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Nature Communications 5: Article number: 4997; DOI: 10.1038/ncomms5997 (2014); Published: September 29 2014; Updated: March 26 2018
The original HTML version of this Article had an incorrect article number of 3997; it should have been 4997. This has now been corrected in the HTML; the PDF version of the Article was correct from the time of publication.
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