipennis Szépligeti, 1901; sibiricus Telenga, 1955] not British, recorded in error (Shaw and Huddleston 1991)

Cenocoeliinae

Cenocoeliinae

Szépligeti, 1901

Notes

Distribution data from Shaw (1999).

Cenocoelius

Haliday, 1840

• LACCOPHRYS Förster, 1863

• PROMACHUS Cresson, 1887 preocc.

• CAENOCOELIUS Marshall, 1894

• POSTPROMACHUS Maes, 1999

Cenocoelius aartseni

(van Achterberg, 1994)

• Promachus aartseni van Achterberg, 1994

Distribution

England

Notes

added by Shaw (1999)

Cenocoelius analis

(Nees, 1834)

• Bracon analis Nees, 1834

• flavifrons Haliday, 1840

• cephalotes (Ratzeburg, 1848, Opius) preocc.

• magdalini (Förster, 1863, Laccophrys)

• hungaricus Kiss, 1927

Distribution

England, Ireland

Lestricus

Reinhard, 1865

Lestricus secalis

(Linnaeus, 1758)

• Ichneumon secalis Linnaeus, 1758

• agricolator (Linnaeus, 1767, Ichneumon)

• rubriceps (Ratzeburg, 1844, Alysia)

• femorator (Tobias, 1973, Cenocoelius)

Distribution

Scotland

Notes

NMS, det. Shaw, added here; removed from the British list by Shaw (1999) but recently found in Scotland.

Charmontinae

Charmontinae

van Achterberg, 1979

Charmon

Haliday, 1833

• PROVANCHERIA Ashmead, 1900

• CYCLOCORMUS Cameron, 1911

• EUBADIZON misident.

Charmon cruentatus

Haliday, 1833

• pectoralis (Nees, 1834, Eubadizon)

• pleuralis (Cresson, 1872, Eubadizon)

• luteus (Cameron, 1911, Cyclocormus)

• Charmon cruentatus ?brevicauda (Hellén, 1958, Eubadizon)

Distribution

England, Scotland, Wales, Ireland, Isle of Man

Notes

Although Achterberg (1979) treated brevicauda as a ‘form’ of cruentatus, the possibility of a separate species requires investigation.

Charmon extensor

(Linnaeus, 1758)

• Ichneumon extensor Linnaeus, 1758

• gracilis (Provancher, 1880, Eubadizon)

• hungaricus (Kiss, 1927, Calyptus)

• striatus (Shestakov, 1940, Eubadizon)

Distribution

England, Scotland, Ireland

Cheloninae

Cheloninae

Förster, 1863

• ADELIINAE Viereck, 1918

• ACAELIINAE Viereck, 1918

• ACOELIINAE Viereck, 1918

Notes

The genus Adelius and related extralimital genera have usually been treated as comprising a separate subfamily, Adeliinae. Recent phylogenetic studies (Belshaw et al. 2000, Belshaw and Quicke 2002) have placed the adeliines within the Cheloninae, as sister taxon to Phanerotoma.

Chelonini

Förster, 1863

Ascogaster

Wesmael, 1835

• CASCOGASTER Baker, 1926

• LEPTODREPANA Shaw, 1983; synonymy by Achterberg (1990)

Notes

Distribution and synonymic data taken from Huddleston (1984).

Species of Ascogster excluded from the British and Irish list:

[bicarinata (Herrich-Schäffer, 1838, Chelonus); syn. mlokossewitschi Kokujev, 1895; syn. rufiventris Telenga, 1941; preocc.] Included by Huddleston (1978) but no British specimens were seen by Huddleston (1984).

[similis (Nees, 1816, Chelonus)] The type is lost and the species is unplaceable Huddleston (1984).

Ascogaster abdominator

(Dahlbom, 1833)

• Chelonus abdominator Dahlbom, 1833

• instabilis Wesmael, 1835

• fulviventris Curtis, 1837

• femoralis (Herrich-Schäffer, 1838, Chelonus)

• rufiventris (Herrich-Schäffer, 1838, Chelonus)

• pallida Ruthe, 1855

Distribution

England, Scotland, Ireland

Ascogaster albitarsus

Reinhard, 1867

• similis (Herrich-Schäffer, 1838, Chelonus)

• leptopus Thomson, 1874

Distribution

Scotland, Ireland

Notes

added by Huddleston (1984)

Ascogaster annularis

(Nees, 1816)

• Sigalphus annularis Nees, 1816

Distribution

England

Ascogaster armata

Wesmael, 1835

• pulchella (Curtis, 1829, Chelonus) nom. nud.

• esenbeckii Curtis, 1837

• luteicornis (Herrich-Schäffer, 1838, Chelonus)

Distribution

England, Wales

Ascogaster bidentula

Wesmael, 1835

• scabriuscula (Zetterstedt, 1838, Sigalphus)

• multiarticulata (Ratzeburg, 1852, Chelonus)

• gibbiscuta Thomson, 1874

• fuscipennis Thomson, 1892

• atamiensis Ashmead, 1906

Distribution

England, Scotland, Wales, Ireland

Notes

added by Huddleston (1984)

Ascogaster brevicornis

Wesmael, 1835

• monilicornis (Herrich-Schäffer, 1838, Chelonus)

Distribution

England, Ireland

Notes

added by Huddleston (1984)

Ascogaster canifrons

Wesmael, 1835

• graniger Thomson, 1892

• zernyana Fahringer, 1925

Distribution

England, Ireland

Ascogaster consobrina

Curtis, 1837

Distribution

England, Scotland, Ireland

Ascogaster dentifer

Tobias, 1976

Distribution

England

Notes

added by Huddleston (1984)

Ascogaster dispar

Fahringer, 1934

• spinifer Tobias, 1964

• kozlovi Tobias, 1972

Distribution

England

Notes

added by Huddleston (1984)

Ascogaster gonocephala

Wesmael, 1835

Distribution

England

Notes

added by Huddleston (1984)

Ascogaster grahami

Huddleston, 1984

Distribution

England

Notes

added by Huddleston (1984)

Ascogaster klugii

(Nees, 1816)

• Sigalphus klugii Nees, 1816

• ruficeps Wesmael, 1835

• neesii Reinhard, 1867

Distribution

England

Notes

added by Huddleston (1984)

Ascogaster quadridentata

Wesmael, 1835

• pallidicornis Curtis, 1837

• impressa (Herrich-Schäffer, 1838, Chelonus)

• quadridens (Herrich-Schäffer, 1838, Chelonus)

• cynipum Thomson, 1892

• nigricornis Thomson, 1892

• egregia Kokujev, 1895

• nigrator (Szépligeti, 1896, Chelonus)

• carpocapsae (Viereck, 1909, Chelonus)

• epinotiae Watanabe, 1937

Distribution

England

Ascogaster rufidens

Wesmael, 1835

• rufipes (Herrich-Schäffer, 1838, Chelonus) preocc.

• laevigator (Ratzeburg, 1852, Chelonus)

Distribution

England, Ireland

Ascogaster rufipes

(Latreille, 1809)

• Sigalphus rufipes Latreille, 1809

• elegans (Nees, 1816, Sigalphus)

• fasciata (Dahlbom, 1833, Chelonus)

• pallipes (Herrich-Schäffer, 1838, Chelonus)

• rubripes (Lucas, 1849, Chelonus)

• rugosula (Goureau, 1861, Chelonus)

• ratzeburgii Marshall, 1885

• arisanica Sonan, 1932

• nigribasis Fahringer, 1934

• soror Telenga, 1941

Distribution

England, Ireland

Ascogaster varipes

Wesmael, 1835

• atriceps (Ratzeburg, 1844, Chelonus)

• tersa Reinhard, 1867

• cavifrons Thomson, 1874

• sternalis Thomson, 1874

• catula (Marshall, 1885, Chelonus); synonymy by Papp (1996c)

• jaroslawensis Kokujev, 1895

Distribution

England, Scotland, Ireland

Chelonus

Jurine, 1801

Chelonus

Jurine, 1801

• Anomala von Block, 1799 nom. ob.

• DAVISANIA La Munyon, 1877

• ARICHELONUS Viereck, 1913

• MEGACHELONUS Baker, 1926

Chelonus (Chelonus) acuminatus

Herrich-Schäffer, 1838

Distribution

England

Notes

NMS, det. Huddleston, added here

Chelonus (Chelonus) annulatus

(Nees, 1816)

• Sigalphus annulatus Nees, 1816

• maculatus Szépligeti, 1896

Distribution

England, Scotland, Ireland

Chelonus (Chelonus) asiaticus

Telenga, 1941

Distribution

England, Ireland

Notes

BMNH, det. Lozan, added here. The name is a junior homonym of Chelonus asiaticus Fahringer, 1932.

Chelonus (Chelonus) canescens

Wesmael, 1835

Distribution

England

Chelonus (Chelonus) carbonator

Marshall, 1885

• asiaticus Fahringer, 1932

Distribution

England

Chelonus (Chelonus) corvulus

Marshall, 1885

• suturatus Szépligeti, 1898

Distribution

England, Scotland

Chelonus (Chelonus) cylindrus

(Klug, 1816)

• Sigalphus cylindrus Klug, 1816

• variabilis Herrich-Schäffer, 1838

• macrocerus Thomson, 1874

• speculator Marshall, 1885

• ebeninus Fahringer, 1934

Distribution

England

Notes

Chelonus speculator was listed as a synonym of oculator by Huddleston (1978).

Chelonus (Chelonus) decorus

Marshall, 1885

• clavipes Curtis, 1837 nom. nud.

• clavipes Fahringer, 1934

• szepligetii Fahringer, 1934

Distribution

England

Chelonus (Chelonus) inanitus

(Linnaeus, 1767)

• Cynips inanita Linnaeus, 1767

• binarius (Fourcroy, 1785, Ichneumon)

• atomos (Rossi, 1790, Ichneumon)

Distribution

England, Ireland

Chelonus (Chelonus) obscuratus

Herrich-Schäffer, 1838

• intermedius Thomson, 1874

Distribution

England

Notes

NMS, det. Huddleston, added here

Chelonus (Chelonus) oculator

(Fabricius, 1785)

• Ichneumon oculator Fabricius, 1785

• integer (von Block, 1799, Anomala)

• mutabilis (Nees, 1816, Sigalphus)

• oculatus (Nees, 1816, Sigalphus)

Distribution

England, Scotland

Chelonus (Chelonus) pusio

Marshall, 1885

Distribution

England

Chelonus (Chelonus) scabrator

(Fabricius, 1793)

• Ichneumon scabrator Fabricius, 1793

• scaber (Nees, 1816, Sigalphus)

• buccatus Thomson, 1874

Distribution

England

Chelonus (Chelonus) submuticus

Wesmael, 1835

• luteipes Thomson, 1874; synonymy by Papp (1995b)

Distribution

England

Chelonus (Chelonus) wesmaelii

Curtis, 1837

• zimini Tobias, 1972

Distribution

England

Microchelonus

Szépligeti, 1908

• CHELONELLA Szépligeti, 1908

• NEOCHELONELLA Hincks, 1943

Notes

Treated as a valid genus by Papp (1996c) and Belokobylskij et al. (2003), sometimes treated as a straight synonym of Chelonus (Achterberg and Polaszek 1996), here treated as a subgenus of Chelonus, following Taxapad (Yu et al. 2012).

Chelonus (Microchelonus) atripes

Thomson, 1874

• cunctator (Papp, 1971, Microchelonus)

• kamtshaticus (Tobias, 1986, Microchelonus)

Distribution

England

Notes

NMS, BMNH, det. Huddleston and Lozan, added here

Chelonus (Microchelonus) basalis

Curtis, 1837

Distribution

England, Scotland

Chelonus (Microchelonus) binus

(Tobias, 1995)

• Microchelonus binus Tobias, 1995

Distribution

England

Notes

BMNH, det. Lozan, added here

Chelonus (Microchelonus) contractus

(Nees, 1816)

• Sigalphus contractus Nees, 1816

• compressiscapus Szépligeti, 1898; synonymy by Papp (1996c)

Distribution

England, Scotland, Wales, Ireland

Notes

some distribution data from Tobias and Shaw (2005)

Chelonus (Microchelonus) depressus

Thomson, 1874

Distribution

England, Scotland, Wales

Notes

NMS, BMNH, det. Huddleston and Lozan, added here

Chelonus (Microchelonus) exilis

Marshall, 1885

• excavatus Tobias, 1972; synonymy by Papp (1995b)

Distribution

England

Chelonus (Microchelonus) fenestratus

(Nees, 1816)

• Sigalphus fenestratus Nees, 1816

• dispar Marshall, 1885

• bimaculatus Ivanov, 1899 preocc.

Distribution

England

Notes

Synonymised under contractus by Papp (1995b) but treated as a separate species by Tobias and Lozan (2003) and listed as such in Taxapad (Yu et al. 2012).

Chelonus (Microchelonus) fumipennis

(Tobias, 1986)

• Microchelonus fumipennis Tobias, 1986

Notes

BMNH, det. Lozan, added here; specimen lacking locality data, Billups coll.

Chelonus (Microchelonus) latrunculus

Marshall, 1885

• Chelonus (Microchelonus) latrunculus ?parcicornis Herrich-Schäffer, 1838

• thomsonii Dalla Torre, 1898

• polonicus Fahringer, 1934

• rectus (Papp, 1971, Microchelonus)

Distribution

England, Scotland

Notes

Although parcicornis would have priority, Papp (1995b) states that the type is lost and its identity unverifiable.

Chelonus (Microchelonus) lugubris

Wesmael, 1835

Distribution

England, Ireland

Notes

NMS, BMNH, det. Huddleston and Lozan, added here

Chelonus (Microchelonus) microphtalmus

Wesmael, 1838

• dilatus Papp, 1971

Distribution

England

Notes

NMS, det. Huddleston, added here

Chelonus (Microchelonus) miscellae

(Tobias & Shaw, 2005)

• Microchelonus miscellae Tobias & Shaw, 2005

Distribution

England

Notes

added by Tobias and Shaw (2005)

Chelonus (Microchelonus) retusus

(Nees, 1816)

• Sigalphus retusus Nees, 1816

• emarginatus Herrich-Schäffer, 1838

• subemarginatus Herrich-Schäffer, 1838

• caudatus Thomson, 1874

• pamiricus Vojnovskaja-Krieger, 1931

Distribution

England

Notes

BMNH, det. Lozan, added here

Chelonus (Microchelonus) risorius

Reinhard, 1867

• fissus Szépligeti, 1900 preocc.

• fissuralis (Tobias, 1964, Neochelonella)

• magnifissus (Tobias, 1986, Microchelonus)

Distribution

England

Chelonus (Microchelonus) sulcatus

Jurine, 1807

Distribution

England

Parachelonus

Tobias, 1995

Chelonus (Parachelonus) gravenhorstii

(Nees, 1816)

• Sigalphus gravenhorstii Nees, 1816

• maculator Dahlbom, 1833

• eurytheca Wesmael, 1838

• adjaricus Tobias, 1976

• tricolor Tobias, 1976

Distribution

England

Chelonus (Parachelonus) pellucens

(Nees, 1816)

• Sigalphus pellucens Nees, 1816

• nitens Reinhard, 1867

• alboannulatus Szépligeti, 1896

• pulchricornis Szépligeti, 1898

• varimaculatus Tobias, 1986

• austriacus Fahringer, 1934

Distribution

England

Notes

added by Papp (2004c)

Stylochelonus

Hellén, 1958

Chelonus (Stylochelonus) pedator

Dahlbom, 1833

• secutor (Marshall, 1885, Chelonus)

Distribution

England, Scotland

Chelonus (Stylochelonus) pusillus

(Szépligeti, 1908)

• Stylochelonus pusillus Szépligeti, 1908

• furtivus (Tobias, 1986, Microchelonus)

• tuberculiventris (Tobias, 1986, Microchelonus)

Distribution

England

Notes

BMNH, det. Lozan, added here

Phanerotomini

Baker, 1926

Adelius

Haliday, 1833

• ACAELIUS Haliday, 1834

• ACOELIUS Haliday, 1835

• PLEIOMERUS Wesmael, 1837

• ANOMOPTERUS Rohwer, 1914

• MYRIOLA Shestakov, 1932

Adelius erythronotus

(Förster, 1851)

• Acoelius erythronotus Förster, 1851

• pyrrhia (Beirne, 1945, Acoelius)

• flavus (Tobias, 1966, Acoelius)

Distribution

England, Ireland

Notes

Adelius pyrrhia was listed as a valid species by O'Connor et al. (1999) but as a synonym of erythronotus by Belokobylskij et al. (2003).

Adelius germanus

(Haliday, 1834)

• Acaelius germanus Haliday, 1834

Distribution

England, Ireland

Adelius subfasciatus

Haliday, 1833

• minutissimus (Zetterstedt, 1840, Bracon)

• parvulus (Förster, 1851, Acoelius)

Distribution

England, Wales, Ireland

Notes

some distribution data from Shaw and Askew (1976)

Adelius viator

(Förster, 1851)

• Acoelius viator Förster, 1851

Distribution

England

Phanerotoma

Wesmael, 1838

Phanerotoma

Wesmael, 1838

• PHANEROGASTER Wesmael, 1838 unavailable

• SULYDUS Du Buysson, 1897

• ICHNEUTIPTERUS Vachal, 1907; synonymy by Achterberg (1990)

• NEOPHANEROTOMA Szépligeti, 1908

• NEOACAMPSIS Szépligeti, 1914; synonymy by Achterberg (1990)

• PHANEROTOMINA Shestakov, 1930

Notes

species of Phanerotoma (Phanerotoma) excluded from the British and Irish list:

[planifrons (Nees, 1816, Sigalphus); syn. blanda Fahringer, 1934; bicolor Snoflák, 1958 preocc.; asini Llopis Minguez, 1968; snoflaki Shenefelt, 1973] Prior to van Achterberg’s (Achterberg 1990) revision, dentata had requently been misidentified as planifrons. All of the British specimens identified as planifrons in the BMNH are in fact dentata.

Phanerotoma (Phanerotoma) acuminata

Szépligeti, 1908

Distribution

England

Notes

added by Achterberg (1990)

Phanerotoma (Phanerotoma) dentata

(Panzer, 1805)

• Chelonus dentatus Panzer, 1805

• dentator (Nees, 1816, Sigalphus)

• rendilea Fahringer, 1934

• minor Šnoflák, 1951; synonymy by Achterberg (1990)

Distribution

England

Notes

Huddleston (1978) listed Sigalphus rufescens Latreille, 1809 as a junior synonym of dentata but this is now considered to be a separate species, not occurring in Britain or Ireland (Achterberg 1990), with historical records being misidentifications.

Phanerotoma (Phanerotoma) leucobasis

Kriechbaumer, 1894

• ocularis Kohl, 1906

• ornatulopsis de Saeger, 1942

• desertorum Hedwig, 1957; synonymy by Achterberg (1990)

• flavitestacea Fischer, 1939; synonymy by Achterberg (1990)

• caboverdensis Hedqvist,1965

Distribution

England

Notes

Added by van Achterberg (1990). Phanerotoma rjabovi Vojnovskaja-Krieger, 1929 and media Shestakov, 1930, synonymised under leucobasis by Achterberg (1990), are now considered to be junior synonyms of fracta Kokujev, 1903 (Belokobylskij 2000b).

Bracotritoma

Csiki, 1909

• TRITOMA Szépligeti, 1908 preocc.; synonymy by Achterberg (1990)

• SZEPLIGETIA Schulz, 1911; synonymy by Achterberg (1990)

• TRITOMIOS Strand, 1921; synonymy by Achterberg (1990)

• UNICA Šnoflák, 1951

Phanerotoma (Bracotritoma) bilinea

Lyle, 1924

• gregori Šnoflák, 1951

Distribution

England

Phanerotoma (Bracotritoma) tritoma

(Marshall, 1898)

• Chelonus tritomus Marshall, 1898

• antennalis Šnoflák, 1951; synonymy by Achterberg (1990)

Distribution

England, Wales, Ireland

Doryctinae

Doryctinae

Förster, 1863

Notes

Tribal classification follows Belokobylskij (1992), but note that recent work suggests that none of the major doryctine tribes are actually monophyletic (Belokobylskij et al. 2004). Generic synonyms for the most part follow Belokobylskij et al. (2004).

Doryctini

Förster, 1863

Dendrosoter

Wesmael, 1838

• EURYBOLUS Ratzeburg, 1848

• CAENOPACHYS Förster, 1863

Notes

Although Mancini et al. (2003) argued that Caenopachys should be treated as a separate genus, Gebiola et al. (2015) presented molecular phylogenetic evidence that that ‘Caenopachys’ species (including hartigii in Britain) are nested within Dendrosoter.

Dendrosoter hartigii

(Ratzeburg, 1848)

• Bracon hartigii Ratzeburg, 1848

• flaviventris Förster, 1878

• caenopachoides Ruschka, 1925; synonymy by Gebiola et al. (2015)

• hartigi misspelling

Distribution

England

Dendrosoter middendorffi

(Ratzeburg, 1848)

• Bracon middendorffi Ratzeburg, 1848

• schimitscheki Fahringer, 1941

Distribution

England

Notes

NMS, det. Shaw, added in Fauna Europaea

Dendrosoter protuberans

(Nees, 1834)

• Bracon protuberans Nees, 1834

• insignis Förster, 1878

Distribution

England

Doryctes

Haliday, 1836

• ISCHIOGONUS Wesmael, 1838

• Neodoryctes Szépligeti, 1914

• UDAMOLCUS Enderlein, 1920

• PRISTODORYCTES Kieffer, 1921

• Paradoryctes Granger, 1949

• PLYCTES Fischer, 1970

Doryctes heydenii

Reinhard, 1865

Distribution

England

Doryctes leucogaster

(Nees, 1834)

• Bracon leucogaster Nees, 1834

Notes

#Probably occurred only in imported timber. Its British status appears to depend solely on specimens reared in July 1908 from Austrian oak imported to a timber yard near Millwall docks (Elliot and Morley 1911).

Doryctes obliteratus

(Nees, 1834)

• Bracon obliteratus Nees, 1834

• mutillator misident.

• tabidus (Haliday, 1836, Rogas)

• brachyurus Marshall, 1888

• strigatus Kokujev, 1900

• petrovskii Kokujev, 1902

Distribution

England

Notes

Regarded as a synonym of striatellus by Belokobylskij et al. (2003).

Doryctes pomarius

Reinhard, 1865

• schimitscheki Fahringer, 1931

Distribution

England

Doryctes rossicus

Telenga, 1941

Distribution

England

Notes

NMS, det. Shaw, added in Fauna Europaea

Doryctes striatellus

(Nees, 1834)

• Bracon striatellus Nees, 1834

• maculipes Curtis, 1837 nom. nud.

• disparator (Ratzeburg, 1844, Bracon)

• rex Marshall, 1897

• striatelloides Strand, 1918

• yogoi Watanabe, 1954

• ambigua Kokujev, 1900

• notatus Kokujev, 1900

Distribution

England

Doryctes undulatus

(Ratzeburg, 1852)

• Bracon undulatus Ratzeburg, 1852

Distribution

England

Gildoria

Hedqvist, 1974

Notes

Removed from synonymy with Dendrosotinus Telenga, 1941 by Achterberg (2003b).

Gildoria similis

(Bouček, 1955)

• Dendrosotinus similis Bouček, 1955

Distribution

England

Notes

added by Shaw (1998)

Ontsira

Cameron, 1900

• DORYCTODES Hellén, 1927

Notes

Zaldívar-Riverón et al. (2008a) did not recover Ontsira as monophyletic and consequently some authors (e.g. Achterberg 2014) prefer to treat these species as belonging to Doryctodes.

Ontsira antica

(Wollaston, 1858)

• Clinocentrus anticus Wollaston, 1858

• gallica (Reinhard, 1865, Doryctes)

• truncorum (Goureau, 1866, Bracon)

• incerta (Ashmead, 1888, Doryctes)

• caudalis (Hellén, 1957, Oncophanes)

Distribution

Scotland

Ontsira ignea

(Ratzeburg, 1852)

• Bracon igneus Ratzeburg, 1852

Distribution

England

Notes

NMS, det. Shaw and van Achterberg, added here

Ontsira imperator

(Haliday, 1836)

• Rogas imperator Haliday, 1836

• zonata (Wesmael, 1838, Ischiogonus)

• praecisa (Ratzeburg, 1852, Bracon)

• cingulata (Provancher, 1880, Syngaster)

• dubia (Kokujev, 1900, Doryctes)

• iranica (Telenga, 1941, Doryctodes)

• niger (Hedwig, 1957, Coeloides)

Distribution

England, Ireland

Rhaconotus

Ruthe, 1854

• HEDYSOMUS Förster, 1863

• HORMIOPTERUS Giraud, 1869

• RHADINOGASTER Szépligeti, 1908

• EURYPHRYMNUS Cameron, 1910

• RHACONOTINUS Hedqvist, 1965

Rhaconotus aciculatus

Ruthe, 1854

• cerdai Docavo Alberti, 1960

• major Tobias, 1964

Distribution

England

Notes

added by Shaw (1998)

Wachsmannia

Szépligeti, 1900

Notes

Belokobylskij (1992) synonymised Wachsmannia under Ontsira but van Achterberg (1995) disagreed and synonymised Wachsmannia under Hypodoryctes Kokujev, 1900. Van Achterberg (pers. comm. and in Fauna Europaea) now regards Wachsmannia as a separate genus again.

