1Molecular and Population Genetics Laboratory University of Oxford, Oxford, OX3 7BN UK
2Clinical Trial Service Unit and Epidemiological Studies Unit, Nuffield Department of Population Health, Old Road Campus, Oxford, OX3 7DQ UK
3Ludwig Institute for Cancer Research, University of Oxford, Old Road Campus Research Building, Roosevelt Drive, Oxford, OX3 7DQ UK
4Translational Gastroenterology Unit, Experimental Medicine Division, Nuffield Department of Clinical Medicine, John Radcliffe Hospital, Oxford, OX3 9DU UK
5Gastrointestinal Stem Cell Biology Laboratory, Oxford Centre for Cancer Gene Research, Oxford, UK
6Oxford NIHR Biomedical Research Centre, University of Oxford, Oxford, UK
7Institute of Cancer and Genomic Sciences, College of Medical and Dental Sciences, University of Birmingham, Vincent Drive, Edgbaston, Birmingham, B15 2TT UK
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