1	Elytra with punctures randomly and uniformly distributed (see Fig. 1)	2
–	Elytra with punctures arranged as well defined longitudinal series, forming striae (see Figs 2, 3)	3
2	Elevation of mesoventrite as a narrow toothlike projection	Q. dentatus Hansen (Fig. 1A–D)
–	Elevation of mesoventrite as a wide, transverse, straight blunt carina	Q. acroreius sp. n. (Fig. 1E–H)
3	Serial elytral punctures simple; the ones on striae similar in size as those on the interstrial surface (Figs 6A, 3E–F)	Q. clusia sp. n.
–	Serial elytral punctures ramified; the ones on striae evidently larger in size than those on the interstrial surface (Fig. 6B)	4
4	Surface of pseudepipleura smooth throughout; dorsal outline of body in lateral view moderately and uniformly convex	Q. reticulatus Hansen (Fig. 3A–D)
–	Surface of pseudepipleura posteriorly markedly canaliculated (e.g. Fig. 2G), dorsal outline of body in lateral view nearly flat	5
5	Apex of parameres widely to narrowly rounded (Figs 7A–D)	Q. depressus Hansen (Fig. 2A–D)
–	Apex of parameres angulate (Fig. 8)	Q. similaris Hansen (Fig. 2E–G)