Wachsmannia spathiiformis

(Ratzeburg, 1848)

• Bracon spathiiformis Ratzeburg, 1848

• maculipennis Szépligeti, 1900; synonymy by Achterberg (1995)

• obliteratus misident.

Distribution

England

Ecphylini

Hellén, 1957

Ecphylus

Förster, 1863

• TERENUSA Marshall, 1885

• PARAECPHYLUS Ashmead, 1900

• SACTOPUS Ashmead, 1900

Notes

Other species are treated as synonyms of silesiacus by Belokobylskij et al. (2003) but van Achterberg (in Belokobylskij et al. 2003) states that there is biological evidence for several species.

Ecphylus eccoptogastri

(Ratzeburg, 1848)

• Bracon eccoptogastri Ratzeburg, 1848

Ecphylus hylesini

(Ratzeburg, 1852)

• Bracon hylesini Ratzeburg, 1852

Distribution

England

Notes

NMS, det. Shaw and van Achterberg, added in Fauna Europaea

Ecphylus pinicola

Hedqvist, 1967

Distribution

England, Scotland

Notes

NMS, det. Shaw and van Achterberg, added in Fauna Europaea

Ecphylus silesiacus

(Ratzeburg, 1848)

• Bracon silesiacus Ratzeburg, 1848

• minutissimus (Ratzeburg, 1848, Bracon) preocc.

Distribution

England

Notes

NMS, det. Shaw and van Achterberg, added in Fauna Europaea; Shaw (1998) recorded silesiacus as British but was using the name in the broader sense (i.e. =eccoptogastri, =hylesini, =pinicola). Listed in Huddleston (1978) as eccoptogastri.

Hecabolini

Förster, 1863

Hecabolus

Curtis, 1834

• ANISOPELMA Wesmael, 1838

Hecabolus sulcatus

Curtis, 1834

• belgicus (Wesmael, 1838, Anisopelma)

Distribution

England, Wales

Monolexis

Förster, 1863

Notes

Species of Monolexis excluded from the British and Irish list:

[fuscicornis Förster, 1863; syn. lycti (Cresson, 1880, Anisopelma); minimus (Cresson, 1880, Anisopelma); utilis (Cresson, 1880, Anisopelma); doderoi (Mantero, 1910, Hecabolus); lavagnei Picard, 1913; atis Nixon, 1943] Seems to have been included on the British list only on the basis of specimens emerging from Oak (Quercus) timber imported from the USA (e.g. Laing 1928).

Heterospilus

Haliday, 1836

• SYNODUS misident.

• TELEBOLUS Marshall, 1888

• KAREBA Cameron, 1905

• ANOCATOSTIGMA Enderlein, 1920

• HARPAGOLACCUS Enderlein, 1920

• LITUANIA Jakimavičius, 1968

Notes

species of Heterospilus excluded from the British and Irish list:

[caesus (Nees, 1834, Bracon) misident.] Excluded by Shaw and Huddleston (1991), see also Shaw (1997).

Heterospilus ater

Fischer, 1960

Distribution

England

Notes

added by Shaw (1998)

Heterospilus fuscexilis

Shaw, 1997

Distribution

England

Notes

added by Shaw (1997)

Spathiini

Förster, 1863

Spathius

Nees, 1818

• STENOPHASMUS Smith, 1859

• EUSPATHIUS Förster, 1863

• PSEUDOSPATHIUS Szépligeti, 1902

• RHACOSPATHIUS Cameron, 1905

Spathius brevicaudis

Ratzeburg, 1844

Distribution

England, Wales

Notes

NMS, det. Shaw, added in Fauna Europaea

Spathius exarator

(Linnaeus, 1758)

• Ichneumon exarator Linnaeus, 1758

• formicatus (Linnaeus, 1767, Ichneumon)

• mutillarius (Fabricius, 1775, Ichneumon)

• mystacatus (Schrank, 1781, Ichneumon)

• affinis (Fabricius, 1793, Ichneumon)

• immaturus (Gravenhorst, 1807, Ichneumon)

• clavatus (Panzer, 1809, Cryptus)

• affinator (Thunberg, 1824, Ichneumon)

• attenuator (Thunberg, 1824, Ichneumon)

• formicator (Thunberg, 1824, Ichneumon) preocc.

• mutillator (Thunberg, 1824, Ichneumon)

• exannulatus Ratzeburg, 1848

• ferrugatus Goureau, 1866

• strandi Fahringer, 1930

• breviterebrantus Dutu-Lacatusu, 1956

Distribution

England, Scotland, Wales, Ireland

Spathius pedestris

Wesmael, 1838

• apterus Wollaston, 1858

• maderi Fahringer, 1930

• hirtus Hedqvist, 1976

Distribution

England

Notes

Recorded as new to Britain by Perkins and Nixon (1939). Huddleston (1978) listed this as a species in need of confirmation, although, being wingless, it is straightforward to identify. There are additional British specimens in BMNH, World Museum Liverpool, Manchester Museum and Leeds City Museum.

Spathius phymatodis

Fischer, 1966

Distribution

England

Notes

NMS, det. Shaw, added in Fauna Europaea

Spathius rubidus

(Rossius, 1794)

• Ichneumon rubidus Rossius, 1794

• umbrator (Thunberg, 1824, Ichneumon) preocc,

• rugosus Ratzeburg, 1848

• sculpturatus Hellén, 1927

• depressus Hedqvist, 1976

• bimaculatus Telenga, 1941

Distribution

England, Scotland, Ireland

Spathius umbratus

(Fabricius, 1798)

• Ichneumon umbratus Fabricius, 1798

• erythrocephalus Wesmael, 1838

• curvicaudis Ratzeburg, 1844

Distribution

England

Notes

Spathius curvicaudis was recorded as new to Britain by Shaw (1988a) but synonymised under erythrocephalus by Belokobylskij and Samartsev (2014) and Achterberg (2014), who also synonymyised erythrocephalus under umbratus.

Euphorinae

Euphorinae

Förster, 1863

Notes

The tribal and generic classification follows Stigenberg et al. (2015).

Centistini

Čapek, 1970

Allurus

Förster, 1863

Allurus lituratus

(Haliday, 1835)

• Leiophron lituratus Haliday, 1835

Distribution

Ireland

Notes

Synonymised under muricatus by Achterberg (1985), later removed from synonymy by Achterberg (1997), with a key to the species.

Allurus muricatus

(Haliday, 1833)

• Ancylus muricatus Haliday, 1833

• armatus (Wesmael, 1835, Leiophron)

• niger (Lyle, 1926, Leiophron)

Distribution

Scotland, Ireland

Centistes

Haliday, 1835

Ancylocentrus

Förster, 1863

Notes

species of Centistes (Ancylocentrus) excluded from the British and Irish list:

[subsulcatus (Thomson, 1895, Leiophron)] Not included in Huddleston (1978) but listed as British in Taxapad (Yu et al. 2012) as it was recorded by Kerrich (1932), as a tentative identification (by Roman), which has not been corroborated.

Centistes (Ancylocentrus) ater

(Nees, 1834)

• Leiophron ater Nees, 1834

• excrucians (Haliday, 1835, Leiophron)

• lativalvis (Jakimavičius, 1972, Allurus)

Distribution

England, Ireland

Centistes (Ancylocentrus) collaris

(Thomson, 1895)

• Leiophron collaris Thomson, 1895

Distribution

England, Ireland

Notes

BMNH, det. Mason, Broad, added here

Centistes (Ancylocentrus) edentatus

(Haliday, 1835)

• Leiophron edentatus Haliday, 1835

Distribution

England, Ireland

Centistes (Ancylocentrus) nasutus

(Wesmael, 1838)

• Eubadizon nasutus Wesmael, 1838

• saxo (Reinhard, 1862, Leiophron)

Distribution

England

Notes

some distribution data from Luff (1976a)

Centistes

Haliday, 1835

• ANCYLUS Haliday, 1833

• ANCYLLUS Haldeman, 1842

• EUPHORIDEA Ashmead, 1900

• LIOSIGALPHUS Ashmead, 1900

Centistes (Centistes) cuspidatus

(Haliday, 1833)

• Ancylus cuspidatus Haliday, 1833

• lucidator (Nees, 1834, Bracon)

Distribution

England, Ireland

Centistes (Centistes) fuscipes

(Nees, 1834)

• Bracon fuscipes Nees, 1834

• fuscipes (Wesmael, 1835, Leiophron)

Distribution

England

Syrrhizus

Förster, 1863

Centistes (Syrrhizus) delusorius

(Förster, 1863)

• Syrrhizus delusorius Förster, 1863

Distribution

England, Scotland

Cosmophorini

Muesebeck & Walkley, 1951

Cosmophorus

Ratzeburg, 1848

• COSMOPHORINUS Viereck, 1925

Cosmophorus cembrae

Ruschka, 1925

Distribution

England

Notes

added by Shaw (1989)

Ropalophorus

Curtis, 1837

• RHOPALOPHORUS Blanchard, 1840 emendation, preocc.

• CORYNOPHORE Blanchard, 1845

• EUSTALOCERUS Förster, 1863

Ropalophorus clavicornis

(Wesmael, 1835)

• Microctonus clavicornis Wesmael, 1835

• wisconsinensis Shenefelt, 1960; synonymy by Yang et al. (2003)

Distribution

England

Dinocampini

Shaw, 1985

Dinocampus

Förster, 1863

Dinocampus coccinellae

(Schrank, 1802)

• Ichneumon coccinellae Schrank, 1802

• terminatus (Nees, 1812, Bracon)

• sculptus (Cresson, 1872, Euphorus)

• americanus (Riley, 1888, Centistes)

Distribution

England, Scotland, Wales, Ireland

Euphorini

Förster, 1863

Leiophron

Nees, 1818

Notes

Distribution and much taxonomic data from Richards (1967), New (1970) and Loan (1974).

Euphorus

Nees, 1834

Notes

Treated as a subgenus of Leiophron by Belokobylskij (2000b).

Leiophron (Euphorus) duploclaviventris

Shenefelt, 1969

• claviventris (Ruthe, 1856, Microctonus) preocc.

• ruthei Loan, 1974

Distribution

England

Leiophron (Euphorus) pallidistigma

(Curtis, 1833)

• Leiophron pallidistigma Curtis, 1833

• pallicornis (Nees, 1834, Euphorus)

• claviventris (Wesmael, 1835, Microctonus)

• intacta (Haliday, 1835, Leiophron)

• parvula (Ruthe, 1856, Microctonus)

Distribution

England, Ireland

Leiophron (Euphorus) similis

(Curtis, 1833)

• Leiophron similis Curtis, 1833

• basalis (Curtis, 1833, Leiophron)

Distribution

England, Wales, Ireland

Notes

Listed twice, under Microctonus and Leiophron, by Huddleston (1978).

Leiophron

Nees, 1818

• Liophron Förster, 1863

Leiophron (Leiophron) apicalis

Haliday, 1833

• apicalis Curtis, 1833 preocc.

• ornata (Marshall, 1887, Euphorus)

Distribution

England, Ireland, Isle of Man

Notes

The type of Euphorus ornatus is lost but Loan (1974) suggested, on the basis of possible type material (collected in England), that ornata may be a synonym of apicalis, which has been formally proposed by Papp (2003).

Leiophron (Leiophron) fascipennis

(Ruthe, 1856)

• Microctonus fascipennis Ruthe, 1856

• fasciipennis misspelling

• aciculata Belokobylskij, 1993

Distribution

England

Leiophron (Leiophron) fulvipes

Curtis, 1833

Distribution

England

Leiophron (Leiophron) heterocordyli

Richards, 1967

Distribution

England

Peristenus

Förster, 1863

Notes

species of Peristenus excluded from the British and Irish list:

[brevicornis (Herrich-Schäffer, 1838, Perilitus)] Listed by Loan (1974) as incertae sedis within Peristenus or Leiophron, by Huddleston (1978) as a doubtfully placed species of Leiophron and by van Achterberg (in Belokobylskij et al. 2003) as a species of Peristenus. We cannot trace any references to this species occurring in Britain or Ireland.

[duplobrevicornis (Shenefelt, 1969, Leiophron); syn. brevicornis (Ruthe, 1856, Microctonus) preocc.] According to Loan (1974), the type of brevicornis (Ruthe) is lost and the species cannot be identified with certainty. Recorded as British by Marshall (1872). Belokobylskij et al. (2003) list it as a species of Peristenus (=Leiophron according to Belokobylskij).

[mitis (Haliday, 1833, Leiophron)] According to van Achterberg (in Belokobylskij et al. 2003) the type is lost and was probably a deformed specimen anyway.

Peristenus accinctus

(Haliday, 1835)

• Leiophron accinctus Haliday, 1835

• laeviventris (Ruthe, 1856, Microctonus)

• intermedius (Ruthe, 1856, Microctonus)

Peristenus antennalis

(Hincks, 1943) preocc.

• Leiophron antennalis Hincks, 1943

• picipes (Haliday, 1835, Leiophron) preocc.

Distribution

Ireland

Notes

Added by Achterberg (1997). Hincks’s was a replacement name for Haliday’s use of the name picipes for a species that is distinct from picipes (Curtis). Preoccupied by Leiophron antennalis Watanabe, 1937 (now classified in Centistes (Ancylocentrus)).

Peristenus facialis

(Thomson, 1892)

• Euphorus facialis Thomson, 1892

• fascialis misspelling

• microcerus (Thomson, 1892, Euphorus)

Distribution

England, Wales

Peristenus grandiceps

(Thomson, 1892)

• Euphorus grandiceps Thomson, 1892

Distribution

England, Ireland

Notes

Not listed by Huddleston (1978), although it was taken out of synonymy with orchesiae by Loan (1974).

Peristenus malatus

Loan, 1976

Distribution

Scotland

Notes

distribution data from Loan (1976)

Peristenus nitidus

(Curtis, 1833)

• Leiophron nitidus Curtis, 1833

Distribution

England

Peristenus orchesiae

(Curtis, 1833)

• Leiophron orchesiae Curtis, 1833

• rufibarbis (Curtis, 1837, Leiophron)

Distribution

England

Peristenus orthotyli

(Richards, 1967)

• Leiophron orthotyli Richards, 1967

Distribution

England

Peristenus pallipes

(Curtis, 1833)

• Leiophron pallipes Curtis, 1833

• barbiger (Wesmael, 1835, Microctonus)

Distribution

England, Scotland, Ireland, Isle of Man

Notes

Van Achterberg has identified a Scottish specimen as barbiger, currently listed as a junior synonym of pallipes.

Peristenus pallipes

(Herrich-Schäffer, 1838) preocc.

• Perilitus pallipes Herrich-Schäffer, 1838

• mellipes (Cresson, 1872, Euphorus)

• punctatus (Provancher, 1883, Microctonus)

• tuberculifer (Marshall, 1887, Euphorus)

• nocturnus (Viereck, 1905, Brachistes)

Peristenus picipes

(Curtis, 1833)

• Leiophron picipes Curtis, 1833

• coactus (Marshall, 1887, Euphorus)

Distribution

England, Ireland

Helorimorphini

Schmiedeknecht, 1907

Chrysopophthorus

Goidanich, 1948

Chrysopophthorus hungaricus

(Kiss, 1927)

• Helorimorpha hungaricus Kiss, 1927

• chrysopimaginis Goidanich, 1948

• elegans Tobias, 1961

Distribution

England

Notes

added by Shaw (1996a)

Wesmaelia

Förster, 1863

Wesmaelia petiolata

(Wollaston, 1858)

• Euphorus petiolatus Wollaston, 1858

• pendula Förster, 1863

• cremasta Marshall, 1872

• americana Myers, 1917

• asiatica Shestakov, 1932

Distribution

England

Myiocephalini

Chen & van Achterberg, 1997

• LOXOCEPHALINI Shaw, 1985 invalid

Myiocephalus

Marshall, 1897

• LOXOCEPHALUS Förster, 1863 preocc.

• SPILOMMA Morley, 1909

Myiocephalus boops

(Wesmael, 1835)

• Microctonus boops Wesmael, 1835

• longipes (Förster, 1863, Loxocephalus)

• laticeps (Provancher, 1886, Gamosecus)

• falconivibrans (Morley, 1909, Spilomma)

• hedini (Fahringer, 1930, Aphidius)

Distribution

England, Ireland

Neoneurini

Bengtsson, 1918

• ELASMOSOMINI Viereck, 1918

Notes

Treated as a separate subfamily in many works (e.g. Shaw and Huddleston 1991, Achterberg 1993b) but recent phylogenetic studies have shown the neoneurines to be nested within the Euphorinae (e.g. Belshaw and Quicke 2002, Dowton et al. 2002).

Elasmosoma

Ruthe, 1858

• PARAMIRAX Ashmead, 1895 unavailable

Elasmosoma berolinense

Ruthe, 1858

Distribution

England

Notes

added by Shaw (2009)

Neoneurus

Haliday, 1838

• ECCLITES Förster, 1863

• SIXIA Vollenhoven, 1867; synonymy by Achterberg (1997)

Neoneurus auctus

(Thomson, 1895)

• Elasmosoma aucta Thomson, 1895

• halidaii Marshall, 1897

• bistigmaticus (Morley, 1909, Euphorus)

Distribution

England, Scotland

Perilitini

Förster, 1863

• MICROCTONINI Shaw, 1985

Microctonus

Wesmael, 1835

• GAMOSECUS Provancher, 1880

Notes

Haeselbarth (2008) described many new species in Perilitus (Microctonus) but as Microctonus was raised to generic rank by Stigenberg et al. (2015), these are effectively new combinations in Microctonus. Synonymy follows Haeselbarth (2008).

Microctonus aciculatus

(Haeselbarth, 2008)

• Perilitus aciculatus Haeselbarth, 2008

Distribution

Ireland

Notes

added by Haeselbarth (2008)

Microctonus aethiops

Nees, 1834

• spurius Ruthe, 1856

• Microctonus aethiops ?brevispina (Thomson, 1892, Euphorus)

• aethiopoides Loan, 1975

Distribution

England, Ireland

Microctonus alticae

(Haeselbarth, 2008)

• Perilitus alticae Haeselbarth, 2008

Distribution

England

Notes

added by Haeselbarth (2008)

Microctonus aphthonae

(Haeselbarth, 2008)

• Perilitus aphthonae Haeselbarth, 2008

Distribution

England, Ireland

Notes

added by Haeselbarth (2008)

Microctonus apiophaga

Loan, 1974

Microctonus areolatus

(Thomson, 1892)

• Perilitus areolatus Thomson, 1892

Distribution

England

Microctonus belokobylskiji

(Haeselbarth, 2008)

• Perilitus belokobylskiji Haeselbarth, 2008

Distribution

Ireland

Notes

added by Haeselbarth (2008)

Microctonus brassicae

(Haeselbarth, 2008)

• Perilitus brassicae Haeselbarth, 2008

Distribution

England

Notes

added by Haeselbarth (2008)

Microctonus brevicollis

(Haliday, 1835)

• Perilitus brevicollis Haliday, 1835

Distribution

England, Ireland

Microctonus cerealium

(Haliday, 1835)

• Perilitus cerealium Haliday, 1835

• secalis (Haliday, 1833, Perilitus) unavailable

Distribution

England, Ireland

Microctonus colesi

Drea, 1968

Distribution

Scotland, Ireland

Notes

added by Haeselbarth (2008)

Microctonus consuetor

(Nees, 1834)

• Perilitus consuetor Nees, 1834

Distribution

Scotland

Notes

added by Haeselbarth (2008)

Microctonus debilis

(Wollaston, 1858)

• Perilitus debilis Wollaston, 1858

• Microctonus debilis ?gracilipes (Thomson, 1892, Perilitus)

Distribution

Ireland

Notes

added by Haeselbarth (2008)

Microctonus fagi

(Haeselbarth, 2008)

• Perilitus fagi Haeselbarth, 2008

Distribution

England, Ireland

Notes

Added by Haeselbarth (2008); specimens tentatively associated with the type material.

Microctonus fittkaui

(Haeselbarth, 2008)

• Perilitus fittkaui Haeselbarth, 2008

Distribution

England

Notes

added by Haeselbarth (2008)

Microctonus flaviventris

(Thomson, 1892)

• Perilitus flaviventris Thomson, 1892

• areolatus (Thomson, 1892, Perilitus)

Distribution

Ireland

Notes

added by Haeselbarth (2008)

Microctonus haszprunari

(Haeselbarth, 2008)

• Perilitus haszprunari Haeselbarth, 2008

Distribution

Ireland

Notes

added by Haeselbarth (2008)

Microctonus lipari

Čapek & Starý, 1995

Distribution

England

Notes

added by Haeselbarth (2008)

Microctonus melanopus

Ruthe, 1856

Distribution

England, Scotland, Ireland

Notes

added by Haeselbarth (2008)

Microctonus parcicornis

Ruthe, 1856

Distribution

England

Microctonus perforatus

(Haeselbarth, 2008)

• Perilitus perforatus Haeselbarth, 2008

Distribution

England

Notes

added by Haeselbarth (2008)

Microctonus podargae

(Haeselbarth, 2008)

• Perilitus podargae Haeselbarth, 2008

Distribution

England

Notes

added by Haeselbarth (2008)

Microctonus retusus

Ruthe, 1856

• lancearius Ruthe, 1856

• caudatus (Thomson, 1892, Perilitus)

Distribution

England, Ireland

Notes

Added by Luff (1976b); overlooked by Huddleston (1978) and O'Connor et al. (1999).

Microctonus silvularis

(Haeselbarth, 2008)

• Perilitus silvularis Haeselbarth, 2008

Distribution

Scotland, Ireland

Notes

added by Haeselbarth (2008)

Microctonus stenocari

(Haeselbarth, 2008)

• Perilitus stenocari Haeselbarth, 2008

Distribution

Scotland

Notes

added by Haeselbarth (2008)

Microctonus strophosomi

(Haeselbarth, 2008)

• Perilitus strophosomi Haeselbarth, 2008

Distribution

Scotland, Ireland

Notes

added by Haeselbarth (2008)

Microctonus thyellae

(Haeselbarth, 2008)

• Perilitus thyellae Haeselbarth, 2008

Distribution

England, Scotland

Notes

added by Haeselbarth (2008)

Perilitus

Nees, 1818

Notes

species of Perilitus excluded from the British and Irish list:

[falciger (Ruthe, 1856, Microctonus)] Richards (1960) states that it is not known with certainty from Britain.

Perilitus areolaris

Gerdin & Hedqvist, 1985

Distribution

England

Notes

added by Haeselbarth (1999)

Perilitus dubius

(Wesmael, 1838)

• Microctonus dubius Wesmael, 1838

• rutilus Herrich-Schäffer, 1838 preocc.

Distribution

England

Perilitus foveolatus

Reinhard, 1862

• sicheli Giard, 1895; synonymy by Haeselbarth (1999)

Distribution

England, Wales

Perilitus marci

Haeselbarth, 1999

Distribution

England

Notes

added by Haeselbarth (1999)

Perilitus rutilus

(Nees, 1811)

• Bracon rutilus Nees, 1811

• luteus Herrich-Schäffer, 1838

• ruralis Herrich-Schäffer, 1838

• strenuus Marshall, 1887; synonymy by Haeselbarth (1999)

• pyri (Viereck, 1917, Dinocampus)

• tuberculus Zaykov, 1981; synonymy by Haeselbarth (1999)

Distribution

England, Ireland

Rilipertus

Haeselbarth, 1996

Rilipertus intricatus

(Ruthe, 1859)

• Microctonus intricatus Ruthe, 1859

• borealis (Thomson, 1892, Perilitus); synonymy by Haeselbarth (1996)

Distribution

Scotland, Ireland

Spathicopis

van Achterberg, 1977

flavocephala

van Achterberg, 1977

Distribution

England

Notes

NMS, det. van Achterberg, added here

Pygostolini

Belokobylskij, 2000

Pygostolus

Haliday, 1833

Notes

Distribution data from Achterberg (1992a) and BMNH.

Pygostolus falcatus

(Nees, 1834)

• Leiophron falcatus Nees, 1834

• testaceus misident.

Distribution

England, Ireland

Pygostolus multiarticulatus

(Ratzeburg, 1852)

• Blacus multiarticulatus Ratzeburg, 1852

• falcatus (Wesmael, 1838, Blacus) preocc.

Distribution

England

Pygostolus otiorhynchi

(Boudier, 1834)

• Bracon otiorhynchi Boudier, 1834

Distribution

England, Scotland

Notes

Achterberg (1992a) separated otiorhynchi from falcatus but Belokobylskij et al. (2003), without comment, treated the two names as synonymous again. This is not followed here as otiorhynchi and falcatus seem to be distinct species.

Pygostolus sticticus

(Fabricius, 1798)

• Ichneumon sticticus Fabricius, 1798

• testaceus (Fallén, 1813, Bassus) preocc.

• sticticator (Thunberg, 1824, Ichneumon)

• barynoti (Boudier, 1834, Bracon)

• gigas (Wesmael, 1835, Blacus)

Distribution

England, Scotland, Ireland, Isle of Man

Syntretini

Shaw, 1985

Syntretus

Förster, 1863

• FALCOSYNTRETUS Tobias, 1965

• PARASYNTRETUS Belokobylskij, 1993

Notes

Distribution and synonymic data from Achterberg and Haeselbarth (2003).

Species of Syntretus excluded from the British and Irish list:

[parvicornis (Ruthe, 1862, Microctonus)] Not listed as British or Irish by Achterberg and Haeselbarth (2003).

Syntretus breviradialis

van Achterberg & Haeselbarth, 2003

Distribution

England

Notes

added by Broad (2009)

Syntretus conterminus

(Nees, 1834)

• Perilitus conterminus Nees, 1834

Distribution

England, Scotland, Ireland

Syntretus elegans

(Ruthe, 1856)

• Microctonus elegans Ruthe, 1856

• transsylvanicus (Kiss, 1927, Perilitus)

Distribution

Ireland

Notes

added by Achterberg and Haeselbarth (2003)

Syntretus flevo

van Achterberg & Haeselbarth, 2003

Distribution

England, Ireland

Notes

added by Achterberg and Haeselbarth (2003)

Syntretus fuscicoxis

van Achterberg & Haeselbarth, 2003

Distribution

England, Ireland

Notes

added by Achterberg and Haeselbarth (2003)

Syntretus fuscivalvis

van Achterberg & Haeselbarth, 2003

Distribution

England, Scotland, Ireland

Notes

added by Achterberg and Haeselbarth (2003)

Syntretus idalius

(Haliday, 1833)

• Perilitus idalius Haliday, 1833

• vernalis (Wesmael, 1835, Microctonus)

• cultus (Marshall, 1887, Microctonus)

Distribution

England, Scotland, Ireland

Syntretus ocularis

van Achterberg & Haeselbarth, 2003

Distribution

England, Wales, Scotland

Notes

added by Achterberg and Haeselbarth (2003)

Syntretus politus

(Ruthe, 1856)

• Microctonus politus Ruthe, 1856

• cynthius (Curtis, 1837, Microctonus) nom. nud.

• cynthius Lyle, 1927

Distribution

England, Ireland

Syntretus pusio

(Marshall, 1898)

• Microctonus pusio Marshall, 1898

Distribution

England, Ireland

Syntretus splendidus

(Marshall, 1887)

• Microctonus splendidus Marshall, 1887

• testaceus (Capron, 1887, Microctonus)

• suffolciensis (Morley, 1933, Dyscritus) new synonymy

• niger Tobias, 1976

Distribution

England, Scotland, Ireland

Notes

The female holotype of Dyscritus suffolciensis Morley in BMNH, described as an aphidiine, has remained uninterpreted, although Starý (1959) recognised that it is a euphorine. It is a normal specimen of Syntretus splendidus.

Syntretus taegeri

van Achterberg & Haeselbarth, 2003

Distribution

England

Notes

added by Achterberg and Haeselbarth (2003)

Syntretus xanthocephalus

(Marshall, 1887)

• Microctonus xanthocephalus Marshall, 1887

• tempestivus (Curtis, 1837, Microctonus) nom. nud.

• lyctaea Cole, 1959

• lyctae misspelling

Distribution

England, Scotland, Ireland

Syntretus zuijleni

van Achterberg & Haeselbarth, 2003

Distribution

England, Scotland, Ireland

Notes

added by Achterberg and Haeselbarth (2003)

Townesilitini

Shaw, 1985

Streblocera

Westwood, 1833

Notes

Cosmophoridia Hedqvist, 1955 and Eutanycerus, usually considered synonymous with Streblocera, were regarded as valid subgenera by Chen and Achterberg (1997).

Streblocera

Westwood, 1833

• LECYTHODELLA Enderlein, 1912

Streblocera (Streblocera) fulviceps

Westwood, 1833

Distribution

England

Streblocera (Streblocera) longiscapha

Westwood, 1882

Distribution

England

Eutanycerus

Förster, 1863

• VILLOCERA Chen & van Achterberg, 1997; synonymy by Belokobylskij (2000a)

Streblocera (Eutanycerus) macroscapa

(Ruthe, 1856)

• Microctonus macroscapus Ruthe, 1856

• halidayana (Förster, 1863, Eutanycerus)

Distribution

England

Townesilitus

Haeselbarth & Loan, 1983

Townesilitus aemulus

(Ruthe, 1856)

• Microctonus aemulus Ruthe, 1856

• punctifrontis (Watanabe, 1955, Microctonus)

Distribution

England, Wales, Ireland

Notes

added by Haeselbarth (1988)

Townesilitus bicolor

(Wesmael, 1835)

• Microctonus bicolor Wesmael, 1835

• breviradialis (Tobias, 1976, Microctonus)

Distribution

England, Wales, Ireland

Townesilitus deceptor

(Wesmael, 1835)

• Microctonus deceptor Wesmael, 1835

Distribution

England, Scotland, Ireland

Notes

added by Haeselbarth (1988)

Townesilitus fulviceps

(Ruthe, 1856)

• Microctonus fulviceps Ruthe, 1856

Distribution

England

Notes

added by Haeselbarth (1988)

Exothecinae

Exothecinae

Förster, 1863

• PHANOMERINAE Fahringer, 1928

Colastes

Haliday, 1833

Notes

Belokobylskij (1998) treats Shawiana and Xenarcha as subgenera of Colastes.

Colastes

Haliday, 1833

• EXOTHECUS Wesmael, 1838

• PHANOMERIS Förster, 1863

• PHAENOMERIS Dalla Torre, 1898

Colastes (Colastes) affinis

(Wesmael, 1838)

• Exothecus affinis Wesmael, 1838

Distribution

England, Scotland, Wales

Notes

NMS, det. Shaw & van Achterberg, added here; treated as a species of Colastes (Xenarcha) by Belokobylskij (1998).

Colastes (Colastes) braconius

Haliday, 1833

• debilis (Wesmael, 1838, Exothecus)

• gracilis Papp, 1975

Distribution

England, Scotland, Wales, Ireland

Notes

some distribution data from Shaw and Askew (1976)

Colastes (Colastes) fragilis

(Haliday, 1836)

• Rogas fragilis Haliday, 1836

• semeyticus Jakimavičius, 1969

Distribution

England, Scotland

Colastes (Colastes) incertus

(Wesmael, 1838)

• Exothecus incertus Wesmael, 1838

Distribution

England, Scotland

Colastes (Colastes) magdalenae

Sterzynski, 1983

Distribution

England

Notes

NMS, det. Shaw & van Achterberg, added on Fauna Europaea

Colastes (Colastes) pubicornis

(Thomson, 1892)

• Exothecus pubicornis Thomson, 1892

Distribution

England, Scotland

Notes

added by Godfray and McGavin (1985)

Colastes (Colastes) vividus

Papp, 1975

Distribution

England

Notes

NMS, det. Shaw, added here

Fungivenator

van Achterberg & Shaw, 2008

Colastes (Fungivenator) sandei

van Achterberg & Shaw, 2008

Distribution

England

Notes

added by van Achterberg and Shaw (2008)

Shawiana

van Achterberg, 1983

• PHANOMERIS misident.

Shawiana catenator

(Haliday, 1836)

• Rogas catenator Haliday, 1836

Distribution

England, Scotland, Wales

Notes

some distribution data from Shaw and Askew (1976)

Shawiana laevis

(Thomson, 1892)

• Exothecus laevis Thomson, 1892

• rugulosus (Hellén, 1959, Colastes)

Distribution

England

Notes

NMS, det. Shaw, added on Fauna Europaea

Xenarcha

Förster, 1863

• ZAMEGASPILUS Ashmead, 1900

Xenarcha abnormis

(Wesmael, 1838)

• Exothecus abnormis Wesmael, 1838

• glabricollis (Thomson, 1892, Exothecus)

Distribution

England, Scotland, Wales

Notes

NMS, det. Shaw, added on Fauna Europaea

Xenarcha lustrator

(Haliday, 1836)

• Rogas lustrator Haliday, 1836

• dimidiatus (Nees, 1834, Bracon) preocc.

• lustratrix Schulz, 1906

• thomsoni (Szépligeti, 1906, Phanomeris)

Distribution

England, Wales, Ireland

Gnamptodontinae

Gnamptodontinae

Fischer, 1970

• GNAPTODONTINAE misspelling

• GNAPTOGASTRINAE Tobias, 1976

Gnamptodon

Haliday, 1836

• GNAPTODON Haliday, 1837 suppressed

• DIRAPHUS Wesmael, 1838

• MESOTAGES Förster, 1863

Gnamptodon decoris

(Förster, 1863)

• Mesotages decoris Förster, 1863

• klemensiewiczii Niezabitowski, 1910

• bachmaieri Fischer, 1957

Distribution

England, Wales

Notes

NMS, det. Shaw, added on Fauna Europaea

Gnamptodon pumilio

(Nees, 1834)

• Bracon pumilio Nees, 1834

• pygmaeus (Wesmael, 1838, Diraphus)

Distribution

England, Scotland, Ireland

Notes

some distribution data from Shaw and Askew (1976)

Helconinae

Helconinae

Förster, 1863

Notes

The Diospilini were removed to the Brachistinae by Sharanowski et al. (2011).

Helconini

Förster, 1863

Notes

Much distribution and taxonomic data from Achterberg (1987).

Helcon

Nees, 1814

• GYMNOSCELUS Förster, 1863

• EDYIA Cameron, 1905

• COELOSTEPHANUS Kieffer, 1911

Helcon claviventris

Wesmael, 1835

Distribution

England

Notes

added by Achterberg (1987)

Helcon tardator

Nees, 1812

• Helcon tardator ?adulterator (Villers, 1789, Ichneumon)

Distribution

England

Helconidea

Viereck, 1914

Helconidea dentator

(Fabricius, 1804)

• Pimpla dentator Fabricius, 1804

• aequator (Nees, 1812, Helcon)

• tentator (Thunberg, 1824, Ichneumon) preocc.

• rugator (Ratzeburg, 1848, Helcon)

• armator (Marshall, 1898, Helcon)

• dentatrix (Schulz, 1906, Pimpla)

Distribution

England

Notes

Helconidea armator (Marshall, 1898, Helcon) removed from synonymy by Achterberg (2014).

Helconidea ruspator

(Linnaeus, 1758)

• Ichneumon ruspator Linnaeus, 1758

• dentator (Nees, 1812, Helcon) preocc.

Distribution

England

Wroughtonia

Cameron, 1899

• DUPORTIA Kieffer, 1921

Wroughtonia spinator

(Lepeletier, 1825)

• Helcon spinator Lepeletier, 1825

• annulicornis (Nees, 1834, Helcon)

Distribution

England

Homolobinae

Homolobinae

van Achterberg, 1979

Homolobus

Förster, 1863

• ZELE misident.

Notes

Shaw (2010) summarises the taxonomy and biology of British and Irish species, including some distribution data.

Apatia

Enderlein, 1920

Homolobus (Apatia) truncator

(Say, 1828)

• Bracon truncator Say, 1828

• calcarator (Wesmael, 1835, Phylax)

• melleus (Cresson, 1872, Phylax)

• crassicalcaratus (Viereck, 1905, Zele)

• calcaratrix (Schulz, 1906, Zele)

• fuscitarsis (Bengtsson, 1918, Phylacter)

• simillimus (Enderlein, 1920, Apatia)

• unicolor (Enderlein, 1920, Zele)

• chlorophthalmus (Nixon, 1938, Zele)

Distribution

England

Notes

Nixon’s description of Zele chlorophthalmus validated the use of the name for this taxon; chlorophthalmus of authors is a misidentification (Bracon chlorophthalmus Spinola, 1808 is actually a species of the true Zele).

Chartolobus

van Achterberg, 1979

Homolobus (Chartolobus) infumator

(Lyle, 1914)

• Zele infumator Lyle, 1914

• wesmaeli (Bengtsson, 1918, Phylacter)

• japonicus (Watanabe, 1932, Zele)

Distribution

England, Scotland, Wales, Ireland

Homolobus

Förster, 1863

Homolobus (Homolobus) discolor

(Wesmael, 1835)

• Phylax discolor Wesmael, 1835

• pectoralis (Herrich-Schäffer, 1838, Rogas); synonymy by Achterberg (1992b)

Distribution

England, Wales, Ireland

Oulophus

van Achterberg, 1979

Homolobus (Oulophus) flagitator

(Curtis, 1837)

• Zele flagitator Curtis, 1837

• geminator (Lyle, 1914, Zele)

Distribution

England, Scotland, Wales, Ireland

Phylacter

Reinhard, 1863

Homolobus (Phylacter) annulicornis

(Nees, 1834)

• Rogas annulicornis Nees, 1834

• testaceator misident.

• simplex (Herrich-Schäffer, 1838, Rogas); synonymy by Achterberg (1992b)

Distribution

England, Scotland, Ireland

Hormiinae

Hormiinae

Förster, 1863

Hormius

Nees, 1818

• CHLIDONIA Herrich-Schäffer, 1838

• HORMIELLUS Enderlein, 1912

• MEDIELLA Hedqvist, 1963

• ANHORMIUS Belokobylskij, 1989

Hormius maderae

Graham, 1986

Distribution

England

Notes

NMS, det. Shaw & van Achterberg, added on Fauna Europaea

Hormius moniliatus

(Nees, 1811)

• Bracon moniliatus Nees, 1811

• brevipennis Hellén, 1957

• dusmeti (Docavo Alberti, 1960, Hormiopterus)

• insularis Hedqvist, 1965

• coniceps Hellén, 1957

Distribution

England, Ireland, Isle of Man

Hormius piciventris

Wesmael, 1838

Distribution

England, Scotland, Wales, Ireland, Isle of Man

Notes

NMS, det. Shaw & van Achterberg, added on Fauna Europaea. Considered here to be a valid species; van Achterberg (in Belokobylskij et al. 2003) considered that piciventris may be a distinct species but it is listed as a synonym of moniliatus in Taxapad (Yu et al. 2012).

Ichneutinae

Ichneutinae

Förster, 1863

Ichneutini

Förster, 1863

Ichneutes

Nees, 1814

Ichneutes brevis

Wesmael, 1835

Distribution

England, Ireland

Ichneutes reunitor

Nees, 1816

• costatus (Zetterstedt, 1838, Microgaster)

• laeviventris Hellén, 1958

• leptostigma Hellén, 1958

Distribution

England, Ireland

Proteropini

van Achterberg, 1976

Proterops

Wesmael, 1835

• ICHNEUTIDEA Ashmead, 1900

• PROTEROPOIDES Viereck, 1909

Proterops nigripennis

Wesmael, 1835

Distribution

England, Scotland, Ireland

Macrocentrinae

Macrocentrinae

Förster, 1863

Austrozele

Roman, 1910

• PANISCOZELE Enderlein, 1920

• PALINZELE Brues, 1922

• LAEVIS Sharma, 1982

Austrozele longipalpis

van Achterberg, 1993

Distribution

England

Notes

added by Achterberg (1993a)

Macrocentrus

Curtis, 1833

• AMICROPLUS Förster, 1863

• AMICROPLITES Dalla Torre, 1898

• FHOGRA Cameron, 1901

• LEPTOZELE Cameron, 1910

• METAPLEURODON Enderlein, 1920

• PSEUDOPHYLACTER Fahringer, 1929

Notes

Distribution data from Eady and Clark (1964) and Achterberg and Haeselbarth (1983).

Macrocentrus bicolor

Curtis, 1833

• limbator (Ratzeburg, 1848, Rogas)

• gracilipes Telenga, 1935

Distribution

England, Scotland, Wales, Ireland, Isle of Man

Notes

Achterberg and Haeselbarth (1983) and Achterberg (1993a) treat gracilipes as a synonym of bicolor but Belokobylskij et al. (2003) list it as a synonym of thoracicus.

Macrocentrus blandus

Eady and Clark, 1964

Distribution

England, Scotland, Ireland

Macrocentrus cingulum

Brischke, 1882

• grandii Goidanich, 1937

• gifuensis misident.

Distribution

England

Macrocentrus collaris

(Spinola, 1808)

• Bracon collaris Spinola, 1808

• ebeninus (Nees, 1834, Bracon)

• dubius (Wesmael, 1835, Eubadizon)

• picipes (Haliday, 1835, Helcon)

• dispar (Kollar, 1852, Bracon) preocc.

• kollari (Rondani, 1877, Bracon); synonymy by Papp (1996b)

• affinis Hedwig, 1961

• affiniqades Shenefelt, 1969

Distribution

England, Ireland

Macrocentrus equalis

Lyle, 1914

Distribution

England

Macrocentrus infirmus

(Nees, 1834)

• Rogas infirmus Nees, 1834

Distribution

England, Scotland, Ireland, Isle of Man

Macrocentrus linearis

(Nees, 1812)

• Bracon linearis Nees, 1812

• abdominalis (Fabricius, 1793, Ichneumon) preocc.

• abdominator (Thunberg, 1824, Ichneumon)

• fissura (Thunberg, 1824, Ichneumon)

• pallidator (Zetterstedt, 1838, Bracon) preocc.

• tenuis (Ratzeburg, 1848, Rogas)

• iridescens French, 1880

• gifuensis Ashmead, 1906

• amicroploides Viereck, 1912

• pallidatorius (Fahringer, 1928, Bracon)

Distribution

England, Scotland, Ireland, Isle of Man

Macrocentrus marginator

(Nees, 1811)

• Bracon marginator Nees, 1811

• rugator (Ratzeburg, 1848, Rogas)

Distribution

England, Scotland, Ireland

Notes

The North American Macrocentrus aegeriae Rohwer, 1915 was removed from synonymy by Achterberg and Haeselbarth (1983).

Macrocentrus nidulator

(Nees, 1834)

• Rogas nidulator Nees, 1834

• longicaudis (Herrich-Schäffer, 1838, Rogas)

• procerus Costa, 1884

• curticaudis Telenga, 1950

Distribution

England, Scotland, Wales, Ireland

Macrocentrus nitidus

(Wesmael, 1835)

• Rogas nitidus Wesmael, 1835

Distribution

England, Scotland

Macrocentrus pallipes

(Nees, 1811)

• Bracon pallipes Nees, 1811

• pallidipes Dalla Torre, 1898

Distribution

England

Macrocentrus resinellae

(Linnaeus, 1758)

• Ichneumon resinellae Linnaeus, 1758

• resinator (Thunberg, 1824, Ichneumon) unavailable

• flavipes (Ratzeburg, 1844, Rogas)

• interstitialis (Ratzeburg, 1844, Rogas)

• obscurator (Ratzeburg, 1848, Rogas)

• intricator (Ratzeburg, 1852, Helcon)

• punctifrons Thomson, 1895

• sublaevis Thomson, 1895

Distribution

England, Scotland

Macrocentrus sylvestrellae

van Achterberg, 2001

Distribution

England

Notes

NMS, det. Shaw, added here

Macrocentrus thoracicus

(Nees, 1811)

• Bracon thoracicus Nees, 1811

• longicornis (Wesmael, 1835, Rogas)

Distribution

England, Wales

Macrocentrus townesi

van Achterberg & Haeselbarth, 1983

Distribution

England, Scotland, Ireland

Notes

added by Achterberg and Haeselbarth (1983)

Meteorinae

Meteorinae

Cresson, 1887

• ZELINAE Ashmead, 1900

• ZEMIOTINAE van Achterberg, 1976

Notes

Stigenberg et al. (2015) included the meteorines as a basal tribe of Euphorinae, rather than as a separate subfamily, as was the preference of many authors, for example, Shaw and Huddleston (1991). In light of their very different developmental biology (parasitoids of larval Lepidoptera and Coleoptera as opposed to parasitising adult insects) and sister-group position to the euphorines, we retain the subfamily rank for Meteorinae.

Meteorus

Haliday, 1835

• SAPROTICHUS Holmgren, 1868

• PACHYTHECUS Cameron, 1912 preocc.

Notes

Distribution and synonymic data from Huddleston (1980), Stigenberg and Ronquist (2011) and Stigenberg & Shaw (2013), except where noted. Taxonomy follows Stigenberg and Ronquist (2011).

Species of Meteorus excluded from the British and Irish list:

[flaviceps (Ratzeburg, 1844, Perilitus) nom. dub.]

Meteorus abdominator

(Nees, 1811)

• Perilitus abdominator Nees, 1811

• brunnipes (Ruthe, 1862, Perilitus)

• bruneipes Dalla Torre, 1898

• brevipesalis Shenefelt, 1969

• Meteorus abdominator ?delator (Haliday, 1835, Perilitus); synonymy by Achterberg (1997)

Distribution

England, Scotland, Wales, Ireland

Meteorus abscissus

Thomson, 1895

Distribution

England, Scotland, Ireland

Meteorus affinis

(Wesmael, 1835)

• Perilitus affinis Wesmael, 1835

• gracilis Ruthe, 1862 preocc.

• punctiventris Ruthe, 1862

• ruthei Schmiedeknecht, 1897

• voloscensis Fischer, 1959

Distribution

England, Scotland, Wales, Ireland

Meteorus alborossicus

Lobodenko, 2000

Distribution

England

Notes

added by Stigenberg and Ronquist (2011)

Meteorus brevicauda

Thomson, 1895

• thuringiacus Schmiedeknecht, 1897

• mongolicus Fahringer, 1935

Distribution

England

Notes

Listed by Huddleston (1978) but Huddleston (1980) could not find any British or Irish material. Shaw (1988b) subsequently recorded it as a British species.

Meteorus cespitator

(Thunberg, 1824)

• Ichneumon cespitator Thunberg, 1824

• atrator (Curtis, 1832, Zele)

• similator (Nees, 1834, Perilitus)

• microcerus (Wesmael, 1835, Perilitus)

• humeralis (Zetterstedt, 1838, Bracon)

• rufipes (Zetterstedt, 1838, Bracon)

• ambiguus Ruthe, 1862

Distribution

England, Scotland, Wales, Ireland

Meteorus cinctellus

(Spinola, 1808)

• Bracon cinctellus Spinola, 1808

• fuscipes (Wesmael, 1835, Perilitus)

Distribution

England, Scotland, Wales

Notes

This has recently been identified as Meteorus necator (e.g. Belokobylskij et al. 2003) but, according to Stigenberg and Ronquist (2011), Ichneumon necator Fabricius, 1777, is actually a species of Microgastrinae.

Meteorus cis

(Bouché, 1834)

• Bracon cis Bouché, 1834

• profligator (Haliday, 1835, Perilitus); synonymy by van Achterberg in Belokobylskij et al. (2003)

Distribution

England, Scotland, Wales, Ireland

Meteorus colon

(Haliday, 1835)

• Perilitus colon Haliday, 1835

• fragilis (Wesmael, 1835, Perilitus)

• fasciatus (Ratzeburg, 1844, Perilitus)

• alternatus Ruthe, 1862

• continuus Ruthe, 1862

• luridus Ruthe, 1862

• pallidus Ruthe, 1862

• trivittatus Ruthe, 1862

Distribution

England, Scotland, Wales, Ireland

Meteorus consimilis

(Nees, 1834)

• Perilitus consimilis Nees, 1834

• brevipes (Wesmael, 1835, Perilitus)

• albicornis Ruthe, 1862

• flagellatus Alexeev, 1971

Distribution

England, Ireland

Meteorus eadyi

Huddleston, 1980

Distribution

England, Wales

Notes

added by Huddleston (1980)

Meteorus eklundi

Stigenberg, 2011

Distribution

England

Notes

added by Stigenberg and Ronquist (2011)

Meteorus filator

(Haliday, 1835)

• Perilitus filator Haliday, 1835

• laticeps (Wesmael, 1835, Perilitus)

• hodisensis Fischer, 1970

Distribution

England, Scotland, Ireland

Meteorus heliophilus

Fischer, 1970

Distribution

England

Notes

added by Huddleston (1980)

Meteorus hirsutipes

Huddleston, 1980

Distribution

England, Ireland

Notes

added by Huddleston (1980)

Meteorus ictericus

(Nees, 1811)

• Bracon ictericus Nees, 1811

• minutor (Thunberg, 1824, Ichneumon)

• lucidator (Trentepohl, 1829, Bracon)

• ephippium (Curtis, 1832, Zele)

• xanthomelas (Wesmael, 1835, Perilitus)

• rubriceps (Ratzeburg, 1844, Perilitus)

• confinis Ruthe, 1862

• consors Ruthe, 1862

• fallax Ruthe, 1862

• liquis Ruthe, 1862

• pleuralis Ruthe, 1862

• crassicrus Thomson, 1895

• lophyriphagus Fahringer, 1934

• dumbletoni Muesebeck, 1939

• adoxophyesi Minamikawa, 1954

• makinoharanus Minamikawa, 1954

Distribution

England, Scotland, Wales, Ireland, Isle of Man

Meteorus jaculator

(Haliday, 1835)

• Perilitus jaculator Haliday, 1835

• obscurellus Ruthe, 1862

• tenuicornis Thomson, 1895

• turcicus Fahringer, 1944

Distribution

England, Scotland, Wales, Ireland

Meteorus limbatus

Maeto, 1989

Distribution

England, Scotland

Notes

added by Stigenberg and Ronquist (2011)

Meteorus lionotus

Thomson, 1895

• ruficoloratus Fischer, 1957

Distribution

England, Scotland

Notes

added by Huddleston (1980)

Meteorus longipilosus

Stigenberg, 2011

Distribution

England, Wales

Notes

added by Stigenberg and Shaw (2013)

Meteorus melanostictus

Capron, 1887

• niger Lyle, 1913

• monachae Tobias, 1986; synonymy by Belokobylskij (2000b)

Distribution

England

Meteorus micropterus

(Haliday, 1835)

• Perilitus micropterus Haliday, 1835

Distribution

England, Scotland, Wales, Ireland

Meteorus obfuscatus

(Nees, 1811)

• Bracon obfuscatus Nees, 1811

• thoracicus (Curtis, 1832, Zele)

• formosus (Wesmael, 1835, Perilitus)

• orchesiae (Boie, 1841, Alysia)

• fodori Papp, 1973

Distribution

England

Meteorus obsoletus

(Wesmael, 1835)

• Perilitus obsoletus Wesmael, 1835

• viridanae Johansson, 1964

Distribution

England, Scotland, Ireland

Notes

added by Huddleston (1980)

Meteorus oculatus

Ruthe, 1862

Distribution

Scotland

Notes

added by Stigenberg and Shaw (2013)

Meteorus pendulus

(Müller, 1776)

• Ichneumon pendulus Müller, 1776

• pendulator (Latreille, 1799, Ichneumon)

• gyrator (Thunberg, 1824, Ichneumon); synonymy by van Achterberg in Belokobylskij et al. (2003)

• ochraceator (Curtis, 1832, Zele) nom. nud.

• scutellator (Nees, 1834, Perilitus)

• petiolator (Zetterstedt, 1838, Bracon)

• parvulus Thomson, 1895

Distribution

England, Scotland, Wales, Ireland

Meteorus pulchricornis

(Wesmael, 1835)

• Perilitus pulchricornis Wesmael, 1835

• striatus Thomson, 1895

• thomsoni Marshall, 1899

• japonicus Ashmead, 1906

• nipponensis Viereck, 1912

• baicalensis Telenga, 1950

• graeffei Fischer, 1957

• macedonicus Fischer, 1957

• tuberculifer Fischer, 1957

Distribution

England, Scotland, Wales, Ireland

Notes

added by Huddleston (1980)

Meteorus rubens

(Nees, 1811)

• Bracon rubens Nees, 1811

• leviventris (Wesmael, 1835, Perilitus)

• islandicus Ruthe, 1859

• medianus Ruthe, 1862

• vulgaris (Cresson, 1872, Perilitus)

• dejanus (Rondani, 1877, Perilitus); synonymy by Papp (1996b)

• scutatus Costa, 1884

• coquilleti Ashmead, 1889

• heteroneurus Thomson, 1895

• mellinervis Viereck, 1903

• mamestrae Viereck, 1913

• szechuanensis Fahringer, 1935

• mesopotamicus Fischer, 1957

Distribution

England, Scotland, Wales, Ireland, Isle of Man

Meteorus ruficeps

Nees, 1834

• pallipes (Wesmael, 1835, Perilitus)

• nigritarsis Ruthe, 1862

• pallidipes Marshall, 1887

Distribution

England, Scotland, Ireland, Isle of Man

Meteorus rufus

(DeGeer, 1773)

• Ichneumon rufus DeGeer, 1773

• rufus (Retzius, 1783, Ichneumon) preocc.

• unicolor (Wesmael, 1835, Perilitus); synonymy by van Achterberg in Belokobylskij et al. (2003)

• chinensis (Holmgren, 1868, Saprotichus)

Distribution

England, Wales

Meteorus sibyllae

Stigenberg, 2011

Distribution

England, Scotland

Notes

added by Stigenberg and Shaw (2013)

Meteorus sulcatus

Szépligeti, 1896

• insignis Muesebeck, 1939

• molorchi Fischer, 1966

Distribution

England

Meteorus tabidus

(Wesmael, 1835)

• Perilitus tabidus Wesmael, 1835

• dubius Ruthe, 1862

• facialis Ruthe, 1862

• pentheri Fischer, 1970

Distribution

England, Scotland, Ireland

Meteorus tenellus

Marshall, 1887

• boreus Tobias, 1986

Distribution

Scotland

Notes

Added by Stigenberg and Shaw (2013). Separated from cinctellus by Stigenberg and Ronquist (2011); Marshall (1887) gave no locality data so the first documented British occurrence was published by Stigenberg and Shaw (2013).

Meteorus versicolor

(Wesmael, 1835)

• Perilitus versicolor Wesmael, 1835

• bimaculatus (Wesmael, 1835, Perilitus)

• unicolor (Hartig, 1838, Perilitus) preocc.

• brevicornis (Ratzeburg, 1844, Perilitus) preocc.

• rugator (Ratzeburg, 1852, Perilitus)

• decoloratus Ruthe, 1862

• ikonomovi Fischer, 1959

• hartigi Shenefelt, 1969

Distribution

England, Scotland, Wales, Ireland

Meteorus vexator

(Haliday, 1835)

• Perilitus vexator Haliday, 1835

Distribution

England, Scotland, Ireland

Zele

Curtis, 1832

• PROTELUS Förster, 1863

• SEMIOTES Förster, 1863

Notes

Synonymic and some distribution data from Achterberg (1979), Achterberg (1984b) and Stigenberg and Shaw (2013). Zele longicauda Curtis, 1832 is apparently a synonym of a species of Eubazus (van Achterberg, pers. comm.).

species of Zele excluded from the British and Irish list:

[annulicrus (Thomson, 1895, Meteorus)] Included in Huddleston’s (Huddleston 1978) checklist but we can trace no specimens or literature citations.

Zele albiditarsus

Curtis, 1832

• testaceator Curtis, 1832

• albitarsis (Nees, 1834, Perilitus)

• dispar (Wesmael, 1835, Perilitus)

• calcitrator (Curtis, 1837, Meteorus)

• wesmaeli (Boie, 1850, Perilitus)

• testaceatrix Schulz, 1906

Distribution

England, Scotland, Wales, Ireland

Zele caligatus

(Haliday, 1835)

• Meteorus caligatus Haliday, 1835

• neesii (Ruthe, 1862, Meteorus)

• alaskensis (Ashmead, 1902, Dyscoletes)

• sibiricus (Fahringer, 1930, Meteorus)

Distribution

England, Scotland, Wales, Ireland

Zele chlorophthalmus

(Spinola, 1808)

• Bracon chlorophthalmus Spinola, 1808

• chrysophthalmus (Nees, 1811, Bracon)

• pallidus (Nees, 1811, Bracon); synonymy by Stigenberg and Ronquist (2011)

• nudator (Thunberg, 1824, Ichneumon)

• splendens (Costa, 1884, Meteorus)

• nigricollis (Thomson, 1895, Meteorus)

Distribution

England, Scotland, Ireland

Zele deceptor

(Wesmael, 1835)

• Perilitus deceptor Wesmael, 1835

• pallitarsis (Cresson, 1872, Perilitus)

• rufulus (Thomson, 1895, Meteorus)

• palliditarsis (Dalla Torre, 1898, Meteorus)

• maximus (Muesebeck, 1923, Meteorus)

• reticulatus (Muesebeck, 1923, Meteorus)

• romani (Fahringer, 1930, Meteorus)

• separandus (Fischer, 1957, Meteorus)

• metallicus (Jakimavičius, 1972, Apanteles)

Distribution

England, Scotland, Wales, Ireland

Microgastrinae

Microgastrinae

Förster, 1863

Notes

The generic classification of Microgastrinae broadly follows Papp (1988), based on Mason’s (Mason 1981) phylogenetic treatment of microgastrines that split up the large genus ‘Apanteles’, which in turn built upon Nixon’s (Nixon 1965) assignment of ‘Apanteles’ into species groups. We do not follow van Achterberg’s (Achterberg 2003c) recent generic revision (reflected in Fauna Europaea), which reassigned many genera to Apanteles and Protapanteles. We consider this reclassification to be premature, being based on very little explicit character evidence and, with its concentration on western Palaearctic species, not really addressing the affinities of most of the world’s microgastrine species. However, there are undoubted merits to van Achterberg’s treatment as not all of the genera currently employed are well-defined. The following genera are recognised as valid by van Achterberg (2003c): Apanteles (=Choeras, Dolichogenidea, Iconella, Illidops, Pholetesor), Cotesia, Deuterixys, Diolcogaster, Hygroplitis, Microgaster, Microplitis, Paroplitis and Protapanteles (=Distatrix, Glyptapanteles, Rasivalva, Sathon). Some distribution data taken from Nixon (1965), Nixon (1968), Nixon (1970), Nixon (1972), Nixon (1973), Nixon (1974), Nixon (1976), remainder from Shaw (2012) and NMS.

Apantelini

Viereck, 1918

Apanteles

Förster, 1863

• UROGASTER Ashmead, 1898

• XESTAPANTELES Cameron, 1910

• ALLAPANTELES Brèthes, 1915

• AREOLATUS Rao & Chalikwar, 1976 unavailable

Notes

The generic placement of several species treated by Nixon (1973) in his metacarpalis group, here largely apportioned between Apanteles and Dolichogenidea following Papp (1988), is questionable (cf. Achterberg 2003c).

species of Apanteles excluded from the British and Irish list:

[anomalon (Curtis, 1830, Microgaster)] This name appeared in Huddleston (1978) but is not listed by Papp (1988) or Achterberg (2003c) and remains uninterpreted.

[nigripes (Ratzeburg, 1844, Microgaster)] This name appeared in Huddleston (1978) but is not listed by Papp (1988) or Achterberg (2003a).

[picipes (Bouché, 1834, Microgaster)] Papp (1987) intended to deal with this name, with a footnote in that paper saying that it would be dealt with under A. xanthostigma, but there is no mention there of picipes. It is presumed that the name picipes is a synonym or a nomen dubium; it is not listed as a valid species by Achterberg (2003c).

Apanteles atreus

Nixon, 1973

Distribution

England

Apanteles brunnistigma

Abdinbekova, 1969

• sotades Nixon, 1976

Distribution

England, Scotland, Wales, Isle of Man

Apanteles carpatus

(Say, 1836)

• Microgaster carpata Say, 1836

• solitarius (Ashmead,1900)

• hawaiiensis (Ashmead, 1901)

• fuscicornis (Cameron, 1910)

• piceoventris Muesebeck, 1921

• igae Watanabe, 1932

• sarcitorius Telenga, 1955

• ultericus Telenga, 1955

Distribution

England

Apanteles chrysis

Nixon, 1973

Distribution

England

Apanteles contaminatus

(Haliday, 1834)

• Microgaster contaminatus Haliday, 1834

Distribution

Scotland, Ireland

Apanteles corvinus

Reinhard, 1880

• lucidus Szépligeti, 1896

• rasteratus Fahringer, 1936

• aptus Papp, 1977

Distribution

England, Scotland

Apanteles galleriae

Wilkinson, 1932

Distribution

England

Notes

added by Shaw (2012)

Apanteles lacteus

(Nees, 1834)

• Microgaster lacteus Nees, 1834

Distribution

England

Notes

We follow Mason (1981) in placing lacteus in Apanteles rather than Papp’s (Papp 1988) placement in Dolichogenidea.

Apanteles lenea

Nixon, 1976

Distribution

England, Scotland, Ireland

Apanteles metacarpalis

(Thomson, 1895)

• Microgaster metacarpalis Thomson, 1895

Distribution

England, Ireland

Apanteles miramis

Nixon, 1976

Distribution

England

Apanteles obscurus

(Nees, 1834)

• Microgaster obscura Nees, 1834

• arenarius (Haliday, 1834)

Distribution

England, Ireland

Apanteles sodalis

(Haliday, 1834)

• Microgaster sodalis Haliday, 1834

• carbonarius (Ratzeburg, 1848) preocc.

• ater (Ratzeburg, 1852); synonymy by Achterberg (1997)

• lugens (Ratzeburg, 1852)

• lindbergi Hedqvist, 1965

Distribution

England

Apanteles xanthostigma

(Haliday, 1834)

• Microgaster xanthostigma Haliday, 1834

• ochrostigma (Wesmael, 1837)

• xanthocarpus Szépligeti, 1901

Distribution

England, Scotland, Wales, Isle of Man

Choeras

Mason, 1981

Choeras arene

(Nixon, 1973)

• Apanteles arene Nixon, 1973

Distribution

England, Scotland, Ireland

Choeras dorsalis

(Spinola, 1808)

• Microgaster dorsalis Spinola, 1808

• cruciatus (Ratzeburg, 1844)

• suffolciensis (Morley, 1902)

Distribution

England, Wales

Choeras parasitellae

(Bouché, 1834)

• Microgaster parasitellae Bouché, 1834

• adjuncta misident.

• flavilabris (Ratzeburg, 1844)

• rufilabris (Ratzeburg, 1844)

• lictorius (Reinhard, 1880)

• polypori (Gautier & Bonnamour, 1930)

Distribution

England, Scotland

Notes

Listed under Apanteles and Microgaster in Huddleston (1978).

Choeras ruficornis

(Nees, 1834)

• Microgaster ruficornis Nees, 1834

• hedymeles (Nixon, 1973)

Distribution

England

Choeras tedellae

(Nixon, 1961)

• Apanteles tedellae Nixon, 1961

• epinotiae (Fischer, 1962)

• epinoticida (Fischer, 1966)

Distribution

England

Choeras tiro

(Reinhard, 1880)

• Microgaster tiro Reinhard, 1880

Distribution

England

Choeras validus

(Thomson, 1895)

• Apanteles validus Thomson, 1895

Distribution

England

Dolichogenidea

Viereck, 1911

Notes

species of Dolichogenidea excluded from the British and Irish list:

[anarsiae (Faure & Alabouvette, 1924, Apanteles)] Listed as a British species by Huddleston (1978) in error, there are no British records.

[cerialis (Nixon, 1976, Apanteles); syn. areolaris (Balevski & Tobias, 1980, Apanteles) preocc.] Listed as a British species by Huddleston (1978) in error, only known from southern and Eastern Europe and Israel.

[ensiformis (Ratzeburg, 1844, Microgaster)] Listed as a British species by Huddleston (1978) in error (see note under Napamus vipio).

[evonymellae (Bouché, 1834, Microgaster); syn. iarbas (Nixon, 1972, Apanteles)] Listed as a British species by Huddleston (1978) in error, there are no British records.

[impura (Nees, 1834, Microgaster)] Notwithstanding Papp (1978), we regard this name as uncertainly interpreted, but in any case we have not seen British material that conforms to Papp's interpretation.

Dolichogenidea agilla

(Nixon, 1972)

• Apanteles agilla Nixon, 1972

• piratica (Papp, 1977)

Distribution

England

Notes

added by Shaw (2012)

Dolichogenidea annularis

(Haliday, 1834)

• Microgaster annularis Haliday, 1834

Distribution

England

Dolichogenidea appellator

(Telenga, 1949)

• Apanteles appellator Telenga, 1949

• litae (Nixon, 1972)

Distribution

England

Notes

Added by Shaw (2012). Papp (1988) notes, but does not follow, the view that appellator may be the valid name (cf. Kotenko and Tobias 1986). Shaw (2012) has found that reared ‘appellator’ and ‘litae’ appear to be conspecific. The situation is complicated by Nixon's (Nixon 1972) treatment of some series (from a different host in Cyprus, and from Egypt) as ‘litae var operculellae’, and it is this that Papp (1988)lists as a junior synonym of appellator.

Dolichogenidea artissima

(Papp, 1971)

• Apanteles artissimus Papp, 1971

• abila (Nixon, 1972)

Distribution

England, Scotland, Wales

Dolichogenidea ate

(Nixon, 1973)

• Apanteles ate Nixon, 1973

Distribution

England

Dolichogenidea bres

(Nixon, 1973)

• Apanteles bres Nixon, 1973

Distribution

England

Dolichogenidea breviventris

(Ratzeburg, 1848)

• Microgaster breviventris Ratzeburg, 1848

• mesoxantha (Ruschka, 1917)

• nilae (Telenga, 1961)

Distribution

England, Scotland, Wales, Ireland

Dolichogenidea britannica

(Wilkinson, 1941)

• Apanteles britannicus Wilkinson, 1941

Distribution

England

Dolichogenidea candidata

(Haliday, 1834)

• Microgaster candidatus Haliday, 1834

• longicauda (Wesmael, 1837); synonymy by Achterberg (1997)

• terebrator (Ratzeburg, 1852)

Distribution

England, Scotland, Wales, Isle of Man

Dolichogenidea coleophorae

(Wilkinson, 1938)

• Apanteles coleophorae Wilkinson, 1938

Distribution

England

Dolichogenidea coniferae

(Haliday, 1834)

• Microgaster coniferae Haliday, 1834

Distribution

England, Isle of Man

Notes

Revised status (Shaw 2012): inadvertently listed as a synonym of candidata by Achterberg (1997) due to a drafting error (van Achterberg, pers. comm.).

Dolichogenidea credne

(Nixon, 1973)

• Apanteles credne Nixon, 1973

Distribution

England

Dolichogenidea cytherea

(Nixon, 1972)

• Apanteles cytherea Nixon, 1972

Distribution

England

Dolichogenidea decora

(Haliday, 1834)

• Microgaster decorus Haliday, 1834

• lineata (Reinhard, 1880)

• sibirica Fahringer, 1938

Distribution

Ireland

Dolichogenidea dilecta

(Haliday, 1834)

• Microgaster dilectus Haliday, 1834

• femoralis (Bouché, 1834)

Distribution

England, Isle of Man

Dolichogenidea drusilla

(Nixon, 1972)

• Apanteles drusilla Nixon, 1972

Distribution

England

Dolichogenidea emarginata

(Nees, 1834)

• Microgaster emarginatus Nees, 1834

• scapularis (Bouché,1834)

Distribution

England, Scotland, Wales

Dolichogenidea exilis

(Haliday, 1834)

• Microgaster exilis Haliday, 1834

Distribution

England

Notes

Shaw (2012) gives a diagnosis; not treated by Nixon or Papp.

Dolichogenidea faucula

(Nixon, 1972)

• Apanteles faucula Nixon, 1972

Distribution

England

Dolichogenidea gagates

(Nees, 1834)

• Microgaster gagates Nees, 1834

Distribution

England

Dolichogenidea glabra

(Papp, 1978)

• Apanteles glaber Papp, 1978

Distribution

England, Scotland

Notes

added by Shaw (2012)

Dolichogenidea gracilariae

(Wilkinson, 1940)

• Apanteles gracilariae Wilkinson, 1940

Distribution

England, Isle of Man

Dolichogenidea halidayi

(Marshall, 1872)

• Apanteles halidayi Marshall, 1872

• albipennis (Haliday, 1834) preocc.

• halidaii misspelling

Distribution

England, Scotland, Isle of Man

Dolichogenidea hilaris

(Haliday, 1834)

• Microgaster hilaris Haliday, 1834

Distribution

Ireland

Dolichogenidea imperator

(Wilkinson, 1939)

• Apanteles imperator Wilkinson, 1939

Distribution

England, Scotland, Wales, Isle of Man

Dolichogenidea infima

(Haliday, 1834)

• Microgaster infimus Haliday, 1834

Distribution

England

Dolichogenidea lacteicolor

(Viereck, 1911)

• Apanteles lacteicolor Viereck, 1911

• conspersae (Fiske, 1911)

Distribution

England

Dolichogenidea lacteipennis

(Curtis, 1830)

• Microgaster lacteipennis Curtis, 1830

• lissonota (Tobias, 1964)

Dolichogenidea laevigata

(Ratzeburg, 1848)

• Microgaster laevigatus Ratzeburg, 1848

• hoplites (Ratzeburg, 1848)

• calcarata (Ivanov, 1899)

Distribution

England, Scotland, Wales

Dolichogenidea laevigatoides

(Nixon, 1972)

• Apanteles laevigatoides Nixon, 1972

Distribution

England

Dolichogenidea laevissima

(Ratzeburg, 1848)

• Microgaster laevissimus Ratzeburg, 1848

• tersa (Papp, 1973)

Distribution

England

Dolichogenidea lemariei

(Nixon, 1961)

• Apanteles lemariei Nixon, 1961

Distribution

England

Dolichogenidea lineipes

(Wesmael, 1837)

• Microgaster lineipes Wesmael, 1837

Distribution

England, Scotland, Isle of Man

Dolichogenidea longicalcar

(Thomson, 1895)

• Apanteles longicalcar Thomson, 1895

Distribution

England

Dolichogenidea longipalpis

(Reinhard, 1880)

• Apanteles longipalpis Reinhard, 1880

• tadzhica (Telenga, 1949)

Distribution

England, Scotland

Notes

Papp (1981) reports that the type series of tadzhica belongs to two species (lacteus and longipalpis) but did not select a lectotype; Belokobylskij et al. (2003) treated the name as a synonym of longipalpis.

Dolichogenidea marica

(Nixon, 1972)

• Apanteles marica Nixon, 1972

Distribution

England

Dolichogenidea myron

(Nixon, 1973)

• Apanteles myron Nixon, 1973

Distribution

England, Scotland

Notes

Transferred from Apanteles in anticipation of publication by Jose Fernandez-Triana.

Dolichogenidea ononidis

(Marshall, 1889)

• Apanteles ononidis Marshall, 1889

Distribution

England

Notes

Shaw (2012) gives a diagnosis; not treated by Nixon or Papp.

Dolichogenidea petrovae

(Walley, 1937)

• Apanteles petrovae Walley, 1937

• dioryctriae (Wilkinson, 1938)

• magna (Telenga, 1955)

• murinanae (Čapek & Zwölfer, 1957)

Distribution

England

Notes

added by Shaw (2012)

Dolichogenidea phaloniae

(Wilkinson, 1940)

• Apanteles phaloniae Wilkinson, 1940

Distribution

England, Scotland, Wales, Ireland

Dolichogenidea phaola

(Nixon, 1972)

• Apanteles phaola Nixon, 1972

Distribution

England

Dolichogenidea praetor

(Marshall, 1885)

• Apanteles praetor Marshall, 1885

Distribution

England

Dolichogenidea princeps

(Wilkinson, 1941)

• Apanteles princeps Wilkinson, 1941

Distribution

England, Scotland, Wales

Dolichogenidea punctiger

(Wesmael, 1837)

• Microgaster punctiger Wesmael, 1837

• itea (Nixon, 1972)

Distribution

England, Scotland

Dolichogenidea sicaria

(Marshall, 1885)

• Apanteles sicarius Marshall, 1885

• chrysosticta (Marshall, 1899)

• crudelis (Papp, 1971)

Distribution

England, Scotland

Dolichogenidea sisenna

(Nixon, 1972)

• Apanteles sisenna Nixon, 1972

Distribution

England

Notes

Recorded as British by Nixon (1972) but omitted by Huddleston (1978).

Dolichogenidea soikai

(Nixon, 1972)

• Apanteles soikai Nixon, 1972

Distribution

England

Notes

added by Shaw (2012)

Dolichogenidea trachala

(Nixon, 1965)

• Apanteles trachalus Nixon, 1965

• sevocata (Papp, 1975)

Distribution

England, Scotland, Wales, Ireland

Notes

#A parasitoid of synanthropic Lepidoptera species and probably introduced to Britain and Ireland (Nixon 1976).

Dolichogenidea ultor

(Reinhard, 1880)

• Apanteles ultor Reinhard, 1880

• lactipennis (Ratzeburg, 1852) preocc.

Distribution

England

Dolichogenidea victor

(Wilkinson, 1941)

• Apanteles victor Wilkinson, 1941

Distribution

England

Illidops

Mason, 1981

Illidops butalidis

(Marshall, 1888)

• Apanteles butalidis Marshall, 1888

Distribution

England, Scotland

Illidops naso

(Marshall, 1885)

• Apanteles naso Marshall, 1885

• contortus (Tobias, 1964)

• crantor (Nixon, 1965)

• evander (Nixon, 1965)

• coresia (Nixon, 1973)

Distribution

England

Illidops suevus

(Reinhard, 1880)

• Apanteles suevus Reinhard, 1880

• minutus (Szépligeti, 1896)

• polonicus (Fahringer, 1936)

• brevisternis (Tobias, 1964)

• suspicax (Tobias, 1964)

• dion (Nixon, 1965)

• sesostris (Nixon, 1976)

Distribution

England

Napamus

Papp, 1993

Notes

species of Napamus excluded from the British and Irish list:

[vipio (Reinhard, 1880, Apanteles)] Mistakenly listed as a British species by Kloet and Hincks (1945) and Huddleston (1978). Shenefelt (1973) listed it as British on the misunderstanding that Morley and Rait-Smith (1933) had produced a catalogue of British Lepidoptera-parasitoid asscociations, whereas their listings also included non-British rearings from hosts that occur in Britain. Morley and Rait-Smith (1933) cite Marshall (1896) as the source of host records for vipio, who does not mention Britain. Apanteles vipio was transferred from Illidops by Papp (1993).

Pholetesor

Mason, 1981

Pholetesor arisba

(Nixon, 1973)

• Apanteles arisba Nixon, 1973

Distribution

England, Scotland

Pholetesor bicolor

(Nees, 1834)

• Microgaster bicolor Nees, 1834

• ardeaepenellae (Bouché, 1834)

• pedias (Nixon, 1973)

• umbellatarum (Haliday, 1834)

• schillei (Niezabitowski, 1910)

• longicauda (Fahringer, 1938)

Distribution

England

Notes

Some distribution data from Shaw and Askew (1976). Van Achterberg’s (Achterberg 1997) synonymy of bicolor under circumscriptus is not followed here (Shaw 2012); it is probable that his reared series represents two species, bicolor and circumscriptus.

Pholetesor circumscriptus

(Nees, 1834)

• Microgaster circumscriptus Nees, 1834

• exiguus (Haliday, 1834); synonymy by Achterberg (1997)

• blancardellae (Bouché, 1834)

• lividipes (Wesmael, 1837)

• flavolimbatus (Ratzeburg, 1848)

• lautellus (Marshall, 1895)

Distribution

England, Scotland, Wales, Ireland

Pholetesor elpis

(Nixon, 1973)

• Apanteles elpis Nixon, 1973

• girkanus (Tobias, 1976)

Distribution

England

Pholetesor errans

(Nixon, 1973)

• Apanteles errans Nixon, 1973

• arenicola (Papp, 1973)

Distribution

England

Pholetesor laetus

(Marshall, 1885)

• Apanteles laetus Marshall, 1885

• exiguus misident.

• salalicus misident.

• metallicus (Jakimavičius, 1972)

Distribution

England, Scotland

Notes

Shaw (2012) tentatively identified exiguus sensu Nixon (1973) as an extreme of morphological variation within laetus.

Pholetesor maritimus

(Wilkinson, 1941)

• Apanteles maritimus Wilkinson, 1941

Distribution

England, Scotland, Wales

Pholetesor moldavicus

(Tobias, 1975)

• Apanteles moldavicus Tobias, 1975

Distribution

England

Notes

Added by Shaw (2012). Although this generic placement (Papp 1988) was followed by Shaw (2012) is seems very likely to be inappropriate.

Pholetesor nanus

(Reinhard, 1880)

• Apanteles nanus Reinhard, 1880

• szoecsi (Papp, 1973)

Distribution

England, Scotland, Wales, Isle of Man

Pholetesor phaetusa

(Nixon, 1973)

• Apanteles phaetusa Nixon, 1973

Distribution

England, Scotland

Pholetesor viminetorum

(Wesmael, 1837)

• Microgaster viminetorum Wesmael, 1837

• fuliginosus (Wesmael, 1837)

Distribution

England, Scotland, Wales, Ireland

Cotesiini

Mason, 1981

Cotesia

Cameron, 1891

• CRYPTAPANTELES Viereck, 1910

• STENOPLEURA Viereck, 1911

Notes

Some taxonomic and distribution data for species parasitizing Lycaenidae taken from Shaw (2007).

species of Cotesia excluded from the British and Irish list:

[acuminata (Reinhard, 1880, Apanteles); cultrator (Marshall, 1885, Apanteles)] Not a British or Irish species. Marshall’s description was based on unprovenanced material reared from a host that, on inspection by MRS, is almost certainly not British. The current synonymy is also in doubt.

[ordinaria (Ratzeburg, 1844, Microgaster); syn. dendrolimi (Matsumura, 1926, Apanteles); dendrolimusi (Matsumura, 1926, Apanteles)] Although listed as a British species by various authors we can find no evidence that it has occurred here.

[saltator (Thunberg, 1824, Ichneumon) preocc.] Appeared in Huddleston (1978) as Roman (1912) incorrectly synonymised tenebrosa under saltator.

[scabricula (Reinhard, 1880, Apanteles); syn. eguchii (Watanabe, 1935, Apanteles)] No evidence that this is a British or Irish species.

Cotesia abjecta

(Marshall, 1885)

• Apanteles abjectus Marshall, 1885

• complanata (Lyle, 1916, Apanteles)

Distribution

England, Scotland

Cotesia affinis

(Nees, 1834)

• Microgaster affinis Nees, 1834

• euphorbiae (Bouché, 1834, Microgaster)

• vinulae (Bouché, 1834, Microgaster)

• harpyiae (Niezabitowski, 1910, Apanteles)

• okamotoi (Watanabe, 1932, Apanteles)

• planus (Watanabe, 1932, Apanteles)

Distribution

England

Cotesia analis

(Nees, 1834)

• Microgaster analis Nees, 1834

• praetextata (Haliday, 1834, Microgaster)

• mediana (Ratzeburg, 1852, Microgaster)

• leucaniae (Wikinson, 1937, Apanteles)

Distribution

England

Cotesia astrarches

(Marshall, 1889)

• Apanteles astrarches Marshall, 1889

• Cotesia astrarches ?arctica (Thomson, 1895, Apanteles)

• genalis (Tobias, 1964, Apanteles)

Distribution

England, Scotland, Wales

Notes

Nixon’s (Nixon 1974) arctica encompassed two species that occur in Britain, astrarches and tenebrosa. It is not clear which, if either, the name arctica is associated with. The name astrarches was placed, erroneosly, in synonymy with arctica by Nixon (1974), which was followed by Huddleston (1978). Shaw (2007) clarified the status of astrarches.

Cotesia bignellii

(Marshall, 1885)

• Apanteles bignellii Marshall, 1885

Distribution

England, Scotland, Wales, Ireland

Cotesia brevicornis

(Wesmael, 1837)

• Microgaster brevicornis Wesmael, 1837

• cleoceridis (Marshall, 1889, Apanteles)

Distribution

England, Scotland

Cotesia cajae

(Bouché, 1834)

• Microgaster cajae Bouché, 1834

• difficilis (Nees, 1834, Microgaster)

Distribution

England

Cotesia callimone

(Nixon, 1974)

• Apanteles callimone Nixon, 1974

• scelerata (Tobias, 1986, Apanteles)

Distribution

Scotland, Wales, Ireland

Cotesia chares

(Nixon, 1965)

• Apanteles chares Nixon, 1965

Distribution

England

Cotesia cleora

(Nixon, 1974)

• Apanteles cleora Nixon, 1974

Distribution

England

Cotesia coryphe

(Nixon, 1974)

• Apanteles coryphe Nixon, 1974

Distribution

England

Notes

Papp (1987) synonymised coryphe under rubripes but this seems unwarranted, given the distinctly different hosts and other aspects of biology.

Cotesia cuprea

(Lyle, 1925)

• Apanteles cupreus Lyle, 1925

Distribution

England, Scotland

Cotesia errator

(Nixon, 1974)

• Apanteles errator Nixon, 1974

Distribution

England, Wales

Cotesia eulipis

(Nixon, 1974)

• Apanteles eulipis Nixon, 1974

• eulipsis misspelling

Distribution

England, Scotland

Cotesia euryale

(Nixon, 1974)

• Apanteles euryale Nixon, 1974

Distribution

England

Notes

It is not clear from Nixon (1974) that this species occurs in Britain, despite its being ‘bred in captivity at Slough’.

Cotesia ferruginea

(Marshall, 1885)

• Apanteles ferrugineus Marshall, 1885

Distribution

England

Cotesia gades

(Nixon, 1974)

• Apanteles gades Nixon, 1974

Distribution

England

Notes

added by Allen (1978)

Cotesia gastropachae

(Bouché, 1834)

• Microgaster gastropachae Bouché, 1834

Distribution

England, Scotland, Isle of Man

Cotesia geryonis

(Marshall, 1885)

• Apanteles geryonis Marshall, 1885

Distribution

England

Cotesia glomerata

(Linnaeus, 1758)

• Ichneumon glomeratus Linnaeus, 1758

• glomerator (Thunberg, 1824, Ichneumon)

• nigriventris (Nees, 1834, Microgaster)

• recondita (Nees, 1834, Microgaster)

• stellatarum (Bouché, 1834, Microgaster)

• crataegi (Ratzeburg, 1844, Microgaster)

• oleracea (Taylor, 1860, Microgaster)

• pieridis (Packard, 1881, Microgaster) preocc.

• pieridivora (Riley, 1882, Microgaster)

• aporiae (Ivanov, 1899, Apanteles)

• nawaii (Ashmead, 1906, Glyptapanteles)

• aporiae (Matsumura, 1908, Apanteles) preocc.

• heterotergis (Fahringer, 1936, Apanteles)

Distribution

England, Scotland, Isle of Man

Cotesia gonopterygis

(Marshall, 1885)

• Apanteles gonopterygis Marshall, 1885

Distribution

England

Cotesia hyphantriae

(Riley, 1887)

• Apanteles hyphantriae Riley, 1887

Distribution

England, Wales

Cotesia inducta

(Papp, 1973)

• Apanteles inductus Papp, 1973

• tenuivalvis (Tobias, 1986, Apanteles)

Distribution

England, Ireland

Notes

added by Revels (2006); Shaw (2007)

Cotesia isolde

(Nixon, 1974)

• Apanteles isolde Nixon, 1974

Distribution

England, Scotland

Cotesia jucunda

(Marshall, 1885)

• Apanteles jucundus Marshall, 1885

• nigrinervis (Thomson, 1895, Microgaster)

Distribution

England, Scotland

Cotesia juniperatae

(Bouché, 1834)

• Microgaster juniperatae Bouché, 1834

Distribution

England, Scotland

Cotesia kurdjumovi

(Telenga, 1955)

• Apanteles kurdjumovi Telenga, 1955

• laverna (Nixon, 1974, Apanteles)

Distribution

England, Scotland

Cotesia limbata

(Marshall, 1885)

• Apanteles limbatus Marshall, 1885

• kawadai (Watanabe, 1934, Apanteles)

Distribution

England, Scotland

Cotesia lineola

(Curtis, 1830)

• Microgaster lineola Curtis, 1830

• gabrielis (Gautier & Riel, 1919, Apanteles)

Distribution

England

Cotesia melanoscela

(Ratzeburg, 1844)

• Microgaster melanoscelus Ratzeburg, 1844

• solitaria (Ratzeburg, 1844, Microgaster)

• creata (Balevski, 1980, Apanteles)

Distribution

England

Cotesia melitaearum

(Wilkinson, 1937)

• Apanteles melitaearum Wilkinson, 1937

• melittaearum misspelling

• ukrainica (Tobias, 1986, Apanteles)

Distribution

England, Scotland, Wales

Cotesia notha

(Marshall, 1885)

• Apanteles nothus Marshall, 1885

Distribution

England, Scotland

Cotesia numen

(Nixon, 1974)

• Apanteles numen Nixon, 1974

Distribution

England, Scotland

Cotesia ofella

(Nixon, 1974)

• Apanteles ofella Nixon, 1974

• Cotesia ofella ?perspicua (Nees, 1834, Microgaster)

Distribution

England

Notes

Microgaster perspicua is listed as a senior synonym of cajae in Taxapad (Yu et al. 2012), following Marshall (1885), who listed cajae as the senior synonym. Papp (2005a) lists perspicua as a tentative synonym (which would have priority) of ofella.

Cotesia onaspis

(Nixon, 1974)

• Apanteles onaspis Nixon, 1974

• avetyanae (Tobias, 1976, Apanteles)

Distribution

England

Cotesia orestes

(Nixon, 1974)

• Apanteles orestes Nixon, 1974

Distribution

England

Cotesia pilicornis

(Thomson, 1890)

• Microgaster pilicornis Thomson, 1890

• piliflagellaris (Tobias, 1986, Apanteles)

Distribution

England, Scotland, Wales, Ireland

Notes

In NMS numerous series reared solitarily from various species of Pterophoridae are probably conspecific but show a great variation in extent of pilosity of the antenna.

Cotesia praepotens

(Haliday, 1834)

• Microgaster praepotens Haliday, 1834

• placida (Haliday, 1834, Microgaster)

• memnon (Nixon, 1974, Apanteles)

• acutivalvis (Balevski, 1980, Apanteles)

• beshtaui (Tobias, 1986, Apanteles)

Distribution

England, Ireland

Notes

Nixon’s (Nixon 1974) praepotens is apparently correctly called sericea (Belokobylskij et al. 2003).

Cotesia risilis

(Nixon, 1974)

• Apanteles risilis Nixon, 1974

Distribution

England

Cotesia rubecula

(Marshall, 1885)

• Apanteles rubecula Marshall, 1885

Distribution

England

Cotesia rubripes

(Haliday, 1834)

• Microgaster rubripes Haliday, 1834

Distribution

England, Scotland

Cotesia ruficrus

(Haliday, 1834)

• Microgaster ruficrus Haliday, 1834

• antipoda (Ashmead, 1900, Apanteles)

• manilae (Ashmead, 1904, Apanteles)

• sydneyensis (Cameron, 1911, Apanteles)

• narangae (Viereck, 1913, Apanteles)

• sesamiae (Risbec, 1956, Apanteles) nom. nud.

Distribution

England, Wales, Isle of Man

Cotesia salebrosa

(Marshall, 1885)

• Apanteles salebrosus Marshall, 1885

• callunae Nixon, 1974

Distribution

England, Scotland

Cotesia saltatoria

(Balevski, 1980)

• Apanteles saltatorius Balevski, 1980

Distribution

England, Scotland

Notes

added by Shaw (2007)

Cotesia sericea

(Nees, 1834)

• Microgaster sericeus Nees, 1834

• praepotens misident.

• brachycera (Thomson, 1895, Apanteles)

Distribution

England, Scotland

Notes

Nixon’s (Nixon 1974) praepotens is apparently correctly called sericea (Belokobylskij et al. 2003).

Cotesia sibyllarum

(Wilkinson, 1936)

• Apanteles sibyllarum Wilkinson, 1936

Distribution

England

Cotesia spuria

(Wesmael, 1837)

• Microgaster spurius Wesmael, 1837

• insidens (Ratzeburg, 1844, Microgaster)

Distribution

England, Scotland, Wales, Isle of Man

Cotesia subordinaria

(Tobias, 1976)

• Apanteles subordinarius Tobias, 1976

Distribution

England

Notes

added by Shaw (2012)

Cotesia telengai

(Tobias, 1972)

• Apanteles telengai Tobias, 1972

• amabilis (Nixon, 1974, Apanteles)

Distribution

England

Cotesia tenebrosa

(Wesmael, 1837)

• Microgaster tenebrosus Wesmael, 1837

• Cotesia tenebrosa ?arctica (Thomson, 1895, Apanteles)

Distribution

England, Scotland

Notes

Added by Shaw (2007); Nixon’s (Nixon 1974) arctica encompassed two species that occur in Britain, astrarches and tenebrosa. It is not clear which, if either, the name arctica is associated with. The name astrarches was placed, erroneosly, in synonymy with arctica by Nixon (1974), which was followed by Huddleston (1978). Shaw (2007) clarified the status of tenebrosa.

Cotesia tetrica

(Reinhard, 1880)

• Apanteles tetricus Reinhard, 1880

• opacula (Thomson, 1895, Microgaster)

Distribution

England, Scotland

Cotesia tibialis

(Curtis, 1830)

• Microgaster tibialis Curtis, 1830

• atrator (Curtis, 1830, Microgaster)

• gracilis (Curtis, 1830, Microgaster)

• congesta (Nees, 1834, Microgaster)

• intricata (Haliday, 1834, Microgaster)

• gracilipes (Thomson, 1895, Microgaster)

• similis (Szépligeti, 1901, Apanteles)

• atratrix (Schulz, 1906, Microgaster)

• aranearum (Goureau, 1908, Apanteles) nom. nud.

• mamestrae (Matsumura, 1908, Apanteles)

• simulans (Lyle, 1917, Apanteles)

• claustrata (Gautier & Bonnamour, 1923, Apanteles)

Distribution

England, Scotland, Wales

Cotesia vanessae

(Reinhard, 1880)

• Apanteles vanessae Reinhard, 1880

Distribution

England

Cotesia vestalis

(Haliday, 1834)

• Microgaster vestalis Haliday, 1834

• plutellae (Kurdjumov, 1912, Apanteles); synonymy by Shaw (2003b)

Distribution

England, Wales

Cotesia villana

(Reinhard, 1880)

• Apanteles villanus Reinhard, 1880

• fasciatae (Gautier & du Dresnay, 1926, Apanteles)

• rubroides (Papp, 1971, Apanteles); synonymy by Papp (2009a)

Distribution

England

Cotesia zygaenarum

(Marshall, 1885)

• Apanteles zygaenarum Marshall, 1885

Distribution

England, Scotland, Wales, Ireland, Isle of Man

Deuterixys

Mason, 1981

Deuterixys carbonaria

(Wesmael, 1837)

• Microgaster carbonarius Wesmael, 1837

• anomala (Lyle, 1925, Apanteles)

Distribution

England, Scotland

Deuterixys plugarui

(Tobias, 1975)

• Apanteles plugarui Tobias, 1975

Distribution

England

Notes

added by Shaw (2012)

Deuterixys rimulosa

(Niezabitowski, 1910)

• Apanteles rimulosus Niezabitowski, 1910

• comes (Wilkinson, 1940, Apanteles)

Distribution

England

Diolcogaster

Ashmead, 1901

• PROTOMICROPLITIS misident.

• ZADIOLOCOGASTER Viereck, 1913

Notes

See Mason (1981) and Fernández-Triana (2015), who restricted Protomicroplitis to a few Nearctic and Neotropical species.

Diolcogaster abdominalis

(Nees, 1834)

• Microgaster abdominalis Nees, 1834

Distribution

England

Diolcogaster alvearia

(Fabricius, 1798)

• Ichneumon aleuarius Fabricius, 1798

• aphidum (Panzer, 1804, Ichneumon)

• alveator (Thunberg, 1824, Ichneumon)

• areolata (Szépligeti, 1896, Ichneumon)

Distribution

England, Scotland

Diolcogaster connexa

(Nees, 1834)

• Microgaster connexus Nees, 1834

• consularis (Haliday, 1834, Microgaster)

• diluta (Ratzeburg, 1852, Microgaster)

Distribution

England, Isle of Man

Diolcogaster flavipes

(Haliday, 1834)

• Microgaster flavipes Haliday, 1834

Diolcogaster hinzi

(Nixon, 1965)

• Protomicroplitis hinzi Nixon, 1965

Distribution

England, Scotland

Notes

added by Shaw (2012)

Diolcogaster minuta

(Reinhard, 1880)

• Microgaster minutus Reinhard, 1880

Distribution

England, Scotland

Diolcogaster scotica

(Marshall, 1885)

• Microgaster scoticus Marshall, 1885

Distribution

England, Scotland

Diolcogaster spreta

(Marshall, 1885)

• Microgaster spretus Marshall, 1885

Distribution

England, Scotland, Wales

Distatrix

Mason, 1981

Distatrix formosa

(Wesmael, 1837)

• Microgaster formosus Wesmael, 1837

• marshallii (Bignell, 1901, Apanteles)

Distribution

England, Wales, Isle of Man

Glyptapanteles

Ashmead, 1905

Notes

species of Glyptapanteles excluded from the British and Irish list:

[thompsoni (Lyle, 1917, Apanteles)] Listed as a British species by Huddleston (1978) in error; there is no evidence that it has occurred here.

Glyptapanteles acasta

(Nixon, 1973)

• Apanteles acasta Nixon, 1973

Distribution

England

Glyptapanteles aliphera

(Nixon, 1973)

• Apanteles aliphera Nixon, 1973

• aliphaera misspelling

• sublateralis (Tobias, 1976, Apanteles)

Distribution

England, Scotland

Glyptapanteles callidus

(Haliday, 1834)

• Microgaster callidus Haliday, 1834

• urolus (Papp, 1983, Apanteles)

Distribution

England, Scotland, Wales

Notes

Achterberg (1997) reinterpreted this name and treated majalis as the valid name for the species called callidus by Nixon (1973) and Papp (1983).

Glyptapanteles compressiventris

(Muesebeck, 1921)

• Apanteles compressiventris Muesebeck, 1921

• liparidis misident.

Distribution

Scotland, Wales

Notes

Wilkinson (1945) dealt with the identity of the real liparidis, which does not occur in Britain or Ireland.

Glyptapanteles fausta

(Nixon, 1973)

• Apanteles fausta Nixon, 1973

Distribution

England, Scotland

Notes

Generic placement in doubt; some authors place both fausta and lateralis in Sathon. Papp (1983) synonymised fausta under Apanteles eugeni Papp, 1972 but later (Papp 1988) rescinded his action. Unfortunately Papp (1972) had chosen a non-reared specimen as holotype but a paratype of eugeni reared from Anthophila fabriciana (Linnaeus) is in BMNH and appears to be conspecific with fausta (described from this host). It seems probable that Papp's (Papp 1983) synonymy was justified but, as this requires further investigation, the name fausta is retained for now.

Glyptapanteles fraternus

(Reinhard, 1881)

• Apanteles fraternus Reinhard, 1881

Distribution

England

Glyptapanteles fulvipes

(Haliday, 1834)

• Microgaster fulvipes Haliday, 1834

Distribution

England, Scotland, Wales, Ireland

Glyptapanteles inclusus

(Ratzeburg, 1844)

• Microgaster inclusus Ratzeburg, 1844

• curvulus (Thomson, 1895, Microgaster)

• rectinervis (Telenga, 1955, Apanteles)

Distribution

England

Glyptapanteles lateralis

(Haliday, 1834)

• Microgaster lateralis Haliday, 1834

Distribution

England, Scotland, Wales

Notes

Generic placement in doubt; some authors place both fausta and lateralis in Sathon.

Glyptapanteles luciana

(Nixon, 1973)

• Apanteles luciana Nixon, 1973

Distribution

Scotland

Notes

Described by Nixon (1973) partly from Scottish material but omitted by Huddleston (1978).

Glyptapanteles majalis

(Wesmael, 1837)

• Microgaster majalis Wesmael, 1837

• callidus misident.

Distribution

England, Scotland

Notes

Achterberg (1997) treated majalis as the valid name for the species called callidus by Nixon (1973) and Papp (1983).

Glyptapanteles menander

(Nixon, 1973)

• Apanteles menander Nixon, 1973

Distribution

Scotland

Glyptapanteles mygdonia

(Nixon, 1973)

• Apanteles mygdonia Nixon, 1973

Distribution

England, Scotland, Ireland

Glyptapanteles octonarius

(Ratzeburg, 1852)

• Microgaster octonarius Ratzeburg, 1852

• stauropodis (Bridgman, 1889, Apanteles)

• lucifugus (Lyle, 1917, Apanteles)

Distribution

England

Glyptapanteles pallipes

(Reinhard, 1880)

• Apanteles pallipes Reinhard, 1880

• pallidipes (Marshall, 1885, Apanteles)

• longicornis (Provancher, 1886, Microgaster)

• radiatus (Ashmead, 1898, Apanteles)

• reinhardi (Wilkinson, 1936, Apanteles)

Distribution

England, Scotland, Wales

Glyptapanteles pinicola

(Lyle, 1917)

• Apanteles pinicola Lyle, 1917

Distribution

England, Scotland

Glyptapanteles porthetriae

(Muesebeck, 1928)

• Apanteles porthetriae Muesebeck, 1928

Distribution

England

Notes

added by Shaw and Skelton (2008)

Glyptapanteles salepus

(Papp, 1983)

• Apanteles salepus Papp, 1983

Distribution

England

Notes

added by Shaw (2012)

Glyptapanteles vitripennis

(Curtis, 1830)

• Microgaster vitripennis Curtis, 1830

• fulcriger (Wesmael, 1837, Microgaster)

• impavidus (Gautier & Dresnyay, 1927, Apanteles)

Distribution

England, Scotland, Isle of Man

Protapanteles

Ashmead, 1898

Protapanteles anchisiades

(Nixon, 1973)

• Apanteles anchisiades Nixon, 1973

Distribution

England, Scotland, Wales, Isle of Man

Protapanteles endemus

(Nixon, 1965)

• Apanteles endemus Nixon, 1965

Distribution

England, Scotland

Protapanteles enephes

(Nixon, 1965)

• Apanteles enephes Nixon, 1965

Distribution

England

Protapanteles hirtariae

(Kotenko & Tobias, 1986)

• Apanteles hirtariae Kotenko & Tobias, 1986

Distribution

Scotland

Notes

added by Shaw (2012)

Protapanteles immunis

(Haliday, 1834)

• Microgaster immunis Haliday, 1834

Distribution

England, Scotland, Wales, Ireland

Protapanteles incertus

(Ruthe, 1859)

• Microgaster incertus Ruthe, 1859

• caberae (Marshall, 1885, Apanteles)

• jugosus (Lyle, 1916, Apanteles)

• mihalyii (Papp, 1973, Apanteles)

Distribution

England, Scotland, Isle of Man

Protapanteles parallelus

(Lyle, 1917)

• Apanteles parallelus Lyle, 1917

• lylei (Shenefelt, 1972, Apanteles)

Distribution

England

Protapanteles popularis

(Haliday, 1834)

• Microgaster popularis Haliday, 1834

Distribution

England, Scotland

Protapanteles triangulator

(Wesmael, 1837)

• Microgaster triangulator Wesmael, 1837

Distribution

England

Notes

Included in Protapanteles by Papp (1988) but Mason (1981) treated it as a species of Glyptapanteles. Nixon (1973) was unable to suggest close relatives.

Rasivalva

Mason, 1981

Rasivalva calceata

(Haliday, 1834)

• Microgaster calceatus Haliday, 1834

• pubescens (Ratzeburg, 1844, Microgaster)

Distribution

England, Scotland

Rasivalva circumvecta

(Lyle, 1918)

• Diolcogaster circumvectus Lyle, 1918

Distribution

England, Scotland

Rasivalva marginata

(Nees, 1834)

• Microgaster marginatus Nees, 1834

Distribution

England, Scotland

Microgastrini

Förster, 1863

Hygroplitis

Thomson, 1895

Hygroplitis pseudorussata

Shaw, 1992

Distribution

England

Notes

added by Shaw (1992b)

Hygroplitis rugulosa

(Nees, 1834)

• Microgaster rugulosus Nees, 1834

• infumata (Haliday, 1834, Microgaster)

• opaca (Ruthe, 1858, Microgaster)

Distribution

England, Wales, Ireland

Hygroplitis russata

(Haliday, 1834)

• Microgaster russatus Haliday, 1834

• dimidiata (Wesmael, 1837, Microgaster)

• basalis (Stephens, 1846, Microgaster)

• aomoriensis (Matsumura, 1910, Microgaster)

Distribution

England, Scotland, Wales

Notes

Specimen from Islay in Hunterian Museum, Glasgow, det. MRS.

Iconella

Mason, 1981

Notes

species of Iconella excluded from the British and Irish list:

[lacteoides (Nixon, 1965, Apanteles); syn. memorabilis (Alexeev, 1971, Apanteles)]

[merula (Reinhard, 1880, Apanteles); syn. ?etiellae (Viereck, 1911, Apanteles)] Both of these species were mistakenly listed by Huddleston (1978); there is no evidence that they are British or Irish.

Iconella aeola

(Nixon, 1965)

• Apanteles aeolus Nixon, 1965

Distribution

England

Notes

added by Shaw (2012)

Microgaster

Latreille, 1804

• LIGANIRA Walker, 1860

• LISSOGASTER Bengtsson, 1926

Notes

The current usage of the name Microgaster was restored by Opinion 1510 (ICZN 1988), after temporarily being applied to the genus here called Microplitis (with Microgaster as currently understood being referred to Lissogaster). This was an unfortunate name change as it coincided with Papp’s (Papp 1976, Papp 1984) revisions of the genera, as well as several other important papers. Unless noted otherwise, distribution data taken from NMS, Nixon (1968) and Shaw (2012). Papp’s (Papp 1976) subsequent revision of the genus added many species to Nixon’s (Nixon 1968) revision, and Achterberg (1997) established the precedence of several Haliday names.

species of Microgaster excluded from the British and Irish list:

[auriculata (Fabricius, 1804, Ichneumon)] Listed as doubtfully British by Huddleston (1978) on the basis of Papp’s (Papp 1976) listing of ‘?England’. No evidence that this is really a British or Irish species.

[deceptor Nixon, 1968] Listed as a British species by Huddleston (1978) in error; no evidence that this is a British or Irish species.

[fischeri Papp, 1960] British specimens, misidentified as fischeri by Nixon (1968), represented an undescribed species which was described by Shaw (2012) as raschkiellae.

[nobilis Reinhard, 1880; syn. compressifemur Fahringer, 1937] Listed as doubtfully British by Huddleston (1978) on the basis of Papp’s (Papp 1976) listing of ‘?England’. No evidence that this is really a British or Irish species.

[postica Nees, 1834; syn. marginella Wesmael, 1837; ?ruficoxis Ruthe, 1858] Recorded as British by Marshall (1885), but probably in error as his diagnosis appears not to have been a Microgaster species in the modern sense. Papp’s (Papp 1976) listing of England probably simply reflects Marshall’s record, as does the listing in Huddleston (1978), and there is no evidence that this is a British or Irish species.

Microgaster acilia

Nixon, 1968

Distribution

England

Notes

Raised from synonymy under meridiana (Papp 1999b) by Shaw (2012).

Microgaster alebion

Nixon, 1968

Distribution

England, Scotland

Notes

distribution data from Shaw (2004)

Microgaster areolaris

Thomson, 1895

Distribution

England, Scotland

Microgaster arctostaphylica

Shaw, 2012

Distribution

Scotland

Notes

added by Shaw (2012)

Microgaster consors

Nixon, 1968

Distribution

England

Microgaster crassicornis

Ruthe, 1860

Distribution

England

Microgaster ductilis

Nixon, 1968

Distribution

England

Notes

added by Shaw (2012)

Microgaster fulvicrus

Thomson, 1895

• striatoscutellaris Kiss, 1927

Distribution

England, Scotland, Ireland, Isle of Man

Microgaster globata

(Linnaeus, 1758)

• Ichneumon globatus Linnaeus, 1758

• laeviscuta Thomson, 1895

• gossypina (Retzius, 1783, Ichneumon)

• globator (Thunberg, 1824, Ichneumon)

• anthomyiarum Bouché, 1834

• amentorum Ratzeburg, 1844

• incurvata Papp, 1976; synonymy by Papp (2002)

Distribution

England, Scotland

Notes

Some distribution data from Papp (1976). It seems that laeviscuta has been included as a junior synonym of hospes in Fauna Europaea, with globata not referred to. Shaw (2012) notes that globata as perceived by Papp (1976) is almost certainly an aggregate.

Microgaster hospes

Marshall, 1885

• comptanae Viereck, 1911

Distribution

England, Scotland, Wales

Microgaster luctuosa

Haliday, 1834

• curvicrus Thomson, 1895; synonymy by Achterberg (1997)

Distribution

England, Scotland, Wales

Microgaster meridiana

Haliday, 1834

• spinolae Haliday, 1834 preocc.; synonymy by Achterberg (1997)

• alexis Haliday, 1834 nom. nud.

• grandis Thomson, 1895; synonymy by Achterberg (1997)

• contubernalis Marshall, 1898

Distribution

England, Scotland, Wales, Ireland, Isle of Man

Microgaster messoria

Haliday, 1834

• tibialis Nees, 1834 preocc.; synonymy by Achterberg (1997)

• vulgaris Ruthe, 1860

• pluto Morley, 1936

Distribution

England, Scotland, Wales, Ireland

Notes

Some Ruthe names traditionally regarded as synonyms of tibialis probably do not belong here Shaw (2012).

Microgaster nigricans

Nees, 1834

Notes

A species of doubtful status. Curtis (1837) listed nigricans as a British species and Papp (1976), who stated that it was known only from the male, therefore listed it as English. Nixon (1968), however, did not deal with the name.

Microgaster nixalebion

Shaw, 2004

Distribution

England, Wales

Notes

added by Shaw (2004)

Microgaster novicia

Marshall, 1885

• swammerdamiae Muesebeck, 1922

Distribution

England, Scotland

Microgaster opheltes

Nixon, 1968

Distribution

Ireland

Microgaster pantographae

Muesebeck, 1922

Distribution

England

Microgaster parvistriga

Thomson, 1895

Distribution

England, Scotland

Microgaster polita

Marshall, 1885

• carinata Bengtsson, 1926 preocc.

• bengtssoni Fahringer, 1937

Distribution

England, Scotland, Ireland

Microgaster procera

Ruthe, 1860

• intermedia Ivanov, 1899

Distribution

Ireland

Microgaster raschkiellae

Shaw, 2012

• fischeri misident.

Distribution

England, Scotland, Wales

Notes

added by Shaw (2012)

Microgaster stictica

Ruthe, 1858

• confusa Papp, 1971

Distribution

England, Scotland, Wales, Isle of Man

Microgaster subcompleta

Nees, 1834

• annulipes Curtis, 1830

• carinata Packard, 1881

Distribution

England, Scotland, Wales, Ireland, Isle of Man

Paroplitis

Mason, 1981

Paroplitis wesmaeli

(Ruthe, 1860)

• Microgaster wesmaeli Ruthe, 1860

• picipes (Wesmael, 1837, Microgaster) preocc.

Distribution

England

Sathon

Mason, 1981

Sathon falcatus

(Nees, 1834)

• Microgaster falcatus Nees, 1834

• equestris (Haliday, 1834, Microgaster)

• gladiator (Szépligeti, 1901, Apanteles)

Distribution

England, Scotland, Wales, Ireland

Microplitini

Mason, 1981

Microplitis

Förster, 1863

• DAPSILOTOMA Cameron, 1906

Notes

Unless noted otherwise, distribution data taken from Nixon (1970). Papp (1984) extensively revised the synonymy (as Microgaster species: unfortunately at that time Microplitis was briefly being regarded as a synonym of Microgaster, but this was subsequently rescinded by Opinion 1510 (ICZN 1988), whose conclusions are largely followed here.

species of Microplitis excluded from the British and Irish list:

[eremitus Reinhard, 1880] No evidence that this species has occurred in Britain or Ireland; probably listed in Shenefelt (1973) and Huddleston (1978) because Nixon (1970) reported its distribution as north-west European.

Microplitis aduncus

(Ruthe, 1860)

• Microgaster aduncus Ruthe, 1860

• brachycerus (Thomson, 1895, Microgaster)

Distribution

Scotland

Microplitis decens

Tobias, 1964

Distribution

Scotland

Notes

Added by Papp (1984) but it is unclear on what basis this species was listed as British (Shaw 2012).

Microplitis deprimator

(Fabricius, 1798)

• Ichneumon deprimator Fabricius, 1798

• ingratus (Haliday, 1834, Microgaster); synonymy by Achterberg (1997)

• sordipes (Nees, 1834, Microgaster)

• tau (Ratzeburg, 1852, Microgaster)

• deprimatrix (Schulz, 1906, Microgaster)

Distribution

England, Scotland

Notes

Micropoltis sordipes is listed as a separate species in Taxapad (Yu et al. 2012) but we follow van Achterberg, in Fauna Europaea. However, the status of sordipes remains uncertain.

Microplitis flavipalpis

(Brullé, 1832)

• Microgaster flavipalpis Brullé, 1832

• ruricola Lyle, 1918

Distribution

England

Microplitis fordi

Nixon, 1970

Distribution

England, Scotland

Notes

Papp (1984) suggests that semicircularis (Ratzeburg, 1844, Microgaster) (type destroyed) may be a senior synonym.

Microplitis fulvicornis

(Wesmael, 1837)

• Microgaster fulvicornis Wesmael, 1837

• calcarata misident.

• pallidicornis Marshall, 1898

Distribution

England, Wales

Microplitis impressus

(Wesmael, 1837)

• Microgaster impressus Wesmael, 1837

• sispes Nixon, 1970

Distribution

England

Notes

added by Shaw (2012)

Microplitis lugubris

(Ruthe, 1860)

• Microgaster lugubris Ruthe, 1860

• borealis Marshall, 1885

• coracinus (Thomson, 1895, Microgaster)

• rutheana Fahringer, 1937

Distribution

Scotland

Microplitis malimbus

(Papp, 1984)

• Microgaster malimba Papp, 1984

• trochanterata misident

Distribution

England

Notes

Added by Shaw (2012). Nixon’s (Nixon 1970) interpretation of trochanterata (not tuberculifer, of which trochanterata is a junior synonym) is actually referable to malimbus (Shaw 2012).

Microplitis mandibularis

Thomson, 1895

Distribution

England, Scotland

Microplitis mediator

(Haliday, 1834)

• Microgaster mediator Haliday, 1834

• medianus (Ruthe, 1860, Microgaster)

• halidayi Fahringer, 1937

• pseudomedianus Fahringer, 1937

Distribution

England, Scotland, Wales, Ireland, Isle of Man

Microplitis moestus

(Ratzeburg, 1852)

• Microgaster moestus Ratzeburg, 1852

Microplitis naenia

Nixon, 1970

Distribution

England

Microplitis ocellatae

(Bouché, 1834)

• Microgaster ocellatae Bouché, 1834

• canaliculatus (Wesmael, 1837, Microgaster)

Distribution

England

Microplitis scrophulariae

Szépligeti, 1898

Distribution

England

Notes

added by Shaw (2012)

Microplitis sofron

Nixon, 1970

• Microplitis sofron ?stigmaticus (Ratzeburg, 1844, Microgaster)

Distribution

England, Scotland, Ireland

Notes

Papp (1984) suggests that stigmaticus may be a senior synonym.

Microplitis spectabilis

(Haliday, 1834)

• Microgaster spectabilis Haliday, 1834

• fossulatus (Bouché, 1834, Microgaster)

• Microplitis spectabilis ?parvulus (Ruthe, 1860, Microgaster)

• seuratii Marshall, 1898

• testaceipes (Cameron, 1906, Dapsilotoma)

Distribution

England, Wales, Ireland

Microplitis spinolae

(Nees, 1834)

• Microgaster spinolae Nees, 1834

• sapporoensis Ashmead, 1906

• radiorimatus Telenga, 1955

• Microplitis spinolae ?quadridentatus (Provancher, 1886, Microgaster)

Distribution

England

Microplitis strenuus

Reinhard, 1880

• gracilis (Ruthe, 1860, Microgaster) preocc.

Distribution

England

Microplitis tristis

(Nees, 1834)

• Microgaster tristis Nees, 1834

• dolens Marshall, 1885

Distribution

England, Scotland, Wales

Microplitis tuberculatus

(Bouché, 1834)

• Microgaster tuberculatus Bouché, 1834

• fumipennis (Ratzeburg, 1852, Microgaster)

Distribution

England, Scotland, Ireland

Microplitis tuberculifer

(Wesmael, 1837)

• Microgaster tuberculifer Wesmael, 1837

• calcaratus (Thomson, 1895, Microgaster)

• trochanteratus (Thomson, 1895, Microgaster)

• manevali Gautier & Bonnamour, 1939

Distribution

England, Scotland

Microplitis viduus

(Ruthe, 1860)

• Microgaster viduus Ruthe, 1860

Distribution

England

Microplitis xanthopus

(Ruthe, 1860)

• Microgaster xanthopus Ruthe, 1860

• tenuipes (Thomson, 1895, Microgaster)

Distribution

Scotland, Wales, Ireland

Microtypinae

Microtypinae

Szépligeti, 1908

Notes

Belokobylskij et al. (2003) include Microtypus, without comment, in the Orgilinae, where it had traditionally been placed prior to recent phylogenetic work. In the molecular phylogeny of Belshaw and Quicke (2002) the genus Microtypus is the sister group to Homolobinae, as also found by Sharanowski et al. (2011) (albeit with this clade the sister group to Orgilinae). This relationship was originally suggested by Achterberg (1984b).

Microtypus

Ratzeburg, 1848

• SIMILEARINUS Glowacki & Karpiński, 1967

Notes

Nomenclature follows Capek and Achterberg (1992).

Microtypus wesmaelii

Ratzeburg, 1848

• dioryctriae Rohwer, 1920

Distribution

England

Notes

added by Shaw (1992a)

Miracinae

Miracinae

Viereck, 1918

Mirax

Haliday, 1833

• CENTISTIDEA Rohwer, 1914

Notes

There are also several unrecognised species in Britain.

Mirax rufilabris

Haliday, 1833

• spartii Haliday, 1835

• dryochares Marshall, 1898

• nanivorae Fischer, 1957

Distribution

England, Scotland

Notes

some distribution data from Shaw and Askew (1976) and NMS

Opiinae

Opiinae

Blanchard, 1845

Notes

The generic and tribal classification of opiines has been largely chaotic (Wharton 1988b). In Fauna Europaea, van Achterberg has enacted many taxonomic changes resulting from his work on Western Palaearctic Opiinae (in prep.), which are followed here. Some changes to the generic classification have been published by Wharton (1988b), van Achterberg (Achterberg 2004a, Achterberg 2004b, and in Belokobylskij et al. 2003) and by Li et al. (2013). Wharton (1988b) argued against recognition of tribes within Opiinae, except perhaps Ademonini (for the genus Ademon), with all the other genera in Opiini; van Achterberg (in prep.) does not recognise Ademonini. Some distribution data from Fischer (1958), Fischer (1967), Fischer (1997).

Ademon

Haliday, 1833

• GIARDINAIA de Stefani-Perez, 1902

• ANALOSTANIA Viereck, 1916

Ademon decrescens

(Nees, 1811)

• Bracon decrescens Nees, 1811

• mutuator (Nees, 1811, Bracon)

Distribution

England, Scotland, Wales, Ireland

Apodesmia

Förster, 1863

• ALLOTYPUS Förster, 1863

• LEMNAPHILOPIUS Fischer, 1972

• AGNOPIUS Fischer, 1982

• CRYPTOGNATHOPIUS Fischer, 1984

Notes

generic synonymy follows Li et al. (2013)

Apodesmia aemula

(Haliday, 1836)

• Opius aemulus Haliday, 1836

• melba (Papp, 1978, Opius)

Distribution

England, Ireland

Apodesmia curvata

(Fischer, 1957)

• Opius curvatus Fischer, 1957

Distribution

England, Scotland

Notes

added by Godfray (1986)

Apodesmia irregularis

(Wesmael, 1835)

• Opius irregularis Wesmael, 1835

• bipustulata (Fischer, 1958, Opius)

Distribution

England, Ireland

Apodesmia ocellata

(Wesmael, 1835)

• Opius ocellatus Wesmael, 1835

• areolaris (Thomson, 1895, Opius)

• hungarica (Szépligeti, 1896, Opius)

• bruta (Papp, 1978, Opius)

Distribution

England, Scotland

Apodesmia posticatae

(Fischer, 1957)

• Opius posticatae Fischer, 1957

• seebensteinensis (Fischer, 1959, Opius)

• hilaris (Fischer, 1963, Opius)

• hostium (Fischer, 1964, Opius)

Distribution

England, Scotland

Apodesmia rufipes

(Wesmael, 1835)

• Opius rufipes Wesmael, 1835

• taeniata Förster, 1863

• taeniata (Fischer, 1957, Opius) preocc.

Distribution

England, Ireland

Apodesmia saeva

(Haliday, 1837)

• Opius saevus Haliday, 1837

Distribution

England, Scotland

Apodesmia saevula

(Fischer, 1958)

• Opius saevulus Fischer, 1958

Distribution

England

Apodesmia similis

(Szépligeti, 1898)

• Opius similis Szépligeti, 1898

• xylostei (Marshall, 1898, Opius)

• similiformis (Fischer, 1957, Opius)

• basirufa (Fischer, 1958, Opius)

• nodata (Fischer, 1958, Opius)

• periclymenii (Fischer, 1964, Opius)

• altimontana (Fischer, 1969, Opius)

• differens (Fischer, 1958, Opius)

• parvipunctum (Fischer, 1958, Opius)

• selkirkensis (Fischer, 2006, Opius); synonymy by van Achterberg (in prep.)

Distribution

England, Scotland, Ireland

Atormus

van Achterberg, 1998

Atormus victus

(Haliday, 1837)

• Opius victus Haliday, 1837

• tarni (Papp, 1982, Opius)

Distribution

England, Scotland, Ireland

Notes

Listed as a synonym of Opius singularis in Huddleston (1978).

Bathystomus

Förster, 1863

• COMPRESSARIA Königsmann, 1959

Notes

Although Wharton (1993) noted that the type (and only included) species of Compressaria is the same species as the type of Bathystomus, he did not formally synonymise these names and the synonymy has not been picked up on by, e.g. Belokobylskij et al. (2003) and Taxapad (Yu et al. 2012). Wharton (1988b) demonstrated that Bathystomus is a valid genus but Belokobylskij et al. (2003) treated it as a synonym of Diachasma. Van Achterberg (2014) formally synonymised Compressaria and pugnatrix. Huddleston (1978) included Compressaria pugnatrix as a species of Rogadinae.

Bathystomus xanthopus

Förster, 1863

• pugnatrix (Marshall, 1895, Mesocrina)

• compressiventris (Fischer, 1964, Opius); synonymy by van Achterberg (in prep.)

Distribution

England

Biophthora

Förster, 1863

Notes

Treated as a valid genus following Wharton (2006).

Biophthora bajula

(Haliday, 1837)

• Opius bajulus Haliday, 1837

• beieri (Fischer, 1968, Sternaulopius); synonymy by Wharton (2006)

Distribution

England, Ireland

Biosteres

Förster, 1863

• RHABDOSPILUS Förster, 1863

• RHINOPLUS Förster, 1863

• STENOSPILUS Förster, 1863

• ZETETES Förster, 1863 preocc.

• OPIELLUS Ashmead, 1900

• CELIESTIELLA Cameron, 1903

Biosteres analis

(Wesmael, 1835)

• Opius analis Wesmael, 1835

• colorativentris (Fischer, 1957, Opius)

Distribution

England, Scotland

Biosteres arenarius

(Stelfox, 1959)

• Opius arenarius Stelfox, 1959

Distribution

Ireland

Biosteres bicolor

Wesmael, 1835

• vagator (Förster, 1863, Stenospilus)

Distribution

Ireland

Biosteres carbonarius

(Nees, 1834)

• Bracon carbonarius Nees, 1834

• impressus (Wesmael, 1835, Opius)

• procerus (Wesmael, 1835, Opius)

• onzi (Fischer, 1959, Opius); synonymy by Achterberg (2014)

Distribution

England, Scotland, Wales, Ireland

Notes

Synonymised under carbonarius by Achterberg (1975), impressus was removed from synonymy by Fischer (1997), a move which is not accepted by van Achterberg (in prep.). Supporting van Achterberg’s synonymy, Godfray (unpublished) has repeatedly reared specimens resembling both ‘species’ from collections of hosts made at the same place and time.

Biosteres haemorrhoeus

(Haliday, 1837)

• Opius haemorrhoeus Haliday, 1837

• castaneiventris (Thomson, 1895, Opius)

• palaearcticus Szépligeti, 1901

Distribution

England, Ireland

Biosteres magnicornis

(Wesmael, 1835)

• Opius magnicornis Wesmael, 1835

Distribution

England, Ireland

Biosteres micans

(Stelfox, 1957)

• Opius micans Stelfox, 1957

• nitidus (Stelfox, 1949, Opius) preocc.

Distribution

Scotland, Ireland

Biosteres placidus

(Haliday, 1837)

• Opius placidus Haliday, 1837

• melanocerus (Wesmael, 1838, Opius)

• tarsator (Thomson, 1895, Opius)

• indotatus Viereck,1905

Distribution

England, Ireland

Biosteres rusticus

(Haliday, 1837)

• Opius rusticus Haliday, 1837

Distribution

England, Scotland, Ireland

Biosteres scabriculus

(Wesmael, 1835)

• Opius scabriculus Wesmael, 1835

Distribution

Ireland

Biosteres spinaciae

(Thomson, 1895)

• Opius spinaciae Thomson, 1895

• pegomyiae (Gahan, 1917, Opius)

• hyoscyamiellus (Viereck, 1925, Opius)

Distribution

England

Notes

added by Godfray (1988)

Biosteres sylvaticus

(Haliday, 1837)

• Opius sylvaticus Haliday, 1837

• clypealis (Thomson, 1895, Opius)

• nitidus Szépligeti, 1896

Distribution

England, Scotland, Ireland

Biosteres wesmaelii

(Haliday, 1837)

• Opius wesmaelii Haliday, 1837

• carbonarius (Wesmael, 1835, Opius) preocc.

• ultor (Förster, 1863, Zetetes)

• ultor (Fischer, 1957, Opius) preocc.

• jonaitisi (Jakimavi?ius, 1977, Opius)

Distribution

England, Scotland

Bitomoides

van Achterberg, 2004

Bitomoides rugosus

(Wesmael, 1838)

• Opius rugosus Wesmael, 1838

• rugiventris (Thomson, 1895, Opius); synonymy by Achterberg (2014)

Distribution

England

Notes

Listed as British by Kloet and Hincks (1945) but omitted by Huddleston (1978). Material in NMS has been identified by van Achterberg.

Chilotrichia

Förster, 1863

• TRICHOPIUS Thomson, 1895

Chilotrichia blanda

(Haliday, 1837)

• Opius blandus Haliday, 1837

Distribution

England, Ireland

Desmiostoma

Förster, 1863

Notes

Regarded by Wharton (Wharton 1983, Wharton 1988b) as a junior synonym of Opius.

Desmiostoma parvulum

(Wesmael, 1835)

• Opius parvulus Wesmael, 1835

• nudiscutum (Fischer, 1964, Opius)

• ziratus (Papp, 1982, Opius)

• temporale (Fischer, 1958, Opius)

Distribution

England, Scotland

Diachasma

Förster, 1863

• Atoreuteus Förster, 1863

• Lytacra Förster, 1863

• Alysopius Tobias, 1976

Diachasma caffer

(Wesmael, 1835)

• Opius caffer Wesmael, 1835

• stygium (Förster, 1863, Lytacra)

Diachasma cephalotes

(Wesmael, 1835)

• Opius cephalotes Wesmael, 1835

Distribution

England

Diachasma fulgidum

(Haliday, 1837)

• Opius fulgidus Haliday, 1837

Distribution

England, Ireland

Eurytenes

Förster, 1863

Eurytenes

Förster, 1863

Eurytenes (Eurytenes) abnormis

(Wesmael, 1835)

• Opius abnormis Wesmael, 1835

Distribution

England, Ireland

Notes

According to Walker and Wharton (2011), Opius paradoxus Ratzeburg, 1848, sometimes treated as a valid name (synonymous with abnormis) should be regarded as invalid as it was first proposed as a synonym of abnormis; Walker and Wharton (2011) also provide some locality data.

Eurytenes (Eurytenes) britannicola

Fischer, 2006

Distribution

England

Notes

Added by Fischer (2006): mis-placed in Eurytenes (van Achterberg, pers. comm.).

Stigmatopoea

Fischer, 1986

Notes

The generic name Stigmatopoea has been treated as a synonym of Xynobius by Achterberg (2004a) but the type species, macrocerus, has been regarded as a species of Eurytenes, a genus which was ignored by Achterberg (2004a). Both Xynobius and Stigmatopoea are treated as subgenera of Eurytenes, following Wharton (2006), although Li et al. (2013) again synonymised Stigmatopoea with Xynobius.

Eurytenes (Stigmatopoea) macrocerus

(Thomson, 1895)

• Opius macrocerus Thomson, 1895

• hians (Stelfox, 1949, Opius)

Distribution

England, Scotland, Ireland

Xynobius

Förster, 1863

• ACLISIS Förster, 1863

• HOLCONOTUS Förster, 1863

• AULONOTUS Ashmead, 1900

• ERISTERNAULAX Viereck, 1914

• XYNOBIOTENES Fischer, 1998

Notes

Generic synonymy from Achterberg (2004a) and Li et al. (2013) but treated as a subgenus of Eurytenes by Wharton (2006).

Eurytenes (Xynobius) aciculatus

(Thomson, 1895)

• Opius aciculatus Thomson, 1895

• tenuicornis (Thomson, 1895, Opius); synonymy by van Achterberg (in prep.)

Distribution

Ireland

Eurytenes (Xynobius) aemuloides

(Fischer, 1958)

• Opius aemuloides Fischer, 1958

Distribution

England

Eurytenes (Xynobius) caelatus

(Haliday, 1837)

• Opius caelatus Haliday, 1837

• isomera (Förster, 1863, Aclisis)

• pallipes Förster, 1863

• pallidipes Dalla Torre, 1898

Distribution

England, Scotland, Ireland

Eurytenes (Xynobius) comatus

(Wesmael, 1835)

• Opius comatus Wesmael, 1835

• sulcifer (Papp, 1967, Dapsilarthra)

Distribution

England, Ireland

Eurytenes (Xynobius) geniculatus

(Thomson, 1895)

• Opius geniculatus Thomson, 1895

• albicoxis (Marshall, 1898, Opius)

Distribution

England, Scotland

Eurytenes (Xynobius) holconotus

(Fischer, 1958)

• Opius holconotus Fischer, 1958

Distribution

England

Eurytenes (Xynobius) maculipes

Wesmael, 1835

• addendus Fischer, 1959

• turcmenicus Fischer, 1959

Distribution

England, Ireland

Eurytenes (Xynobius) polyzonius

(Wesmael, 1835)

• Opius polyzonius Wesmael, 1835

Distribution

England, Scotland

Eurytenes (Xynobius) silenis

(Fischer, 1967)

• Diachasma silenis Fischer, 1967

Distribution

Wales

Notes

Described from Welsh material (Fischer 1967) but omitted by Huddleston (1978).

Eurytenes (Xynobius) thomsoni

(Fischer, 1971)

• Opius thomsoni Fischer, 1971

• annulicornis (Thomson, 1895, Opius) preocc.

Distribution

England

Eurytenes (Xynobius) sp. A

van Achterberg, in prep.

Distribution

England

Notes

reared by Godfray, in NMS

Neopius

Gahan, 1917

Notes

Raised from synonymy with Phaedrotoma by Li et al. (2013).

Neopius rudis

(Wesmael, 1835)

• Opius rudis Wesmael, 1835

• carinaticeps Gahan, 1917

Distribution

England, Ireland

Opiognathus

Fischer, 1972

Notes

Raised from synonymy with Phaedrotoma by Li et al. (2013).

Opiognathus pactus

(Haliday, 1837)

• Opius pactus Haliday, 1837

Distribution

England, Ireland

Opiostomus

Fischer, 1972

• SNOFLAKOPIUS Fischer, 1972

• JUCUNDOPIUS Fischer, 1984

• OETZALOTENES Fischer, 1998

• OPIOTENES Fischer, 1998

Notes

Raised from synonymy with Opius by Li et al. (2013), who report the generic synonymy.

Opiostomus aureliae

(Fischer, 1957)

• Opius aureliae Fischer, 1957

Distribution

England, Scotland

Notes

added by Godfray and Achterberg (2015)

Opiostomus campanariae

(Fischer, 1959)

• Opius campanariae Fischer, 1959

Distribution

Scotland

Notes

added by Godfray and Achterberg (2015)

Opiostomus griffithsi

(Fischer, 1962)

• Opius griffithsi Fischer, 1962

Distribution

England

Opiostomus leptostigma

(Wesmael, 1835)

• Opius leptostigma Wesmael, 1835

• percontator (Fischer, 1964, Opius)

Distribution

England, Scotland

Notes

Recorded as a British species by various authors, latterly by Fischer (1967), and reared recently by Godfray (unpublished), but omitted by Huddleston (1978).

Opius

Wesmael, 1835

• CRYPTONASTES Förster, 1863

• HYPOCYNODUS Förster, 1863

• HYPOLABIS Förster, 1863

• MISOPHTHORA Förster, 1863

• DESMATOPHORUS Thomson, 1895

• ALLOPHLEBUS Fischer,1972

• NOSOPAEOPIUS Fischer, 1972

• OPIOTHORAX Fischer, 1972

• PENDOPIUS Fischer, 1972

• STOMOSEMA Fischer, 1972

• ODONTOPOEA Fischer, 1987

Notes

Restricted by Achterberg and Salvo (1997) to a group of species with distinctive mandibles.

Opius agromyzicola

Fischer, 1967

Distribution

England

Opius ambiguus

Wesmael, 1835

• celsus Haliday, 1837; synonymy by Achterberg (2014)

• longipes Fischer, 1957

• phytomyzae Fischer, 1957

Distribution

England, Scotland, Ireland

Opius brevipalpis

Thomson, 1895

• mutus Fischer, 1964

• gyoerfii Fischer, 1958; synonymy by Achterberg (2014)

Distribution

England

Opius caudifer

Fischer, 1958

• longicornis misident.

Distribution

England

Notes

Listed as a synonym of longicornis Thomson, 1895, in Taxapad (Yu et al. 2012), regarded as a separate, valid species by van Achterberg (in prep.). Recorded as British by Fischer (1967) as longicornis but omitted by Huddleston (1978).

Opius cingulatus

Wesmael, 1835

• dentifer Thomson, 1895

• stramineipes Thomson, 1895

Opius compar

Marshall, 1894

• pulchrithorax Fischer, 1958

Opius crassipes

Wesmael, 1835

Distribution

England

Opius flammeus

Fischer, 1959

Distribution

England

Opius funebris

Wesmael, 1835

• latipes misident.

Distribution

England, Ireland

Notes

Listed as a synonym of pygmaeator in Taxapad (Yu et al. 2012), regarded as a separate, valid species (=latipes sensu Fischer) by van Achterberg (in prep.).

Opius fuscipennis

Wesmael, 1835

Distribution

England

Notes

Fischer (1960) mistakenly listed this species as fuscipennis (Szépligeti, 1914, Rhinoplus) (van Achterberg, pers. comm.), a separate species now placed in the genus Pseudorhinoplus Fischer, 1972.

Opius gracilis

Fischer, 1957

• csikii Fischer, 1957

• minor Fischer, 1957

• nigrithorax Fischer, 1958 preocc.

Distribution

England

Opius instabilis

Wesmael, 1835

Opius levis

Wesmael, 1835

• apiculator (Nees, 1834, Bracon)

• filicornis Thomson, 1895

• varipes Szépligeti, 1898

Distribution

England, Wales, Ireland

Notes

Opius filicornis was recorded as new to Britain by Godfray (1986) but synonymised with levis by van Achterberg (in Belokobylskij et al. 2003).

Opius lucidus

Szépligeti, 1896

Distribution

England, Scotland

Opius lugens

Haliday, 1837

• abscissus Thomson, 1895

• obscurus Szépligeti, 1901

• adveniens Fischer, 1960

Distribution

England, Ireland

Opius nigricoloratus

Fischer, 1958

• dureseaui Fischer, 1975; synonymy by Achterberg (2014)

Distribution

Scotland, Wales

Notes

added by Godfray and Achterberg (2015)

Opius ochrogaster

Wesmael, 1835

• nigriceps Szépligeti, 1898

• neopusillus Fischer, 1957

Distribution

England, Scotland, Ireland

Opius orbiculator

(Nees, 1811)

• Bracon orbiculator Nees, 1811

• breviscapus Thomson, 1895

Distribution

Ireland

Opius pallipes

Wesmael, 1835

• exilis Haliday, 1837; synonymy by Achterberg (1997)

• pallidipes (Marshall, 1872, Hypolabis)

• liopleuris Thomson, 1895

• piceus Thomson, 1895

• adaequator (Fischer, 1964, Hypolabis) nom. nud.

• lividipes (Fischer, 1964, Hypolabis) nom. nud.

• subsulcatus (Fischer, 1964, Hypolabis) nom. nud.

• extusus Papp, 1981

• cisromensis Papp, 1982

Distribution

England, Scotland, Wales, Ireland

Opius pendulus

Haliday, 1837

• latipes Fischer, 1958

Distribution

England, Ireland

Opius phytobiae

Fischer, 1959

Distribution

England

Notes

added by Godfray (1986)

Opius propodealis

Fischer, 1958

Distribution

England

Opius pygmaeator

(Nees, 1811)

• Bracon pygmaeator Nees, 1811

• ruminans Fischer, 1957

• dilatatus Fischer, 1960

• meracus Fischer, 1960

Distribution

England, Ireland

Opius pygmaeus

Fischer, 1962

Distribution

England

Opius singularis

Wesmael, 1835

• clarus Haliday, 1836; synonymy by Fischer (1997)

• spretus Haliday, 1836; synonymy by Fischer (1997)

• vindex Haliday, 1837; synonymy by van Achterberg (in prep.)

• arenosus Szépligeti, 1898

Distribution

England, Ireland

Notes

Both clarus and spretus were also synonymised by Achterberg (1997) but Fischer’s publication pre-dated this.

Opius soenderupianus

Fischer, 1967

Distribution

England

Notes

added by Godfray and Achterberg (2015)

Opius tenellae

Fischer, 1969

Distribution

Scotland, Ireland

Phaedrotoma

Förster, 1863

• EUTRICHOPSIS Förster, 1863

• NOSOPOEA Förster, 1863

• TOLBIA Cameron, 1907

• BRACHYCENTRUS Szépligeti, 1907

• COELOREUTEUS Roman, 1910

• HEXAULAX Cameron, 1910

• BAEOCENTRUM Schulz, 1911

• NEODIOSPILUS Szépligeti, 1911

• NEOPIUS Fischer, 1965

• EUOPIUS Fischer, 1967

• GASTROSEMA Fischer, 1972

• GERIUS Fischer, 1972

• GRIMNIRUS Fischer, 1972

• HOENIRUS Fischer, 1972

• MEROTRACHYS Fischer, 1972

• MIMIRUS Fischer, 1972

• PHLEBOSEMA Fischer, 1972

• NEOEPHEDRUS Samanta, Tamili, Saha & Raychaudhuri, 1983

• ADONTOPIUS Fischer, 1984

• KAINOPAEOPIUS Fischer, 1987

• MILLENIOPIUS Fischer, 1996

• NEOTROPOPIUS Fischer, 1999

Notes

Phaedrotoma was raised from synonymy with Opius by Achterberg and Salvo (1997) to accommodate many species previously placed in Opius. Generic synonymy follows Li et al. (2013).

species of Phaedrotoma excluded from the British and Irish list:

[viennensis (Fischer, 1959, Opius)] Listed by Huddleston (1978) but we cannot trace any published records or specimens.

Phaedrotoma aethiops

(Haliday, 1837)

• Opius aethiops Haliday, 1837

Distribution

England, Ireland

Phaedrotoma caesa

(Haliday, 1837)

• Opius caesus Haliday, 1837

• punctiventris (Thomson, 1895, Opius)

• subtilis (Szépligeti, 1898, Opius)

• hydrelliae (Rimsky-Korsakov, 1925, Opius)

• hydrelliae (Muesebeck, 1933, Opius) preocc.

• aquatica (Muesebeck, 1967, Opius)

• hydrelliana (Fischer, 1971, Opius)

Distribution

England, Ireland

Phaedrotoma curvata

(Fischer, 1957)

• Opius curvatus Fischer, 1957

Distribution

Scotland

Notes

added by Godfray (1986)

Phaedrotoma decorata

(Stelfox, 1949)

• Opius decoratus Stelfox, 1949

Distribution

Ireland

Phaedrotoma depeculator

(Förster, 1863)

• Opius depeculator Förster, 1863

• semiaciculata (Stelfox, 1949, Opius)

Distribution

England, Ireland

Phaedrotoma diversa

(Szépligeti, 1898)

• Opius diversus Szépligeti, 1898

Distribution

England

Notes

Recorded as British by Fischer (1960) but omitted by Huddleston (1978).

Phaedrotoma exigua

(Wesmael, 1835)

• Opius exiguus Wesmael, 1835

Distribution

England, Ireland

Phaedrotoma fallax

(Szépligeti, 1896)

• Opius fallax Szépligeti, 1896

Distribution

England

Notes

Listed as a synonym of Opius instabilis in Taxapad (Yu et al. 2012) but regarded as a valid species, in Phaedrotoma, by van Achterberg (in prep.).

Phaedrotoma fasciata

(Thomson, 1895)

• Opius fasciatus Thomson, 1895

• comparanda (Fischer, 1958, Opius)

Distribution

England

Phaedrotoma heringi

(Fischer, 1962)

• Opius heringi Fischer, 1962

Distribution

Ireland

Phaedrotoma instabiloides

(Fischer, 1959)

• Opius instabiloides Fischer, 1959

Distribution

Ireland

Phaedrotoma minusculae

(Fischer, 1967)

• Opius minusculae Fischer, 1967

Distribution

England

Phaedrotoma monticola

(Szépligeti, 1898)

• Opius monticola Szépligeti, 1898

Distribution

England

Notes

added by Fischer (1997)

Phaedrotoma munda

(Förster, 1863)

• Eutrichopsis munda Förster, 1863

• munda (Fischer, 1957, Opius) preocc.

Distribution

England

Phaedrotoma nitidulator

(Nees, 1834)

• Opius nitidulator Nees, 1834

• vittata (Ruschka, 1915, Opius) preocc.

Phaedrotoma novojariae

(Fischer, 2006)

• Opius novojariae Fischer, 2006

• stigmatocauda (Fischer, 2006, Eurytenes); synonymy by van Achterberg (in prep.)

Distribution

England

Notes

added by Fischer (2006)

Phaedrotoma paraphytomyzae

(Fischer, 2005)

• Opius paraphytomyzae Fischer, 2005

Distribution

England

Notes

added by Fischer (2005)

Phaedrotoma pulchriceps

(Szépligeti, 1898)

• Opius pulchriceps Szépligeti, 1898

• ilicis (Nixon, 1939, Opius)

• pulcherrimus (Fischer, 1958, Opius)

• pulchriventris (Fischer, 1958, Opius); synonymy by Achterberg (2014)

• vexator (Fischer, 1964, Opius)

• affectus (Papp, 1981, Opius)

Distribution

England, Ireland

Phaedrotoma recondes

van Achterberg, 2004

• reconditor misident.

Notes

Added by van Achterberg in Fauna Europaea; described by van Achterberg because the material named Opius reconditor (now classified in Rhogadopsis) by authors (e.g. Fischer 1972, Fischer and Koponen 1999) is not conspecific with the type. Both species apparently occur in Britain.

Phaedrotoma reptantis

(Fischer, 1957)

• Opius reptantis Fischer, 1957

Distribution

England, Scotland

Notes

Listed as a synonym of Opius ambiguus in Taxapad (Yu et al. 2012) but regarded as a valid species, in Phaedrotoma, by van Achterberg (in prep.).

Phaedrotoma rex

(Fischer, 1958)

• Opius rex Fischer, 1958

Distribution

England

Phaedrotoma rudiformis

(Fischer, 1958)

• Opius rudiformis Fischer, 1958

• uligiloci (Fischer, 2006, Opius); synonymy by Achterberg (2014)

Distribution

England

Phaedrotoma staryi

Fischer, 1958

• sitagrus Papp, 1982

Distribution

England, Scotland

Phaedrotoma tacita

(Haliday, 1837)

• Opius tacitus Haliday, 1837

Distribution

England

Notes

omitted by Huddleston (1978)

Phaedrotoma variegata

(Szepligeti, 1896)

• Opius variegatus Szepligeti, 1896

Distribution

England, Scotland, Ireland

Rhogadopsis

Brèthes, 1913

• LISSOSEMA Fischer, 1972

Notes

Raised from synonymy with Phaedrotoma by Li et al. (2013).

Rhogadopsis reconditor

(Wesmael, 1835)

• Opius reconditor Wesmael, 1835

• docilis (Haliday, 1837, Opius)

• parvungula Thomson, 1895

Distribution

England, Ireland

Notes

Opius docilis was synonymised with reconditor by Achterberg (1997) but listed as a separate species in Taxapad (Yu et al. 2012); re-synonymised by Achterberg (2014).

Utetes

Förster, 1863

• THEROBOLUS Förster, 1863

• FREKIUS Fischer, 1971; synonymy by Wharton (2006)

Notes

species of Utetes excluded from the British and Irish list:

[christenseni (Papp, 1982, Opius)] Listed in Fauna Europaea as occurring in Britain, but presumably mistakenly as no literature or specimen records can be located. Known only from Georgia, Greece and Ukraine (data from Yu et al. 2012).

Utetes caudatus

(Wesmael, 1835)

• Opius caudatus Wesmael, 1835

• exsertus (Thomson, 1895, Opius)

Distribution

England

Notes

Listed as English by Fischer and Koponen (1999), presumably following Fischer (1958), but not listed in Huddleston (1978).

Utetes coracinus

(Thomson, 1895)

• Opius coracinus Thomson, 1895

Distribution

England

Notes

BMNH, det. Fischer, added in Fauna Europaea; there may be a literature citation for its occurrence in Britain (van Achterberg, pers. comm.) but we have been unable to trace it.

Utetes fulvicollis

(Thomson, 1895)

• Opius fulvicollis Thomson, 1895

• cupidus (Gahan, 1919, Opius)

Distribution

England

Notes

added by Godfray (1986)

Utetes rotundiventris

(Thomson, 1895)

• Opius rotundiventris Thomson, 1895

Distribution

England

Utetes ruficeps

(Wesmael, 1835)

• Opius ruficeps Wesmael, 1835

Distribution

England

Utetes testaceus

(Wesmael, 1838)

• Opius testaceus Wesmael, 1838

Distribution

England

Utetes truncatus

(Wesmael, 1838)

• Opius truncatus Wesmael, 1838

Distribution

England

Utetes zelotes

(Marshall, 1891)

• Opius zelotes Marshall, 1891

• insertus (Fischer, 1971, Opius)

• incertus misspelling

Distribution

England, Scotland

Orgilinae

Orgilinae

Ashmead, 1900

Orgilus

Haliday, 1833

• ISCHIUS Wesmael, 1837

• MACROPALPUS Ratzeburg, 1844

• ORESIMUS Ashmead, 1900

• ORGILOMORPHA Ashmead, 1900

• ISCHIOLUS Hellén, 1958

Notes

Taxonomic and much distribution data from Taeger (1989).

Orgilus achterbergi

Taeger, 1989

Distribution

England

Notes

added by Taeger (1989)

Orgilus dovnari

Tobias, 1986

• ukrainicus Tobias, 1986

Distribution

England

Notes

NMS, det. Taeger, added here

Orgilus interjectus

Taeger, 1989

Distribution

England

Notes

added by Taeger (1989)

Orgilus ischnus

Marshall, 1898

• subtilirugosus Papp, 1971

Distribution

England, Wales

Orgilus leptocephalus

Hartig, 1838

• obscurator misident.

• rugulosus Fahringer, 1937

• hyperboreus Hellén, 1958

Distribution

Ireland

Notes

Although Taeger (1989) clarified the identity of obscurator auctt. as leptocephalus, this species was recorded as obscurator by O'Connor et al. (1999).

Orgilus minor

Taeger, 1989

Distribution

England

Notes

NMS, det. Taeger, added here

Orgilus parvipennis

Thomson, 1895

• micropterus Morley, 1907

• macroptera (Rudow, 1917, Aptesis)

• decoratus Hellén, 1946

• curtipennis Fischer, 1958

• discolor Hellén, 1958

Distribution

England, Scotland

Orgilus pimpinellae

Niezabitowski, 1910

• laevigator misident.

Distribution

England, Scotland, Ireland

Notes

Added by Taeger (1989); apparently recorded as Irish under the name Microdus laevigator Nees, 1812 (O'Connor et al. 1999), which is listed as a species inquirendae by Taeger (1989).

Orgilus punctulator

(Nees, 1811)

• Microdus punctulator Nees, 1811

• rufiventris Fahringer, 1937

Distribution

England

Notes

added by Taeger (1989)

Orgilus rugosus

(Nees, 1834)

• Microgaster rugosus Nees, 1834

Distribution

England

Notes

NMS, det. Taeger, added here

Orgilus tobiasi

Taeger, 1989

Distribution

Wales, Ireland

Notes

added by Taeger (1989)

Pambolinae

Pambolinae

Marshall, 1885

Chremylini

Hellén, 1957

Chremylus

Haliday, 1833

• PENECERUS Wesmael, 1838

• PARAMESOCRINA Nagamori, 1925

Chremylus elaphus

Haliday, 1833

• rubiginosus (Nees, 1834, Hormius)

• transversus (Say, 1836, Bracon)

• nigriceps Ashmead, 1893

• terminalis Ashmead, 1893

• japonicus Ashmead, 1896

• tineavorus (Nagamori, 1925, Paramesocrina)

Distribution

England, Ireland

Pambolini

Marshall, 1885

Dimeris

Ruthe, 1854

• PARAPTESIS Magretti, 1884

Dimeris mira

Ruthe, 1854

• melanocephala (Marshall, 1870, Pambolus)

• flavipes (Magretti, 1884, Paraptesis)

• aptera Marshall, 1885

• inermis Fitch, 1885

Distribution

England

Pambolus

Haliday, 1836

Phaenodus

Förster, 1863

Pambolus (Phaenodus) pallipes

(Förster, 1863)

• Phaenodus pallipes Förster, 1863

• flavipes (Förster, 1863, Araphis)

• pallidipes (Marshall, 1897, Phaenodus)

• chalveri (Docavo, 1960, Phaenodus)

Distribution

England, Scotland

Notes

added by Belokobylskij (1986)

Pambolus

Haliday, 1836

• ARAPHIS Ruthe, 1854

• ARRHAPHIS misspelling

• ARHAPHIS misspelling

• FOLCHINIA Kieffer, 1906

• PARAMBOLUS Dahl, 1912

Notes

species of Pambolus (Pambolus) excluded from the British and Irish list:

[biglumis (Haliday, 1836, Rogas); syn. rosenhaueri (Ratzeburg, 1852, Pezomachus); dubius (Fitch, 1885, Araphis); imminens (Fitch, 1885, Araphis)] Included as British in Fauna Europaea and Taxapad (Yu et al. 2012) (but not by Huddleston 1978); this seems to be on the basis of Fitch’s descriptions of the synonymous names dubius and imminens in Marshall’s (Marshall 1885) monograph of British Braconidae. However, the types of Fitch’s species are from the Ruthe collection of German material and we have seen no British or Irish specimens of biglumis.

Rhysipolinae

Rhysipolinae

Belokobylskij, 1984

Pseudavga

Tobias, 1964

Pseudavga flavicoxa

Tobias, 1964

• rustus (Papp, 1991, Rhysipolis)

Distribution

England

Notes

added by Shaw and Sims (2015)

Rhysipolis

Förster, 1863

Notes

Most synonymy has been omitted; published taxonomy has been confused and there are several more species present than have been recorded in the literature, with the application of names not yet settled.

Rhysipolis decorator

(Haliday, 1836)

• Rogas decorator Haliday, 1836

• ruficeps (Wesmael, 1838, Exothecus)

• ruficornis (Szépligeti, 1896, Xenarcha)

Distribution

England, Scotland, Isle of Man

Notes

some distribution data from Shaw and Askew (1976)

Rhysipolis hariolator

(Haliday, 1836)

• Rogas hariolator Haliday, 1836

• barbatus (Wesmael, 1838, Exothecus)

Distribution

England, Scotland

Notes

some distribution data from Shaw and Askew (1976)

Rhysipolis meditator

(Haliday, 1836)

• Rogas meditator Haliday, 1836

Distribution

England, Scotland

Rhysipolis variabilis

(Szépligeti, 1896)

• Xenarcha variabilis Szépligeti, 1896

Distribution

England

Notes

NMS, det. Shaw & van Achterberg, added here; treated as a synonym of meditator by Belokobylskij and Tobias (1986) and Belokobylskij et al. (2003) but here treated as a valid species.

Rhysipolis varicoxa

(Thomson, 1892)

• Exothecus varicoxa Thomson, 1892

Distribution

England

Notes

NMS, det. Shaw & van Achterberg, added here; treated as a synonym of meditator by Belokobylskij and Tobias (1986) and Belokobylskij et al. (2003)​ but here treated as a valid species.

Rhyssalinae

Rhyssalinae

Förster, 1863

• HISTEROMERINAE Fahringer, 1930

Acrisidini

Hellén, 1957

Acrisis

Förster, 1863

• EUCHASMUS Marshall, 1888

• EPISIGALPHUS Ashmead, 1900

Acrisis exiguus

(Marshall, 1888)

• Euchasmus exiguus Marshall, 1888

Distribution

England

Proacrisis

Tobias, 1983

Notes

Treated as a synonym of Acrisis in Fauna Europaea, as a separate genus by Belokobylskij et al. (2003).

Proacrisis acutus

Tobias, 1983

Distribution

England

Notes

NMS, det. Shaw & van Achterberg, added here

Proacrisis rarus

Tobias, 1983

Distribution

England

Notes

NMS, det. Shaw & van Achterberg, added on Fauna Europaea

Histeromerini

Fahringer, 1930

Notes

Zaldívar-Riverón et al. (2006) and Sharanowski et al. (2011) both found, on the basis of molecular phylogenetic results, that the morphologically and biologically aberrant genus Histeromerus belongs in Rhyssalinae; however, there is as yet no indication as to how Histeromerus can be accommodated within the existing tribal classification of Rhyssalinae, so we simply use Histeromerini for now.

Histeromerus

Wesmael, 1838

• MITHOTYNIA Hedqvist, 1976

Histeromerus mystacinus

Wesmael, 1838

• apterus (Hedqvist, 1976, Mithotynia)

Distribution

England, Wales, Ireland

Notes

distribution data from Shaw (1995) and the Horniman Museum

Rhyssalini

Förster, 1863

Dolopsidea

Hincks, 1944

• DOLOPS Marshall, 1889 preocc.

• EXONTSIRA Belokobylskij, 1982

Dolopsidea indagator

(Haliday, 1836)

• Rogas indagator Haliday, 1836

• tuberculata (Wesmael, 1838, Exothecus)

• aculeator (Marshall, 1889, Dolops)

• hastifer (Marshall, 1889, Dolops)

• caucasica (Tobias, 1976, Doryctodes)

• rhodopea (Zaykov, 1980, Rhyssalus)

Distribution

England, Scotland, Ireland

Notes

some distribution data from Shaw (1993)

Oncophanes

Förster, 1863

• EPIRHYSSALUS Ashmead, 1900

Oncophanes minutus

(Wesmael, 1838)

• Exothecus minutus Wesmael, 1838

• lanceolator (Nees, 1834, Bracon) preocc.

• laevigatus (Ratzeburg, 1852, Bracon)

Distribution

England, Wales, Scotland

Notes

Traditionally treated as a species separate from minutus (e.g. Huddleston 1978; Fauna Europaea), laevigatus was synonymised by Belokobylskij (1998); we follow this, although it seems likely that there are two species involved.

Pseudobathystomus

Belokobylskij, 1986

Pseudobathystomus funestus

(Haliday, 1836)

• Rogas funestus Haliday, 1836

• schmiedeknechti (Fahringer, 1930, Bathystomus)

Distribution

England, Scotland

Notes

Listed as a species of Rhysipolis in Huddleston (1978).

Pseudobathystomus tobiasi

(Zaykov, 1980)

• Oncophanes tobiasi Zaykov, 1980

Distribution

England, Scotland

Notes

NMS, det. Shaw & van Achterberg, added on Fauna Europaea

Pseudobathystomus vernalis

Belokobylskij, 1994

Distribution

England, Scotland

Notes

NMS, det. Shaw & van Achterberg, added on Fauna Europaea

Rhyssalus

Haliday, 1833

• EURHOPTROCENTRUS Tobias,1977

Rhyssalus clavator

Haliday, 1833

Distribution

England, Scotland, Ireland, Isle of Man

Rhyssalus longicaudis

(Tobias & Belokobylskij, 1991)

• Eurhoptrocentrus longicaudis Tobias & Belokobylskij, 1991

Distribution

Wales

Notes

NMS, det. Shaw & van Achterberg, added on Fauna Europaea

Rogadinae

Rogadinae

Förster, 1863

Notes

Distribution data mostly from NMS and BMNH.

Aleiodini

Muesebeck, 1928

Notes

Resurrected by Zaldívar-Riverón et al. (2008b).

Aleiodes

Wesmael, 1838

• ROGAS misident.

• RHOGAS misident.

• PETALODES Wesmael, 1838

• NEORHOGAS Szépligeti, 1906

• CHELONORHOGAS Enderlein, 1912

Notes

Aleiodes species are further subdivided in Taxapad (Yu et al. 2012) into the subgenera Aleiodes, Chelonorhogas, Neorhogas and Heterogamus (for those occurring in Britain); the latter is considered here to be a distinct genus (see note under Heterogamus).

species of Aleiodes excluded from the British and Irish list:

[arcticus (Thomson, 1892, Rogas)] MRS has seen the specimens that this record is based upon (in Ipswich Museum) and they are not arcticus.

Aleiodes albitibia

(Herrich-Schäffer, 1838)

• Rogas albitibia Herrich-Schäffer, 1838

• heterogaster Wesmael, 1838

Distribution

England, Scotland, Wales, Ireland

Aleiodes alternator

(Nees, 1834)

• Rogas alternator Nees, 1834

• geniculator misident.

• balteatus (Curtis, 1834, Rogas)

Distribution

England, Scotland, Wales, Ireland

Aleiodes apicalis

(Brullé, 1832)

• Bracon apicalis Brullé, 1832

• ductor misident., in part

• reticulator (Nees, 1834, Rogas)

Distribution

England

Notes

to be added by van Achterberg & Shaw (in prep.); just one British specimen of this Mediterranean species, considered to be an erratic. Synonymy anticipates van Achterberg & Shaw (in prep.).

Aleiodes apiculatus

(Fahringer, 1932)

• Rhogas apiculatus Fahringer, 1932

Distribution

England

Notes

to be added by van Achterberg & Shaw (in prep.)

Aleiodes assimilis

(Nees, 1812)

• Bracon assimilis Nees, 1812

• bicolor misident., in part

• zygaenae (Nees, 1834, Rogas)

Distribution

England, Scotland, Ireland

Notes

added by Schwarz and Shaw (2000)

Aleiodes aterrimus

(Ratzeburg, 1852)

• Bracon aterrimus Ratzeburg, 1852

• grandis Giraud, 1857

Distribution

England

Aleiodes bicolor

(Spinola, 1808)

• Bracon bicolor Spinola, 1808

Distribution

England, Scotland, Wales, Ireland

Aleiodes cantherius

(Lyle, 1919)

• Rhogas cantherius Lyle, 1919

Distribution

England

Aleiodes circumscriptus

(Nees, 1834)

• Rogas circumscriptus Nees, 1834

Distribution

England, Scotland, Wales, Ireland

Notes

Sensu neotype (to be published by van Achterberg & Shaw, in prep.).

Aleiodes compressor

(Herrich-Schäffer, 1838)

• Rogas compressor Herrich-Schäffer, 1838

• unicolor (Wesmael, 1838, Petalodes) preocc.

Distribution

England, Scotland, Wales

Aleiodes coxalis

(Spinola, 1808)

• Rogas coxalis Spinola, 1808

• ater (Curtis, 1834, Rogas)

• tristis Wesmael, 1838

Distribution

England, Scotland, Isle of Man

Notes

Anticipates synonymy to be published by van Achterberg & Shaw (in prep.).

Aleiodes crassipes

(Thomson, 1892)

• Rogas crassipes Thomson, 1892

Distribution

Wales

Notes

to be added by van Achterberg & Shaw (in prep.)

Aleiodes cruentus

(Nees, 1834)

• Rogas cruentus Nees, 1834

Distribution

England

Aleiodes dissector

(Nees, 1834)

• Rogas dissector Nees, 1834

• aestivalis (Vollenhoven, 1858, Phylax)

Distribution

England, Scotland

Aleiodes fortipes

(Reinhard, 1863)

• Rogas fortipes Reinhard, 1863

• freyi (Hellén, 1927, Rhogas)

Distribution

England

Notes

to be added by van Achterberg & Shaw (in prep.)

Aleiodes gastritor

(Thunberg, 1824)

• Ichneumon gastritor Thunberg, 1824

• circumscriptus misident., in part

• testaceus misident., in part

Notes

Distribution data are not given as this name covers an aggregate of at least four species in Britain (van Achterberg & Shaw, in prep.). Further, it is not clear to which the name gastritor should be applied. Bracon testaceus Spinola, 1808 is actually Rogas luteus, and ‘Aleiodes testaceus (Spinola)’ is not a valid taxon, although it has been applied to several small, orangeish Aleiodes, epecially in the gastritor aggregate.

Aleiodes grassator

(Thunberg, 1824)

• Ichneumon grassator Thunberg, 1824

• carbonarius Giraud, 1857

• flavipalpis (Thomson, 1892, Rogas)

Distribution

England, Scotland

Notes

Anticipates synonymy to be published by van Achterberg & Shaw (in prep.).

Aleiodes heterostigma

(Stelfox, 1953)

• Rogas heterostigma Stelfox, 1953

Distribution

Wales, Ireland

Aleiodes hirtus

(Thomson, 1892)

• Rogas hirtus Thomson, 1892

Distribution

England, Scotland

Notes

To be added by van Achterberg & Shaw (in prep.); treated as a synonym of pallidicornis by Belokobylskij et al. (2003), following Papp (1985).

Aleiodes modestus

(Reinhard, 1863)

• Rogas modestus Reinhard, 1863

• piceus (Fahringer, 1932, Rhogas)

Distribution

England, Scotland, Wales

Aleiodes nigriceps

Wesmael, 1838

Distribution

England, Scotland, Wales, Ireland

Notes

to be added by van Achterberg & Shaw (in prep.); listed as a subspecies of circumscriptus in Taxapad (Yu et al., 2012), following Papp (1999c) and other authors.

Aleiodes nigricornis

Wesmael, 1838

Distribution

England, Scotland, Wales, Ireland

Aleiodes nobilis

(Haliday, 1834)

• Rogas nobilis Haliday, 1834

• ductor misident., in part

• medianus (Thomson, 1892, Rogas)

Distribution

England, Scotland, Ireland

Notes

Brought out of synonymy with ductor by Achterberg (1997).

Aleiodes pallidator

(Thunberg, 1824)

• Ichneumon pallidator Thunberg, 1824

• ochraceus (Curtis, 1834, Rogas)

• unicolor Wesmael, 1838

• Aleiodes pallidator ?pellucens (Telenga, 1941, Rhogas)

Distribution

England

Notes

some distribution data from Shaw (1977), Shaw (1981)

Aleiodes pallidicornis

(Herrich-Schäffer, 1838)

• Rogas pallidicornis Herrich-Schäffer, 1838

• ductor misident., in part

Distribution

Scotland

Notes

to be added by van Achterberg & Shaw (in prep.); specimen in BMNH (seen by MRS).

Aleiodes pictus

(Herrich-Schäffer, 1838)

• Rogas pictus Herrich-Schäffer, 1838

• borealis misident., in part

• circumscriptus misident., in part

• nigriceps misident., in part

Distribution

England, Scotland, Wales, Ireland

Notes

to be added by van Achterberg & Shaw (in prep.)

Aleiodes praetor

(Reinhard, 1863)

• Rogas praetor Reinhard, 1863

• luteus (Szépligeti, 1906, Neorhogas)

Distribution

England

Aleiodes pulchripes

Wesmael, 1838

Distribution

England, Ireland, Isle of Man

Notes

some distribution data from Shaw (1979)

Aleiodes punctipes

(Thomson, 1892)

• Rogas punctipes Thomson, 1892

Distribution

Scotland, Wales, Ireland

Aleiodes ruficornis

(Herrich-Schäffer, 1838)

• Rogas ruficornis Herrich-Schäffer, 1838

• dimidiatus misident.

• gasterator misident.

Distribution

England, Ireland

Aleiodes rugulosus

(Nees, 1811)

• Bracon rugulosus Nees, 1811

• pictus (Kokujev, 1898, Rhogas) preocc.

Distribution

England, Scotland, Wales, Ireland

Aleiodes seriatus

(Herrich-Schäffer, 1838)

• Rogas seriatus Herrich-Schäffer, 1838

• vittiger Wesmael, 1838

• kuslitzkyi (Tobias, 1976, Rogas)

Distribution

England, Ireland

Aleiodes signatus

(Nees, 1812)

• Bracon signatus Nees, 1812

• geniculator (Nees, 1834, Rogas)

• annulipes (Herrich-Schäffer, 1838, Rogas)

• esseni Hellén, 1927

Distribution

England, Wales

Aleiodes similis

(Curtis, 1834)

• Rogas similis Curtis, 1834

• circumscriptus misident., in part

• testaceus misident., in part

• spathuliformis (Curtis, 1834, Rogas)

• subucola (Curtis, 1834, Rogas)

• armatus Wesmael, 1838

Distribution

England, Scotland, Ireland

Notes

The name armatus has been misapplied to a range of undescribed, medium-sized, orange species, at least three of which occur in Britain and Ireland and will be described by van Achterberg & Shaw (in prep.).

Aleiodes testaceus

(Telenga, 1941)

• Heterogamus testaceus Telenga, 1941

Distribution

England, Wales

Notes

to be added by van Achterberg & Shaw (in prep.)

Aleiodes ungularis

(Thomson, 1892)

• Rogas ungularis Thomson, 1892

Distribution

England, Wales, Ireland

Notes

To be added by van Achterberg & Shaw (in prep.); English and Welsh records from specimens seen by MRS in, respectively, the Hope Department, Oxford, and Doncaster Museum.

Aleiodes unipunctator

(Thunberg, 1824)

• Ichneumon unipunctator Thunberg, 1824

• ductor (Thunberg, 1824, Ichneumon)

• irregularis Wesmael, 1838

Distribution

England, Scotland, Wales, Ireland

Notes

Anticipates synonymy to be published by van Achterberg & Shaw (in prep.).

Aleiodes sp. L

van Achterberg & Shaw, in prep.

• borealis misident.

• circumscriptus misident., in part

• nigriceps misident., in part

Distribution

England, Scotland, Wales, Ireland, Isle of Man

Notes

To be added by van Achterberg & Shaw (in prep.); this species, whose name is not yet published (van Achterberg & Shaw, in prep.), is very common in the British Isles and M.R. Shaw det. labels (as borealis) have been left in several British collections (up to 2007). The true A. borealis (Thomson, 1892, Rogas) has not been found in the British Isles.

Heterogamus

Wesmael, 1838

Notes

Usually treated as a subgenus of Aleiodes (as in Taxapad: Yu et al. 2012) but Zaldivar-Riverón et al. (Zaldívar-Riverón et al. 2004, Zaldívar-Riverón et al. 2008b) have shown that, on the basis of venom apparatus characters and molecular sequence data, respectively, Heterogamus species form a clade distinct from Aleiodes species. Murray (1939) and Shaw (2000a) published some distribution records.

Heterogamus dispar

(Haliday, 1833)

• Rogas dispar Haliday, 1833

• dispar (Curtis, 1834, Rogas) preocc.

• crypticornis (Wesmael, 1838, Aleiodes)

Distribution

England, Scotland, Wales, Ireland

Heterogamus excavatus

Telenga, 1941

• farmakena Maláč, 1941

Distribution

England

Notes

Added by Shaw (2000a); Irish records (O'Connor et al. 1999) are erroneous (Shaw 2000a).

Clinocentrini

van Achterberg, 1991

Clinocentrus

Haliday, 1833

• CAMPTOCENTRUS Kriechbaumer, 1894

• MICRORHOGAS Cameron, 1910

• NEORHYSSALUS Baker, 1917

Notes

Nomenclature and distribution data from Belokobylskij (1995) and NMS.

species of Clinocentrus excluded from the British and Irish list:

[tenuicornis (Thomson, 1891, Exothecus)] A species of uncertain status (Belokobylskij, pers. comm. to MRS), listed as a synonym of excubitor in Taxapad (Yu et al. 2012), following Szépligeti (1906).

Clinocentrus brevicalcar

(Thomson, 1891)

• Exothecus brevicalcar Thomson, 1891

Distribution

England, Scotland

Clinocentrus cunctator

(Haliday, 1836)

• Rogas cunctator Haliday, 1836

• analis (Wesmael, 1838, Exothecus)

• gracilipes (Thomson, 1892, Exothecus)

Distribution

England, Scotland, Wales, Ireland

Clinocentrus excubitor

(Haliday, 1836)

• Rogas excubitor Haliday, 1836

• marginellus (Wesmael, 1838, Exothecus)

Distribution

England, Scotland, Wales, Ireland

Clinocentrus exsertor

(Nees, 1811)

• Bracon exsertor Nees, 1811

• orbitator (Nees, 1834, Bracon)

• striolatus (Thomson, 1891, Exothecus)

• tarsalis Ashmead, 1894

Distribution

England, Scotland

Clinocentrus hungaricus

Szépligeti, 1906

Distribution

England, Scotland

Notes

NMS, BMNH, det. Shaw, added here

Clinocentrus umbratilis

Haliday, 1833

• petiolaris (Thomson, 1891, Exothecus)

• polonicus Fahringer, 1931

Distribution

England, Scotland, Ireland

Clinocentrus vestigator

(Haliday, 1836)

• Rogas vestigator Haliday, 1836

• stigmaticus Marshall, 1897

• jaroshevskyi Telenga, 1941

• obsoletus (Hellén, 1957, Oncophanes)

Distribution

England, Scotland, Ireland, Isle of Man

Rogadini

Förster, 1863

• PELECYSTOMINI Viereck, 1918

Rogas

Nees, 1818

• PELECYSTOMA Wesmael, 1838

• RHOGAS Agassiz, 1846

Rogas luteus

Nees, 1834

• testaceus (Fabricius, 1798, Ichneumon) preocc.

• testaceator (Thunberg, 1824, Ichneumon)

Triraphis

Ruthe, 1855

Notes

species of Triraphis excluded from the British and Irish list:

[tricolor (Wesmael, 1838, Pelecystoma); syn. solitarius (Watanabe, 1970, Pelecystoma); synonymy by Papp (1995a)] Listed by Huddleston (1978), probably on the basis of the host-parasitoid catalogue of Morley and Rait-Smith (1933), but this publication included non-British rearings from Lepidoptera that occur in Britain and there is no evidence that tricolor, a parasitoid of Apoda limacodes (Hufnagel) (Lepidoptera: Limacodidae), has ever been found in Britain or Ireland.

Sigalphinae

Sigalphinae

Haliday, 1833

Acampsini

van Achterberg & Austin, 1992

Acampsis

Wesmael, 1835

Acampsis alternipes

(Nees, 1816)

• Sigalphus alternipes Nees, 1816

Distribution

England

Sigalphini

Haliday, 1833

Sigalphus

Latreille, 1802

• SPHAEROPYX Illiger, 1807

• RHITIGASTER Wesmael, 1835

• RHYTIDOGASTER Agassiz, 1846

Notes

species of Sigalphus excluded from the British and Irish list:

[irrorator (Fabricius, 1775, Ichneumon); syn. niger (Retzius, 1783, Ichneumon); syn. globulifer (Geoffroy, 1785, Ichneumon); irroratrix (Schulz, 1906, Sphaeropyx)] Shaw and Huddleston (1991) were unable to trace any British specimens.

Supplementary Material

Supplementary material 1

Checklist of British and Irish Hymenoptera

Broad, G.R., Shaw, M.R. & Godfray, H.C.J.

Data type: formatted text file

Brief description: Word document version of the checklist

File: oo_84392.docx

Supplementary material 2

Checklist of British and Irish Braconidae

Broad, G.R., Shaw, MR. & Godfray, H.C.J.

Data type: spreadsheet

Brief description: Excel spreadsheet version of the checklist

File: oo_84393.xlsx

Author contributions

The sections on subfamilies Microgastrinae and Rogadinae are mainly the work of MRS, the section on Dacnusini (Alysiinae) is mainly the work of HCJG, who also contributed to the Alysiini and Opiinae sections. The rest of the work was a joint effort between GRB and MRS, with the editing done by GRB.